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Coastal fish, also called inshore fish or neritic fish, inhabit the sea between the
shoreline A shore or a shoreline is the fringe of land at the edge of a large body of water, such as an ocean, sea, or lake. In physical oceanography, a shore is the wider fringe that is geologically modified by the action of the body of water past ...
and the edge of the continental shelf. Since the continental shelf is usually less than deep, it follows that
pelagic The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or w ...
coastal fish are generally
epipelagic fish Pelagic fish live in the pelagic zone of ocean or lake waters—being neither close to the bottom nor near the shore—in contrast with demersal fish that do live on or near the bottom, and reef fish that are associated with coral reefs ...
, inhabiting the sunlit
epipelagic zone The photic zone, euphotic zone, epipelagic zone, or sunlight zone is the uppermost layer of a body of water that receives sunlight, allowing phytoplankton to perform photosynthesis. It undergoes a series of physical, chemical, and biological pro ...
.Moyle and Cech, 2004, page 585 Coastal fish can be contrasted with ''oceanic fish'' or ''offshore fish'', which inhabit the deep seas beyond the continental shelves. Coastal fish are the most abundant in the world. They can be found in tidal pools,
fjord In physical geography, a fjord or fiord () is a long, narrow inlet with steep sides or cliffs, created by a glacier. Fjords exist on the coasts of Alaska, Antarctica, British Columbia, Chile, Denmark, Germany, Greenland, the Faroe Islands, Icela ...
s and
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environme ...
, near sandy shores and rocky coastlines, around
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. C ...
s and on or above the continental shelf. Coastal fish include
forage fish Forage fish, also called prey fish or bait fish, are small pelagic fish which are preyed on by larger predators for food. Predators include other larger fish, seabirds and marine mammals. Typical ocean forage fish feed near the base of the ...
and the
predator fish Predatory fish are hypercarnivorous fish that actively prey upon other fish or aquatic animals, with examples including shark, billfish, barracuda, pike/muskellunge, walleye, perch and salmon. Some omnivorous fish, such as the red-bellied pira ...
that feed on them. Forage fish thrive in inshore waters where high
productivity Productivity is the efficiency of production of goods or services expressed by some measure. Measurements of productivity are often expressed as a ratio of an aggregate output to a single input or an aggregate input used in a production proce ...
results from
upwelling Upwelling is an oceanographic phenomenon that involves wind-driven motion of dense, cooler, and usually nutrient-rich water from deep water towards the ocean surface. It replaces the warmer and usually nutrient-depleted surface water. The nut ...
and shoreline run off of nutrients. Some are partial residents that spawn in streams, estuaries and bays, but most complete their life cycles in the zone.Moyle and Cech, 2004, page 572


Coastal habitats

Coastal fish are found in the waters above the continental shelves that extend from the continental
shoreline A shore or a shoreline is the fringe of land at the edge of a large body of water, such as an ocean, sea, or lake. In physical oceanography, a shore is the wider fringe that is geologically modified by the action of the body of water past ...
s, and around the
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. C ...
s that surround volcanic islands. The total world shoreline extends for World
The World Factbook, CIA. Retrieved 26 February 2010.
and the continental shelves occupy a total area of 24.3 million km2 (9 376 million sq mi). This is nearly 5% of the world's total area of 510 million km2.


Nearshore fish

Nearshore fish, sometimes called littoral fish, live close to the shore. They are associated with the
intertidal zone The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species ...
, or with
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environme ...
,
lagoon A lagoon is a shallow body of water separated from a larger body of water by a narrow landform, such as reefs, barrier islands, barrier peninsulas, or isthmuses. Lagoons are commonly divided into '' coastal lagoons'' (or ''barrier lagoons' ...
s,
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. C ...
s,
kelp forest Kelp forests are underwater areas with a high density of kelp, which covers a large part of the world's coastlines. Smaller areas of anchored kelp are called kelp beds. They are recognized as one of the most productive and dynamic ecosystems on E ...
s,
seagrass meadow A seagrass meadow or seagrass bed is an underwater ecosystem formed by seagrasses. Seagrasses are marine (saltwater) plants found in shallow coastal waters and in the brackish waters of estuaries. Seagrasses are flowering plants with stems and ...
s, or rocky or sandy bottoms, usually in shallow waters less than about deep.


