Common Scale-backed Antbird
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The common scale-backed antbird (''Willisornis poecilinotus'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
bird in subfamily Thamnophilinae of family
Thamnophilidae The antbirds are a large passerine bird family, Thamnophilidae, found across subtropical and tropical Central and South America, from Mexico to Argentina. There are more than 230 species, known variously as antshrikes, antwrens, antvireos, fire ...
, the "typical antbirds". It is found in
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
,
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
,
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
,
French Guiana French Guiana ( or ; french: link=no, Guyane ; gcr, label=French Guianese Creole, Lagwiyann ) is an overseas departments and regions of France, overseas department/region and single territorial collectivity of France on the northern Atlantic ...
,
Guyana Guyana ( or ), officially the Cooperative Republic of Guyana, is a country on the northern mainland of South America. Guyana is an indigenous word which means "Land of Many Waters". The capital city is Georgetown. Guyana is bordered by the ...
,
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
,
Suriname Suriname (; srn, Sranankondre or ), officially the Republic of Suriname ( nl, Republiek Suriname , srn, Ripolik fu Sranan), is a country on the northeastern Atlantic coast of South America. It is bordered by the Atlantic Ocean to the north ...
, and
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
.


Taxonomy and systematics

The common scale-backed antbird has a complicated taxonomic history. It was described and illustrated by the German ornithologist
Jean Cabanis Jean Louis Cabanis (8 March 1816 – 20 February 1906) was a German ornithologist. Cabanis was born in Berlin to an old Huguenot family who had moved from France. Little is known of his early life. He studied at the University of Berlin from 18 ...
in 1847 and given the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Hypocnemis poecilinotus''. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
is from the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''poikilonōtos'' "with variegated back" (from ''poikilos'' "spotted" and ''nōton'' "back"). It was subsequently included in the genus ''
Hylophylax ''Hylophylax'' is a genus of bird in the family Thamnophilidae. The genus ''Hylophylax'' was erected by the American ornithologist Robert Ridgway in 1909 with the spotted antbird as the type species. It contains three species: * Spotted antbird ...
'', but was found to not be closely related to the other species in the genus and then was briefly placed in genus ''Dichropogon''. This name is preoccupied by a genus of asilid flies (''Dichropogon'' Bezzi, 1910) so the current genus ''Willisornis'' was created for it. The common scale-backed antbird has these five subspecies: * ''W. p. poecilinotus'' (
Cabanis Cabanis is the surname of: * George Cabanis (1815-1892), American politician *Jean Cabanis (1816–1906), German ornithologist * José Cabanis (1922–2000), French writer, historian and magistrate *Pierre Jean George Cabanis Pierre Jean Georges ...
, 1847)
* ''W. p. duidae'' (
Chapman Chapman may refer to: Businesses * Chapman Entertainment, a former British television production company * Chapman Guitars, a guitar company established in 2009 by Rob Chapman * Chapman's, a Canadian ice cream and ice water products manufacturer ...
, 1923)
* ''W. p. lepidonota'' ( Sclater, PL & Salvin, 1880) * ''W. p. griseiventris'' ( Pelzeln, 1868) * ''W. p. gutturalis'' (
Todd Todd or Todds may refer to: Places ;Australia: * Todd River, an ephemeral river ;United States: * Todd Valley, California, also known as Todd, an unincorporated community * Todd, Missouri, a ghost town * Todd, North Carolina, an unincorporated ...
, 1927)
The
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
s of the subspecies differ greatly, leading to speculation that some of them should be recognized as full species. The South American Classification Committee of the
American Ornithological Society The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its m ...
is seeking a formal proposal to consider the issue.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, G. Del-Rio, A. Jaramillo, D. F. Lane, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 4 March 2024. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved March 5, 2024 What are now the two subspecies of the Xingu scale-backed antbird (''W. vidua'') were previously included in ''W. poecilinotus''.


Description

The common scale-backed antbird is long and weighs . Adult males of the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
''W. p. poecilinotus'' are mostly gray; their upperparts are darker than their underparts. They have a white patch between the
scapulars The scapula (plural scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on either ...
. Their lower back has white-edged black patches that give it its English name. Their wings and tail are black with white edges and tips on many feathers. Adult females have a reddish yellow-brown crown and face. Their upperparts are mostly olive-brown with buff-edged brownish black patches on the lower back. They also have a white patch between the scapulars. Their flight feathers are blackish brown with reddish brown edges and their tail is dark olive-brown with white spots and a blackish band near the end. Their throat is pale grayish white. Their underparts are gray with a reddish yellow-brown tinge on the flanks.Zimmer, K., M.L. Isler, and G. M. Kirwan (2020). Common Scale-backed Antbird (''Willisornis poecilinotus''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.scbant3.01 retrieved June 13, 2024 Females of subspecies ''W. p. duidae'' have a black lower back and rump with wide white feather edges, a white-spotted black tail, reddish yellow-brown underparts with a cinnamon-rufous center to the belly, and olive-brown tinged flanks. Females of ''W. p. lepidonota'' are similar to those of ''duidae'' but paler and with a browner crown and a light buff center to the belly. Females of ''W. p. griseiventris'' have a rufous-buff face, a plain lower back and rump, a pale gray throat, and gray underparts. Males of ''W. p. gutturalis'' have a black throat; females resemble female ''lepidonota'' but with a pale olive center to the belly.Schulenberg, T.S., D.F. Stotz, D.F. Lane, J.P. O’Neill, and T.A. Parker III. 2010. ''Birds of Peru''. Revised and updated edition. Princeton University Press, Princeton, New Jersey. Plate 175


