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Halkieria
The halkieriids are a group of fossil organisms from the Lower to Middle Cambrian. Their eponymous genus is ''Halkieria'' , which has been found on almost every continent in Lower to Mid Cambrian deposits, forming a large component of the small shelly fossil assemblages. The best known species is ''Halkieria evangelista'', from the North Greenland Sirius Passet Lagerstätte, in which complete specimens were collected on an expedition in 1989. The fossils were described by Simon Conway Morris and John Peel in a short paper in 1990 in the journal ''Nature''. Later a more thorough description was undertaken in 1995 in the journal ''Philosophical Transactions of the Royal Society of London'' and wider evolutionary implications were posed. The group is sometimes equated to Sachitida, although as originally envisaged, this group includes the wiwaxiids and is thus equivalent to the Halwaxiida. History of discovery Armor plates called "sclerites" had long been known as elements of t ...
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Halwaxiid
Halwaxiida or halwaxiids is a proposed clade equivalent to the older orders Sachitida He 1980 and Thambetolepidea Jell 1981, loosely uniting scale-bearing Cambrian animals, which may lie in the stem group to molluscs or lophotrochozoa. Some palaeontologists question the validity of the Halwaxiida clade. Definition and members The name "halwaxiid" was formed by combining the names of two members of the proposed group, ''Halkieria'' and ''Wiwaxia''. The group was defined as a set of Early to Mid Cambrian animals that had: a "chain mail" coat of three concentric bands of small armor plates that are called sclerites; in some cases, a small cap-like shell at the front end and in some cases both ends. Some scientists are unhappy with this loose definition, arguing that such traits may have arisen convergently rather than being inherited from a common ancestor. This objection implies that the group may not be monophyletic, and that their similarities are not biologically equivalent b ...
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Small Shelly Fossil
The small shelly fauna, small shelly fossils (SSF), or early skeletal fossils (ESF) are mineralized fossils, many only a few millimetres long, with a nearly continuous record from the latest stages of the Ediacaran to the end of the Early Cambrian Period. They are very diverse, and there is no formal definition of "small shelly fauna" or "small shelly fossils". Almost all are from earlier rocks than more familiar fossils such as trilobites. Since most SSFs were preserved by being covered quickly with phosphate and this method of preservation is mainly limited to the late Ediacaran and early Cambrian periods, the animals that made them may actually have arisen earlier and persisted after this time span. Some of the fossils represent the entire skeletons of small organisms, including the mysterious '' Cloudina'' and some snail-like molluscs. However, the bulk of the fossils are fragments or disarticulated remains of larger organisms, including sponges, molluscs, slug-like halkierii ...
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Sirius Passet
Sirius Passet is a Cambrian Lagerstätte in Peary Land, Greenland. The Sirius Passet Lagerstätte was named after the Sirius sledge patrol that operates in North Greenland. It comprises six places in Nansen Land, on the east shore of J.P. Koch Fjord in the far north of Greenland. It was discovered in 1984 by A. Higgins of the Geological Survey of Greenland. A preliminary account was published by Simon Conway Morris and others in 1987 and expeditions led by J. S. Peel and Conway Morris have returned to the site several times between 1989 and the present. A field collection of perhaps 10,000 fossil specimens has been amassed. It is a part of the Buen Formation. Age The fauna is inevitably compared to that of the Burgess Shale, although it is probably ten to fifteen million years older – vs. ) – and more closely contemporaneous with the fauna of the Maotianshan shales from Chengjiang, which are dated to . Preservation The preservation of the Sirius Passet is traditionall ...
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Small Shelly Fossils
The small shelly fauna, small shelly fossils (SSF), or early skeletal fossils (ESF) are mineralized fossils, many only a few millimetres long, with a nearly continuous record from the latest stages of the Ediacaran to the end of the Early Cambrian Period. They are very diverse, and there is no formal definition of "small shelly fauna" or "small shelly fossils". Almost all are from earlier rocks than more familiar fossils such as trilobites. Since most SSFs were preserved by being covered quickly with phosphate and this method of preservation is mainly limited to the late Ediacaran and early Cambrian periods, the animals that made them may actually have arisen earlier and persisted after this time span. Some of the fossils represent the entire skeletons of small organisms, including the mysterious '' Cloudina'' and some snail-like molluscs. However, the bulk of the fossils are fragments or disarticulated remains of larger organisms, including sponges, molluscs, slug-like halki ...
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Sclerites
A sclerite (Greek , ', meaning "hard") is a hardened body part. In various branches of biology the term is applied to various structures, but not as a rule to vertebrate anatomical features such as bones and teeth. Instead it refers most commonly to the hardened parts of arthropod exoskeletons and the internal spicules of invertebrates such as certain sponges and soft corals. In paleontology, a scleritome is the complete set of sclerites of an organism, often all that is known from fossil invertebrates. Sclerites in combination Sclerites may occur practically isolated in an organism, such as the sting of a cone shell. Also, they can be more or less scattered, such as tufts of defensive sharp, mineralised bristles as in many marine Polychaetes. Or, they can occur as structured, but unconnected or loosely connected arrays, such as the mineral "teeth" in the radula of many Mollusca, the valves of Chitons, the beak of Cephalopod, or the articulated exoskeletons of Arthropoda. When sc ...