Intertidal fish

Intertidal fish are fish that move in and out with the
tide Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another. Tide tables can ...
in the
intertidal zone The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species ...
of the seashore, or are found in rock pools or under rocks. The
intertidal zone The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species ...
of rocky shores can contain indentations which trap pools of salty water, called rock pools. Living in these habitats are communities of hardy plant and animal species specially adapted for coping with the volatile environment around them. The plants and animals interact with each other and with the rock pool to form miniature
ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syst ...
s, easily accessible to students and a source of fascination for young children. Plants such as
seaweed Seaweed, or macroalgae, refers to thousands of species of macroscopic, multicellular, marine algae. The term includes some types of ''Rhodophyta'' (red), ''Phaeophyta'' (brown) and ''Chlorophyta'' (green) macroalgae. Seaweed species such as ke ...
s, cnidarians such as sea anemones, arthropods like
barnacle A barnacle is a type of arthropod constituting the subclass Cirripedia in the subphylum Crustacea, and is hence related to crabs and lobsters. Barnacles are exclusively marine, and tend to live in shallow and tidal waters, typically in erosive ...
s, and molluscs such as the common
limpet Limpets are a group of aquatic snails that exhibit a conical gastropod shell, shell shape (patelliform) and a strong, muscular foot. Limpets are members of the class Gastropoda, but are polyphyletic, meaning the various groups called "limpets" ...
and the
common periwinkle The common periwinkle or winkle (''Littorina littorea'') is a species of small edible whelk or sea snail, a marine gastropod mollusc that has gills and an operculum, and is classified within the family Littorinidae, the periwinkles.Reid, Dav ...
can be permanent residents of rock pools. But most rock pool animals, such as crabs,
shrimp Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are refer ...
and fish are just temporary residents, occupying a rock pool only until the next tide takes them to a new location. Some rock pool fish which are temporary residents include the long-spined sea scorpion, the pipefish worm, the rock goby and the common
lumpsucker The Cyclopteridae are a family of marine fishes, commonly known as lumpsuckers or lumpfish, in the order Scorpaeniformes. They are found in the cold waters of the Arctic, North Atlantic, and North Pacific oceans. The greatest number of species a ...
. However some other rock pool fish are territorial in nature, and will stay with the same pool for extended periods. Examples are the common blenny and its near relative the
butterfish Butterfish may refer to: * Stromateidae, found in coastal waters off the Americas, western Africa and in the Indo-Pacific * Sablefish The sablefish (''Anoplopoma fimbria'') is one of two members of the fish family Anoplopomatidae and the onl ...
. * The common
blenny Blenny (from the Greek and , mucus, slime) is a common name for many types of fish, including several families of percomorph marine, brackish, and some freshwater fish sharing similar morphology and behaviour. Six families are considered "true ...
, also known as the shanny, is found in northern temperate waters. They hide under rocks and in crannies in rock pools when the tide is out. They feed on green seaweed and invertebrates such as barnacles. They can crawl on dry land, using their paired fins. About long, they have smooth skin, without scales, and are covered with soft slime. The slime prevents them drying if they are stranded on a shore between tides. So long as their skin stays moist, they can breathe out of water. They are sometimes called "sea frogs" because they bask in the sun on weeds outside the water, and like frogs, jump to safety when disturbed. They can change their colour to match their surroundings. The female lays eggs in crevices or under stones and the male guards them until they hatch. In the winter, when storms can be severe, they move out of their rock pools into the shallows. The common blenny is bold with strong teeth, and will bite humans if it feels threatened. * The rock goby is a small fish, about , found in northern temperate waters. It is coloured black with white blotches, and hides under stones and amongst seaweed. It is a temporary resident of rock pools when the tide is out. The female rock goby lays eggs on the underside of rocks and shells and then leaves them. The male guards the eggs until they hatch. First-year rock gobies often visit rock pools in winter when the older fish have left. File:Australian blenny.jpg, Australian
blenny Blenny (from the Greek and , mucus, slime) is a common name for many types of fish, including several families of percomorph marine, brackish, and some freshwater fish sharing similar morphology and behaviour. Six families are considered "true ...
