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Cochliopidae is a
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of small
freshwater snail Freshwater snails are gastropod mollusks which live in fresh water. There are many different families. They are found throughout the world in various habitats, ranging from ephemeral pools to the largest lakes, and from small seeps and springs ...
s with
gills A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are ...
and an operculum, aquatic
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
s. Paludestrina d'Orbigny, 1840 is an archaic synonym, and has been placed on the Official Index of Rejected and Invalid Names by
ICZN The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the I ...
Opinion 2202.MolluscaBase eds. (2021). MolluscaBase. Cochliopidae Tryon, 1866. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=411706 on 2021-05-13 This family is in the superfamily
Truncatelloidea Truncatelloidea is a superfamily of snails, gastropod mollusks in the clade Caenogastropoda.Bouchet, P. (2013). ''Truncatelloidea''. Retrieved through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=72 ...
and in the order
Littorinimorpha Littorinimorpha is a large order of snails, gastropods, consisting primarily of sea snails ( marine species), but also including some freshwater snails ( aquatic species) and land snails (terrestrial species).Bouchet P. & Rocroi J.-P. (Ed.); Frà ...
(according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).


Description

Cochliopidae snails are characterized by sharp, elongated spire-like structures, and can be found in rivers or brackish water.


2005 taxonomy

The family Cochliopidae consists of 3 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005): * Cochliopinae Tryon, 1866 - synonyms: Mexithaumatinae D. W. Taylor, 1966, Paludiscalinae D. W. Taylor, 1966 * Littoridininae Thiele, 1928 * Semisalsinae Guiusti & Pezzoli, 1980 - synonym: Heleobiini Bernasconi, 1991