Distribution and habitat

The subspecies of the common scale-backed antbird are found thus: * ''W. p. poecilinotus'': central and southeastern Venezuela, the
Guiana The Guianas, sometimes called by the Spanish loan-word ''Guayanas'' (''Las Guayanas''), is a region in north-eastern South America which includes the following three territories: * French Guiana, an overseas department and region of France * ...
s, and northeastern Brazil from
Roraima Roraima (, ) is one of the 26 states of Brazil. Located in the country's North Region, it is the northernmost and most geographically and logistically isolated state in Brazil. It is bordered by the state of Pará to the southeast, Amazonas ...
to
Amapá Amapá () is one of the 26 states of Brazil. It is in the northern region of Brazil. It is the second least populous state and the eighteenth largest by area. Located in the far northern part of the country, Amapá is bordered clockwise by Fr ...
* ''W. p. duidae'': east-central Colombia, southwestern Venezuela, and northwestern Brazil along the Rio Negro * ''W. p. lepidonota'': southeastern Colombia, eastern Ecuador, and northeastern and east-central Peru * ''W. p. griseiventris'': south-central and southeastern Peru, northern Bolivia, and southwestern Amazonian Brazil between the rios Juruá and
Madeira ) , anthem = ( en, "Anthem of the Autonomous Region of Madeira") , song_type = Regional anthem , image_map=EU-Portugal_with_Madeira_circled.svg , map_alt=Location of Madeira , map_caption=Location of Madeira , subdivision_type=Sovereign st ...
south into western
Mato Grosso Mato Grosso ( – lit. "Thick Bush") is one of the states of Brazil, the third largest by area, located in the Central-West region. The state has 1.66% of the Brazilian population and is responsible for 1.9% of the Brazilian GDP. Neighboring ...
* ''W. p. gutturalis'': northeastern Peru and western Amazonian Brazil to the Rio Juruá The common scale-backed antbird primarily inhabits the understorey of humid ''
terra firme Terra may often refer to: * Terra (mythology), primeval Roman goddess * An alternate name for planet Earth, as well as the Latin name for the planet Terra may also refer to: Geography Astronomy * Terra (satellite), a multi-national NASA scienti ...
''
evergreen forest An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, Live oak, acacias, magnolia, and banksia in more temperate zone ...
. It also occurs in '' várzea'' and ''
igapó Igapó (, from Old Tupi: "root forest") is a word used in Brazil for blackwater-flooded forests in the Amazon biome. These forests and similar swamp forests are seasonally inundated with freshwater. They typically occur along the lower reaches of ...
'' forest. In elevation it reaches in Brazil, in Venezuela, and in Colombia. In Ecuador it mostly occurs below but does reach . In Peru it occurs locally to but is mostly below .


Behavior


Movement

The common scale-backed antbird is believed to be a year-round resident throughout its range.


Feeding

The common scale-backed antbird feeds mostly on a wide variety of
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s; it has also been noted eating small lizards. It typically forages individually, in pairs, and in family groups, usually within about of the ground though sometimes as high as and rarely to . In some parts of its range it joins
mixed-species feeding flock A mixed-species feeding flock, also termed a mixed-species foraging flock, mixed hunting party or informally bird wave, is a flock of usually insectivorous birds of different species that join each other and move together while foraging. These ar ...
s. It follows swarms of
army ant The name army ant (or legionary ant or ''marabunta'') is applied to over 200 ant species in different lineages. Because of their aggressive predatory foraging groups, known as "raids", a huge number of ants forage simultaneously over a limit ...
s that pass through its territory to capture prey fleeing the ants, but it just as often forages away from ant swarms. It captures prey mostly by short sallies from a perch to the ground but also to foliage, branches, and vines. It sometimes stays on the ground probing and flicking leaf litter. At ant swarms it defers to other species.


Breeding

The breeding season of the common scale-backed antbird varies greatly across its range, for instance spanning November to March in French Guiana and probably December to July in Ecuador. It possibly breeds in any month in Brazil. It has been documented building several types of nests, including an open cup of dry leaves and other fibers in an understorey palm, a cavity lined with dead leaves, an unlined hollow at the top of a stump, and a cup of dried grass on the ground. The usual clutch is two eggs. The female alone is believed to incubate at night but both parents do so during the day. Both parents also brood and provision nestlings. The incubation period and time to fledging are not known.


Vocalization

The common scale-backed antbird's songs are generally " a series...of long upslurred notes...with little space between them, each note rising in pitch and gaining in intensity, except final note or notes that decrease in intensity". The number of notes and some other features differ among individuals and also among the subspecies. In Venezuela it has been written as "''pureeeee, pureeeee, pureeeee...''", in Ecuador as "teeuw, tuweeé? tuweeé? tuweeé? tuweeé?", and in Peru as "''hew hui hui hui? hui? hui? hui? hui? hui?''". Its calls include a "sharp 'psit'...a usually descending 'chirr', more abrupt chitter, short whistle quickly repeated 3–4 times, ndalso a longer whistle that falls and then rises in pitch, sounding like 'cherri' ".


Status

The
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
has assessed the common scale-backed antbird as being of Least Concern. It has a very large range. Its population size is not known and is believed to be stable. No immediate threats have been identified. It is considered fairly common throughout its range, which includes many large protected areas. "Regions occupied by this species also encompass extensive intact habitat which, although not formally protected, seems to be at little near-term risk."


References


External links


Scale-backed antbird photo gallery
VIREO
Photo
chandra.as.utexas.edu {{Taxonbar, from=Q1063238 Willisornis Birds of the Amazon rainforest Birds of the Guiana Shield Birds described in 1847 Taxonomy articles created by Polbot