File:Gobius_paganellus.jpg, Rock goby File:Long_Spined_Scorpion_Fish.jpg, Long-spined sea scorpion File:smooth lumpsucker.jpg, Smooth
lumpsucker The Cyclopteridae are a family of marine fishes, commonly known as lumpsuckers or lumpfish, in the order Scorpaeniformes. They are found in the cold waters of the Arctic, North Atlantic, and North Pacific oceans. The greatest number of species a ...
inflated in a defensive response
* The long-spined sea scorpion, a small stout fish which grows about long, is another temporary resident of rocky pools. They have large black eyes, a large mouth, and four long spines—two on each side on the gill cover—that stick out when the fish is removed from the water. They also have an organ like a finger on each side of their mouths which helps them catch prey. Because of their broad heads, they are also called "bullheads". They have a variety of effective camouflaged colours ranging from shades of browns with cream blotches, to orange and red with white blotches. They can also change their body colour to match their surroundings. They are found around the coasts of Northern Europe in shallow rocky waters hiding amongst seaweed. They are also found in rock pools and sometimes in waters deep. Long-spined sea scorpions lay eggs amongst seaweed or attached to rock crevices. The young hatch after two or three weeks, and go through several development stages before maturing into adults. *
Lumpsucker The Cyclopteridae are a family of marine fishes, commonly known as lumpsuckers or lumpfish, in the order Scorpaeniformes. They are found in the cold waters of the Arctic, North Atlantic, and North Pacific oceans. The greatest number of species a ...
s are found in temperate northern waters. They live on the seafloor, and are temporary residents of rocky pools in late winter and early spring when they spawn. The body of the lumpsucker is scaleless and covered with small lumps. They have a large sucking disc on their underside which they use to cling to surfaces. They are normally a blue to slate-grey colour, and are effectively camouflaged to look like stones. They are portly, nearly spherical, poor swimmers, reaching lengths up to . After the female lumpsucker lays eggs, the male takes over, clamping itself to a rock where it guards the eggs. When they hatch, lumpsuckers look like tiny tadpoles. They remain in shallow water and rock pools, hiding amongst seaweed and rocks, until they grow up.


Estuarine fish

Estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environme ...
are partly enclosed coastal bodies of water with one or more
river A river is a natural flowing watercourse, usually freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the ...
s or streams flowing into them, and with a free connection to the open sea.Pritchard, D. W. (1967) ''What is an estuary: physical viewpoint''. p. 3–5 ''in:'' G. H. Lauf (ed.) ''Estuaries'', A.A.A.S. Publ. No. 83, Washington, D.C. These
brackish water Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
habitats form a transition zone between river environments and ocean environments, and
ecological succession Ecological succession is the process of change in the species structure of an ecological community over time. The time scale can be decades (for example, after a wildfire) or more or less. Bacteria allows for the cycling of nutrients such as car ...
s can form along the way. Estuaries are subject to both marine influences, such as tides, waves, and the influx of saline water; and riverine influences, such as flows of fresh water and sediment. The inflow of both seawater and freshwater provide high levels of nutrients in both the water column and sediment, making estuaries productive natural habitats.McLusky, D.S. and Elliott, M. (2004) "The Estuarine Ecosystem: ecology, threats and management." New York: Oxford University Press Inc. Fishes that spend time in
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environme ...