Genera

Liu et al. (2001) have recognized 34 genera with more than 260 species within the subfamily Cochliopinae. Strong et al. (2008)Strong E. E., Gargominy O., Ponder W. F. & Bouchet P. (2008). "Global Diversity of Gastropods (Gastropoda; Mollusca) in Freshwater". ''
Hydrobiologia ''Hydrobiologia'', ''The International Journal of Aquatic Sciences'', is a peer-reviewed scientific journal publishing 21 issues per year, for a total of well over 4000 pages per year. ''Hydrobiologia'' publishes original research, reviews and opi ...
'' 595: 149-166. .
have recognized 246 freshwater species within Cochliopidae. Genera within the family Cochliopidae include: subfamily Cochliopinae *'' Aphaostracon'' F. G. Thompson, 1968 *'' Coahuilix''
Taylor Taylor, Taylors or Taylor's may refer to: People * Taylor (surname) **List of people with surname Taylor * Taylor (given name), including Tayla and Taylah * Taylor sept, a branch of Scottish clan Cameron * Justice Taylor (disambiguation) Plac ...
, 1966
*''
Cochliopa ''Cochliopa'' is a genus of freshwater snails, specifically cave snails, aquatic animal, aquatic gastropod mollusks in the family Cochliopidae. ''Cochliopa'' is the type genus of the family Cochliopidae. This genus is not to be confused with th ...
'' Stimpson, 1865 - type genus of the family Cochliopidae *'' Cochliopina'' Morrison, 1946 subfamily Littoridininae *''Antrobia'' Hubricht, 1971 - with the only species ''
Antrobia culveri The Tumbling Creek cavesnail (''Antrobia culveri'') is a species of freshwater cave snail with gills and an operculum, an aquatic gastropod mollusk in the family Amnicolidae.Bouchet, P. (2014). Antrobia culveri Hubricht, 1971. Accessed through ...
'' Hubricht, 1971Kabat A. R. & Hershler R. (1993). "The prosobranch snail family Hydrobiidae (Gastropoda: Rissooidea): review of classification and supraspecific taxa". ''
Smithsonian Contributions to Zoology The Smithsonian Contributions and Studies Series is a collection of serial periodical publications produced by the Smithsonian Institution, detailing advances in various scientific and societal fields to which the Smithsonian Institution has made c ...
'' 547: 1-94
PDF
-
Tumbling Creek cavesnail The Tumbling Creek cavesnail (''Antrobia culveri'') is a species of freshwater cave snail with gills and an operculum, an aquatic gastropod mollusk in the family Amnicolidae.Bouchet, P. (2014). Antrobia culveri Hubricht, 1971. Accessed throug ...
*''
Littoridina ''Littoridina'' is a genus of small freshwater snails with a gill and an operculum, aquatic gastropod mollusks in the family Cochliopidae Cochliopidae is a family of small freshwater snails with gills and an operculum, aquatic gastropod ...
'' Souleyet, 1852 *'' Pseudotryonia'' Hershler, 2001 **''
Pseudotryonia adamantina ''Pseudotryonia'' is a genus of gastropods belonging to the family Cochliopidae. The species of this genus are found in Northern America. Species: *'' Pseudotryonia adamantina'' *'' Pseudotryonia alamosae'' *'' Pseudotryonia brevissima'' *'' ...
'' **''
Pseudotryonia alamosae ''Pseudotryonia'' is a genus of gastropods belonging to the family Cochliopidae. The species of this genus are found in Northern America. Species: *''Pseudotryonia adamantina'' *'' Pseudotryonia alamosae'' *'' Pseudotryonia brevissima'' *'' ...
'' **''
Pseudotryonia brevissima ''Pseudotryonia'' is a genus of gastropods belonging to the family Cochliopidae. The species of this genus are found in Northern America. Species: *''Pseudotryonia adamantina'' *''Pseudotryonia alamosae'' *'' Pseudotryonia brevissima'' *'' P ...
'' **''
Pseudotryonia grahamae ''Pseudotryonia'' is a genus of gastropods belonging to the family Cochliopidae. The species of this genus are found in Northern America. Species: *''Pseudotryonia adamantina'' *''Pseudotryonia alamosae'' *''Pseudotryonia brevissima'' *'' Ps ...
'' **''
Pseudotryonia mica ''Pseudotryonia'' is a genus of gastropods belonging to the family Cochliopidae Cochliopidae is a family of small freshwater snails with gills and an operculum, aquatic gastropod mollusks. Paludestrina d'Orbigny, 1840 is an archaic synonym ...
'' Hershler, Liu & Landye, 2011Hershler R., Liu H.-P. & Landye J. J. (2011). "Two new genera and four new species of freshwater cochliopid gastropods (Rissooidea) from northeastern Mexico". ''
Journal of Molluscan Studies The ''Journal of Molluscan Studies'' is the peer-reviewed scientific journal of the Malacological Society of London, covering research in malacology.
'' 77(1): 8-23. .
**''
Pseudotryonia pasajae ''Pseudotryonia'' is a genus of gastropods belonging to the family Cochliopidae. The species of this genus are found in Northern America. Species: *''Pseudotryonia adamantina'' *''Pseudotryonia alamosae'' *''Pseudotryonia brevissima'' *''Pse ...
'' Hershler, Liu & Landye, 2011 subfamily Semisalsinae - there are three genera in the subfamily SemisalsinaeKroll O., Hershler R., Albrecht C., Terrazas E. M., Apaza R., Fuentealba C., Wolff C. & Wilke T. (2012). "The endemic gastropod fauna of Lake Titicaca: correlation between molecular evolution and hydrographic history". ''
Ecology and Evolution ''Ecology and Evolution'' is a biweekly open-access scientific journal covering all areas of ecology, evolution, and conservation. The Editors in Chief of this journal are Allen Moore, Andrew Beckerman, Jenn Firn, Chris Foote, and Gareth Jenkins. ...
'' 2(7): 1517-1530. .
*''
Heleobia ''Heleobia'' is a genus of small freshwater and brackish water snails with a gill and an operculum, aquatic gastropod mollusks in the family Cochliopidae and the superfamily Truncatelloidea.Bouchet, P.; Gofas, S. (2015). Heleobia. Accessed thro ...
'' Stimpson, 1865 *''
Heleobops ''Heleobops'' is a genus of very small aquatic snails, operculate gastropod mollusks in the family Cochliopidae. ''Heleobops'' is one of three genera (together with ''Heleobia'' and '' Semisalsa'') within the subfamily Semisalsinae. Species Sp ...
'' Thompson, 1968 *'' Semisalsa'' Radoman, 1974 subfamily ? (either Cochliopinae or Littoridininae) *'' Aroapyrgus'' H. B. Baker, 1931 *'' Balconorbis'' Hershler & Longley, 1986 *† '' Carinulorbis'' Yen, 1949 *'' Chorrobius'' Hershler, Liu & Landye, 2011 *'' Dyris'' Conrad, 1871 - it has extant species and also 26 species in Miocene Pebas Formation *'' Emmericiella''
Pilsbry Henry Augustus Pilsbry (7 December 1862 – 26 October 1957) was an American biologist, malacologist and carcinologist, among other areas of study. He was a dominant presence in many fields of invertebrate taxonomy for the better part of a centu ...
, 1909
*'' Eremopyprgus'' Hershler, 1999Hershler R., Liu H.-P. & Landye J. J. (2002). "A new species of ''Eremopyprgus'' (Hydrobiidae: Cochliopinae) from the Chihuahuan desert, Mexico: Phylogentic relationships and biogeography". ''
Journal of Molluscan Studies The ''Journal of Molluscan Studies'' is the peer-reviewed scientific journal of the Malacological Society of London, covering research in malacology.
'' 68: 7-13
PDF
Hershler R. (1999). "A systematic review of the hydrobiid snails (Gastropoda: Rissoidea) of the Great Basin, western United States. Part II. Genera ''Colligyrus'', ''Fluminicola'', ''Pristinicola'', and ''Tryonia''". ''The Veliger'' 42(4): 306-337
PDF
*† '' Feliconcha'' Wesselingh, Anderson & Kadolsky, 2006 - two species from Miocene of the
Pebas Formation The Pebas Formation is a lithostratigraphic unit of Miocene age, found in western Amazonia. The formation extends over , including parts of Brazil, Peru, Ecuador and Colombia.Wesselingh et al., 2006 It is interpreted as representing the deposits ...
Wesselingh F. P., Anderson L. C. & Kadolsky D. (2006). "Molluscs from the Miocene Pebas Formation of Peruvian and Colombian Amazonia". ''
Scripta Geologica ''Scripta Geologica'' is a peer-reviewed scientific journal that publishes on vertebrate and invertebrate palaeontology, palaeobotany/palynology, stratigraphy, petrology, and mineralogy, including gemmology with a focus on systematics. It is publis ...
'' 1333: 19-290
PDF
**'' Feliconcha feliconcha'' Wesselingh, Anderson & Kadolsky, 2006 **'' Feliconcha reticulata'' Wesselingh, Anderson & Kadolsky, 2006 *(probably extant) '' Glabertryonia'' Wesselingh, Anderson & Kadolsky, 2006 - three species **† '' Glabertryonia glabra'' Wesselingh, Anderson & Kadolsky, 2006 - from Miocene of the Pebas Formation **† ''Glabertryonia'' sp. 1 - from Pliocene of the Las Piedras Formation **(probably extant) ''Glabertryonia'' sp. 2 - from Holocene of Surinam, probably extant *'' Juturnia'' Hershler, Liu & Stockwell, 2002 *'' Lithococcus''
Pilsbry Henry Augustus Pilsbry (7 December 1862 – 26 October 1957) was an American biologist, malacologist and carcinologist, among other areas of study. He was a dominant presence in many fields of invertebrate taxonomy for the better part of a centu ...
, 1911
*'' Mesobia'' F. G. Thompson & Hershler, 1991 *'' Mexipyrgus''
Taylor Taylor, Taylors or Taylor's may refer to: People * Taylor (surname) **List of people with surname Taylor * Taylor (given name), including Tayla and Taylah * Taylor sept, a branch of Scottish clan Cameron * Justice Taylor (disambiguation) Plac ...
, 1966
*''
Minckleyella ''Minckleyella'' is a monotypic genus of gastropods belonging to the family Cochliopidae. The only species is ''Minckleyella balnearis''. This freshwater snail genus is named in honor of Wendell L. Minckley.Hershler R., Liu H.-P. & Landye J. J. ...
'' Hershler, Liu & Landye, 2011 - with the only species '' Minckleyella balnearis'' Hershler, Liu & Landye, 2011 * '' Onobops'' Thompson, 1968 * ''
Pyrgophorus ''Pyrgophorus'' is a genus of very small freshwater snails with a gill and an operculum, aquatic gastropod molluscs in the family Hydrobiidae. Species Species in the genus ''Pyrgophorus'' include: * '' Sioliella'' Haas, 1949 * '' Spurwinkia'' Davis, Mazurkiewicz & Mandracchia, 1982 * '' Texadina'' Abbott & Ladd, 1951 * '' Thalassobia'' Bourguignat in Mabille, 1877


