The Coelosclerithophorans are a
polyphyletic
A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of converg ...
group of organisms bearing hollow
sclerites
A sclerite (Greek , ', meaning "hard") is a hardened body part. In various branches of biology the term is applied to various structures, but not as a rule to vertebrate anatomical features such as bones and teeth. Instead it refers most commonly ...
made of
aragonite
Aragonite is a carbonate mineral, one of the three most common naturally occurring crystal forms of calcium carbonate, (the other forms being the minerals calcite and vaterite). It is formed by biological and physical processes, including prec ...
, and with a supposedly distinctive microstructure.
Their skeletons may be homologous to those of the molluscs,
and ''
Halkieria
The halkieriids are a group of fossil organisms from the Lower to Middle Cambrian. Their eponymous genus is ''Halkieria'' , which has been found on almost every continent in Lower to Mid Cambrian deposits, forming a large component of the smal ...
'' looks very much like a
polyplacophoran
Chitons () are marine (ocean), marine molluscs of varying size in the class (biology), class Polyplacophora (), formerly known as Amphineura. About 940 Extant taxon, extant and 430 fossil species are recognized.
They are also sometimes known ...
. The shells of this group and the molluscs appear to have originated in the same fashion.
[ However, there are several reasons to doubt this relationship,] many of which may not pass muster.
There are two groups within the coeloscleritophorans: the Sachitids, to which ''Halkieria
The halkieriids are a group of fossil organisms from the Lower to Middle Cambrian. Their eponymous genus is ''Halkieria'' , which has been found on almost every continent in Lower to Mid Cambrian deposits, forming a large component of the smal ...
'' and ''Wiwaxia
''Wiwaxia'' is a genus of soft-bodied animals that were covered in carbonaceous scales and spines that protected it from predators. ''Wiwaxia'' fossils – mainly isolated scales, but sometimes complete, articulated fossils – are known from ear ...
'' belong, and the Chancellorids
The Chancelloriids are an extinct family of superficially sponge-like animals common in sediments from the Early Cambrian to the early Late Cambrian. Many of these fossils consists only of spines and other fragments, and it is not certain that t ...
. The Ediacaran fossil '' Ausia'' has been touted as an ancestral 'intermediate' between both of these groups.
These are common constituents of Lower Cambrian
The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...
small shelly fossil
The small shelly fauna, small shelly fossils (SSF), or early skeletal fossils (ESF) are mineralized fossils, many only a few millimetres long, with a nearly continuous record from the latest stages of the Ediacaran to the end of the Early Cambri ...
assemblages.
References
Cambrian invertebrates
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