(or
river mouth A river mouth is where a river flows into a larger body of water, such as another river, a lake/reservoir, a bay/ gulf, a sea, or an ocean. At the river mouth, sediments are often deposited due to the slowing of the current reducing the carryin ...
s) need to be
euryhaline Euryhaline organisms are able to adapt to a wide range of salinities. An example of a euryhaline fish is the molly (''Poecilia sphenops'') which can live in fresh water, brackish water, or salt water. The green crab (''Carcinus maenas'') is an ...
(tolerant to a range of salinities). Estuaries provide an unstable environment for fish, where the salinity changes and the waters are often muddy and turbulent. In warmer climates, estuaries have
mangrove A mangrove is a shrub or tree that grows in coastal saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evolution in several ...
s around their edges. At times there may be only a few different fish species present in an estuary, but seasonal migrants, including eels,
salmonid Salmonidae is a family of ray-finned fish that constitutes the only currently extant family in the order Salmoniformes . It includes salmon (both Atlantic and Pacific species), trout (both ocean-going and landlocked), chars, freshwater whitefis ...
s, and some
forage fish Forage fish, also called prey fish or bait fish, are small pelagic fish which are preyed on by larger predators for food. Predators include other larger fish, seabirds and marine mammals. Typical ocean forage fish feed near the base of the ...
such as
herring Herring are forage fish, mostly belonging to the family of Clupeidae. Herring often move in large schools around fishing banks and near the coast, found particularly in shallow, temperate waters of the North Pacific and North Atlantic Ocea ...
s and
sprat Sprat is the common name applied to a group of forage fish belonging to the genus '' Sprattus'' in the family Clupeidae. The term also is applied to a number of other small sprat-like forage fish ('' Clupeoides'', '' Clupeonella'', '' Corica'' ...
s increase the diversity in the estuary. River estuaries form important staging points during the migration of anadromous and catadromus fish species, such as
salmon Salmon () is the common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of ...
and eels, giving them time to form social groups and to adjust to the changes in salinity. Salmon are anadromous, meaning they live in the sea but ascend rivers to spawn; eels are catadromous, living in rivers and streams, but returning to the sea to breed. Besides the species that migrate through estuaries, there are many other fish that use them as "nursery grounds" for spawning or as places young fish can feed and grow before moving elsewhere. For example,
herring Herring are forage fish, mostly belonging to the family of Clupeidae. Herring often move in large schools around fishing banks and near the coast, found particularly in shallow, temperate waters of the North Pacific and North Atlantic Ocea ...
and
plaice Plaice is a common name for a group of flatfish that comprises four species: the European, American, Alaskan and scale-eye plaice. Commercially, the most important plaice is the European. The principal commercial flatfish in Europe, it is ...
are two commercially important species that use the Thames Estuary for this purpose. Mangrove swamps are associated brackish water habitats. Many, though not all, mangrove swamps fringe estuaries and lagoons where the salinity changes with each tide. Among the most specialised residents of mangrove forests are mudskippers, fish that forage for food on land, and archer fish, perch-like fish that "spit" at insects and other small animals living in the trees, knocking them into the water where they can be eaten. Like estuaries, mangrove swamps are important breeding grounds for many fish, with species such as snappers,
halfbeak Hemiramphidae is a family of fishes that are commonly called halfbeaks, spipe fish or spipefish. They are a geographically widespread and numerically abundant family of epipelagic fish inhabiting warm waters around the world. The halfbeaks a ...
s, and tarpon spawning or maturing among them.


Coral reef fish

In tropical waters, coral reef fish live amongst or in close relation to
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. C ...
s. Coral reefs form complex
ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syst ...
s with tremendous
biodiversity Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic ('' genetic variability''), species ('' species diversity''), and ecosystem ('' ecosystem diversity' ...