Cladogram

A
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...
based on sequences of mitochondrial cytochrome-c oxidase I (COI) genes showing phylogenic relations of species within Cochliopidae:Liu H.-P., Hershler R. & Thompson F. G. (2001). "Phylogenetic Relationships of the Cochliopinae (Rissooidea: Hydrobiidae): An Enigmatic Group of Aquatic Gastropods". ''
Molecular Phylogenetics and Evolution ''Molecular Phylogenetics and Evolution'' is a peer-reviewed scientific journal of evolutionary biology and phylogenetics. The journal is edited by E.A. Zimmer. Indexing The journal is indexed in: *EMBiology *Journal Citation Reports *Scopus ...
'' 21(1): 17-25. .


References


Further reading

* Hershler R., Davis C. L., Kitting C. L & Liu H.-P. (2007). "Discovery of introduced and cryptogenic cochliopid gastropods in the San Francisco Estuary, California". ''
Journal of Molluscan Studies The ''Journal of Molluscan Studies'' is the peer-reviewed scientific journal of the Malacological Society of London, covering research in malacology.
'' 73(4): 323-332
PDF
* Hershler R. & Thompson F. G. (1992). "A Review of the Aquatic Gastropod Subfamily Cochliopinae (Prosobranchia: Hydrobiidae)". ''
Malacological Review Malacology is the branch of invertebrate zoology that deals with the study of the Mollusca (mollusks or molluscs), the second-largest phylum of animals in terms of described species after the arthropods. Mollusks include snails and slugs, clams, ...
'', Supplement 5: 1-140. pages 16–18.
Wilke T., Haase M., Hershler R., Liu H.-P., Misof B. & Ponder W. (2013) ''Pushing short DNA fragments to the limit: Phylogenetic relationships of ‘hydrobioid’ gastropods (Caenogastropoda: Rissooidea)''. Molecular Phylogenetics and Evolution, 66: 715-736


External links


Cochliopidae on WoRMS
{{Taxonbar, from=Q969214