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Sclerite
A sclerite (Greek , ', meaning "hard") is a hardened body part. In various branches of biology the term is applied to various structures, but not as a rule to vertebrate anatomical features such as bones and teeth. Instead it refers most commonly to the hardened parts of arthropod exoskeletons and the internal spicules of invertebrates such as certain sponges and soft corals. In paleontology, a scleritome is the complete set of sclerites of an organism, often all that is known from fossil invertebrates. Sclerites in combination Sclerites may occur practically isolated in an organism, such as the sting of a cone shell. Also, they can be more or less scattered, such as tufts of defensive sharp, mineralised bristles as in many marine Polychaetes. Or, they can occur as structured, but unconnected or loosely connected arrays, such as the mineral "teeth" in the radula of many Mollusca, the valves of Chitons, the beak of Cephalopod, or the articulated exoskeletons of Arthropoda. When ...
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Mollusc
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied. Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat. The phylum is typically divided into 7 or 8 taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known invertebrate species. The gastropods ...
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Chitin
Chitin ( C8 H13 O5 N)n ( ) is a long-chain polymer of ''N''-acetylglucosamine, an amide derivative of glucose. Chitin is probably the second most abundant polysaccharide in nature (behind only cellulose); an estimated 1 billion tons of chitin are produced each year in the biosphere. It is a primary component of cell walls in fungi (especially basidiomycetes and filamentous fungi), the exoskeletons of arthropods such as crustaceans and insects, the radulae, cephalopod beaks and gladii of molluscs and in some nematodes and diatoms. It is also synthesised by at least some fish and lissamphibians. Commercially, chitin is extracted from the shells of crabs, shrimps, shellfishes and lobsters, which are major by-products of the seafood industry. The structure of chitin is comparable to cellulose, forming crystalline nanofibrils or whiskers. It is functionally comparable to the protein keratin. Chitin has proved useful for several medicinal, industrial and biotechnological purpos ...
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Botomian
Cambrian Stage 4 is the still unnamed fourth stage of the Cambrian and the upper stage of Cambrian Series 2. It follows Cambrian Stage 3 and lies below the Wuliuan. The lower boundary has not been formally defined by the International Commission on Stratigraphy. One proposal is the first appearance of two trilobite genera, ''Olenellus'' or ''Redlichia''. Another proposal is the first appearance of the trilobite species '' Arthricocephalus chauveaui''. Both proposals will set the lower boundary close to million years ago. The upper boundary corresponds to the beginning of the Wuliuan. Naming The International Commission on Stratigraphy The International Commission on Stratigraphy (ICS), sometimes referred to unofficially as the "International Stratigraphic Commission", is a daughter or major subcommittee grade scientific daughter organization that concerns itself with stratigra ... has not named the fourth stage of the Cambrian yet. In the widely used Siberian nomenclature stage ...
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Growth Ring
Dendrochronology (or tree-ring dating) is the scientific method of dating tree rings (also called growth rings) to the exact year they were formed. As well as dating them, this can give data for dendroclimatology, the study of climate and atmospheric conditions during different periods in history from wood. Dendrochronology derives from Ancient Greek (), meaning "tree", (), meaning "time", and (), "the study of". Dendrochronology is useful for determining the precise age of samples, especially those that are too recent for radiocarbon dating, which always produces a range rather than an exact date. However, for a precise date of the death of the tree a full sample to the edge is needed, which most trimmed timber will not provide. It also gives data on the timing of events and rates of change in the environment (most prominently climate) and also in wood found in archaeology or works of art and architecture, such as old panel paintings. It is also used as a check in radiocarb ...
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Olfaction
The sense of smell, or olfaction, is the special sense through which smells (or odors) are perceived. The sense of smell has many functions, including detecting desirable foods, hazards, and pheromones, and plays a role in taste. In humans, it occurs when an odor binds to a receptor within the nasal cavity, transmitting a signal through the olfactory system. Glomeruli aggregate signals from these receptors and transmit them to the olfactory bulb, where the sensory input will start to interact with parts of the brain responsible for smell identification, memory, and emotion. There are many different causes for alteration, lack, or disturbance to a normal sense of smell, and can include damage to the nose or smell receptors, or central problems affecting the brain. Some causes include upper respiratory infections, traumatic brain injury, and neurodegenerative disease. History of study Early scientific study of the sense of smell includes the extensive doctoral dissertation of ...
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Radula
The radula (, ; plural radulae or radulas) is an anatomical structure used by molluscs for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food enters the esophagus. The radula is unique to the molluscs, and is found in every class of mollusc except the bivalves, which instead use cilia, waving filaments that bring minute organisms to the mouth. Within the gastropods, the radula is used in feeding by both herbivorous and carnivorous snails and slugs. The arrangement of teeth ( denticles) on the radular ribbon varies considerably from one group to another. In most of the more ancient lineages of gastropods, the radula is used to graze, by scraping diatoms and other microscopic algae off rock surfaces and other substrates. Predatory marine snails such as the Naticidae use the radula plus an acidic secretion to bore through the shell of other molluscs. Other predatory marine snails ...
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