. Coral reef fish can be particularly colourful and interesting to watch. Hundreds of species can exist in a small area of a healthy reef, many of them hidden or well camouflaged. Reef fish have developed many ingenious specialisations adapted to survival on the reefs. Coral reefs occupy less than one percent of the surface area of the world oceans, yet they provide a home for 25 percent of all marine fish species. Coral reefs often depend on other habitats in the surrounding area for the supply of nutrients, such as
seagrass meadow A seagrass meadow or seagrass bed is an underwater ecosystem formed by seagrasses. Seagrasses are marine (saltwater) plants found in shallow coastal waters and in the brackish waters of estuaries. Seagrasses are flowering plants with stems and ...
s and mangrove forests. Seagrass and mangroves supply dead plants and animals which are rich in nitrogen and also serve to feed fish and animals from the reef by supplying wood and vegetation. Reefs in turn protect mangroves and seagrass from waves and produce
sediment Sediment is a naturally occurring material that is broken down by processes of weathering and erosion, and is subsequently transported by the action of wind, water, or ice or by the force of gravity acting on the particles. For example, sa ...
for the mangroves and seagrass to root in.
Anthias Anthias are members of the family Serranidae and make up the subfamily Anthiinae. Anthias make up a sizeable portion of the population of pink, orange, and yellow reef fishes seen swarming in most coral reef photography and film. The name An ...
are members of the family
Serranidae The Serranidae are a large family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, in some c ...
and make up the subfamily Anthiinae. They are widespread in tropical waters. They have been called the "quintessential
reef fish Coral reef fish are fish which live amongst or in close relation to coral reefs. Coral reefs form complex ecosystems with tremendous biodiversity. Among the myriad inhabitants, the fish stand out as colourful and interesting to watch. Hundreds ...
", and make up a sizeable portion of the colourful
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% ...
es seen swarming in
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. C ...
photography. Anthias are mostly small, peaceful, beautiful and popular as ornamental fish. They are mainly
zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
feeders. Anthias shoal and school in large numbers, operating more intimate "harems" within the schools. These harems contain a dominant and colourful male, between 2 and 12 females — who operate a hierarchy among themselves — and one or two "subdominant" males, often less brightly coloured and non-territorial. Within the swarm of females, territorial males perform acrobatic U-swim displays and vigorously defend an area of the reef and its associated harem. Anthias are
protogynous Sequential hermaphroditism (called dichogamy in botany) is a type of hermaphroditism that occurs in many fish, gastropods, and plants. Sequential hermaphroditism occurs when the individual changes its sex at some point in its life. In particular, ...
hermaphrodite In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have ...
s. All anthias are born female; if a dominant male perishes, the largest female of the group will often change into a male to take its place. This may lead to squabbling between the next largest male and the transforming female, whose hormones are now surging with testosterone. This can turn quite vicious in the limited confines of captivity. Butterflyfish are group of about 120 species belonging to the family Chaetodontidaeof Perchiformes. They include bannerfish and
coralfish The butterflyfish are a group of conspicuous tropical marine fish of the family Chaetodontidae; the bannerfish and coralfish are also included in this group. The approximately 129 species in 12 genera are found mostly on the reefs of the Atla ...
. They are widespread on coral reefs. Butterflyfish are mostly between in length. The largest species, the lined butterflyfish and saddle butterflyfish, grow to . Many species are brightly coloured and strikingly patterned, though other species are dull in colour. Many have eyespots on their flanks and dark bands across their eyes, not unlike the patterns seen on
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group comprises ...
wings. Their deep, laterally narrow bodies are easily noticed through the profusion of reef life. The conspicuous colouration of butterflyfish may be intended for interspecies communication. Butterflyfish have uninterrupted
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...
s with tail fins that may be rounded or truncated, but are never forked. Generally diurnal and frequenting waters of less than (though some species descend to ), butterflyfish stick to particular home ranges. The
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and secre ...
livores are especially territorial, forming mated pairs and staking claim to a specific coral head. Contrastingly, the
zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
feeders form large conspecific groups. By night butterflyfish hide in reef crevices and exhibit markedly different colouration. Their colouration also makes butterflyfish popular aquarium fish. However, most species feed on
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and secre ...
polyps and sea anemones, which can result in problems for the hobby aquarists.
Clownfish Clownfish or anemonefish are fishes from the subfamily Amphiprioninae in the family Pomacentridae. Thirty species of clownfish are recognized: one in the genus '' Premnas'', while the remaining are in the genus '' Amphiprion''. In the wild, ...
,
anemonefish Clownfish or anemonefish are fishes from the subfamily Amphiprioninae in the family Pomacentridae. Thirty species of clownfish are recognized: one in the genus ''Premnas'', while the remaining are in the genus ''Amphiprion''. In the wild, the ...
and
damselfish Damselfish are those within the subfamilies Abudefdufinae, Chrominae, Lepidozyginae, Pomacentrinae, and Stegastenae within the family Pomacentridae. Most species within this group are relatively small, with the largest species being about 30 ...
are among about 360 species classified in the family
Pomacentridae Pomacentridae is a family of ray-finned fish, comprising the damselfishes and clownfishes. This family were formerly placed in the order Perciformes but are now regarded as being ''incertae sedis'' in the subseries Ovalentaria in the clade ...
. Most Pomacentrids are associated with coral reefs in the Indo-West Pacific, with a few species occurring in temperate waters. Some species are native to freshwater or brackish
estuarine An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environmen ...
environments. Most live in shallow water, from , although some species are found below . Most species are specialists, living in specific parts of the reef, such as sandy lagoons, steep reef slopes, or areas exposed to strong wave action. In general, the coral is used as shelter, and many species can only survive in its presence. The bottom-dwelling species are territorial, occupying and defending a portion of the reef, often centred around an area of shelter. By keeping away other species of fish, some pomacentrids encourage the growth of thick mats of algae within their territories, leading to the common name farmerfish. Different species display a wide range of colours, although some are relatively drab. Pomacentrids are omnivorous or herbivorous, feeding off algae,
plankton Plankton are the diverse collection of organisms found in water (or air) that are unable to propel themselves against a current (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they provide a cr ...
, and small bottom-dwelling
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean gro ...
s. A small number eat coral. Goatfishes are a family Mullidae of about 55 species of perciform fishes, associated worldwide with tropical
reef A reef is a ridge or shoal of rock, coral or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic component, abiotic processes—deposition (geology), deposition of ...
s. They are typically about 20 cm long, though the
dash-and-dot goatfish The dash-and-dot goatfish (''Parupeneus barberinus'') is a species of goatfish native to the Indian and Pacific oceans. Description The dash-and-dot goatfish has the twin chin barbels typical of goatfishes. The body color is white with a ...
, grows to 55 cm. Goatfish are tireless
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
feeders, possess a pair of long chemosensory
barbels In fish anatomy and turtle anatomy, a barbel is a slender, whiskerlike sensory organ near the mouth. Fish that have barbels include the catfish, the carp, the goatfish, the hagfish, the sturgeon, the zebrafish, the black dragonfish and s ...
("whiskers") protruding from their chins resembling a goat's beard. They use these to rifle through the sediments in search of a meal. Like goats, they seek anything edible;
worm Worms are many different distantly related bilateral animals that typically have a long cylindrical tube-like body, no limbs, and no eyes (though not always). Worms vary in size from microscopic to over in length for marine polychaete worm ...
s,
crustacea Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean gro ...
ns, molluscs and other small
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s are staples. Many species of goatfish are conspicuously coloured and have the ability to change their colouration depending on their current activity. By day, many form large inactive (non-feeding) schools: these aggregates may contain both conspecifics and heterospecifics. For example, the yellowfin goatfish school with blue-striped snappers. When they do that, the yellowfins changes its colouration to match that of the snapper. By night the schools disperse and individual goatfish head their separate ways to loot the sands. The diurnal goldsaddle goatfish changes from a lemon-yellow to a pale cream when feeding. Other nocturnal feeders will shadow the active goatfish, waiting patiently for overlooked morsels. Goatfish stay within the shallows, going no deeper than about 110 metres. Most species do not tolerate
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
water, so they do not enter
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environme ...
or the mouths of
river A river is a natural flowing watercourse, usually freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the ...
s.


Other nearshore fish

Other nearshore or shallow water fish live near the shore in depths of less than 10 metres. They occupy the areas over sandy or rocky bottoms, and can be associated with
seagrass meadow A seagrass meadow or seagrass bed is an underwater ecosystem formed by seagrasses. Seagrasses are marine (saltwater) plants found in shallow coastal waters and in the brackish waters of estuaries. Seagrasses are flowering plants with stems and ...
s and
kelp forest Kelp forests are underwater areas with a high density of kelp, which covers a large part of the world's coastlines. Smaller areas of anchored kelp are called kelp beds. They are recognized as one of the most productive and dynamic ecosystems on E ...
s. They can be divided into
demersal fish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They o ...
and
pelagic fish Pelagic fish live in the pelagic zone of ocean or lake waters—being neither close to the bottom nor near the shore—in contrast with demersal fish that do live on or near the bottom, and reef fish that are associated with coral re ...
. Demersal fish live on or near the sea floor, while pelagic fish live in the
water column A water column is a conceptual column of water from the surface of a sea, river or lake to the bottom sediment.Munson, B.H., Axler, R., Hagley C., Host G., Merrick G., Richards C. (2004).Glossary. ''Water on the Web''. University of Minnesota-D ...
away the sea floor. Examples of such shallow water demersal fish, found in both tropical and temperate waters around the world, are triplefins, seahorses, wrasse and flounder. As demersal fish, all these fish spend most of their time on or near the sea floor. *
Flatfish A flatfish is a member of the ray-finned demersal fish order Pleuronectiformes, also called the Heterosomata, sometimes classified as a suborder of Perciformes. In many species, both eyes lie on one side of the head, one or the other migratin ...
are superbly adapted
groundfish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They o ...
, found on muddy and sandy sea floors. In many species both eyes lie on one side of the head, one or the other migrating through and around the head during development. Some species face their "left" side upward, some face their "right" side upward, and others face either side upward. Some flatfish can camouflage themselves on the ocean floor. *
Wrasse The wrasses are a family, Labridae, of marine fish, many of which are brightly colored. The family is large and diverse, with over 600 species in 81 genera, which are divided into 9 subgroups or tribes. They are typically small, most of them ...
are a large family of mainly small fish, usually less than long. Most wrasse are loners that prefer habitats such as coral reefs and rocky shores. They live close to the substrate, eating small invertebrates and almost anything else that lurks on the bottom. Many are brightly coloured. They have thick lips and use their sharp teeth to pick small creatures off the rocks. Many smaller wrasses follow the feeding trails of larger fish, picking up invertebrates disturbed by their passing. *
Triplefin Threefin or triplefin blennies are blenniiforms, small percomorph marine fish of the family Tripterygiidae. Found in tropical and temperate waters of the Atlantic, Pacific and Indian Oceans, the family contains about 150 species in 30 genera ...
s are a
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of fish. They are usually found around
coral reefs A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. Co ...
and rocks, usually in shallow, clear sunlit waters such as lagoons and seaward reefs. Triplefins have three
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...
s (hence the name). They are small fish, usually less than six cm long. Brightly coloured, often for reasons of camouflage, they are nervous and retreat to rock crevices at any perceived threat. *
Seahorse A seahorse (also written ''sea-horse'' and ''sea horse'') is any of 46 species of small marine fish in the genus ''Hippocampus''. "Hippocampus" comes from the Ancient Greek (), itself from () meaning "horse" and () meaning "sea monster" or " ...
s are a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of fish. They prefer sheltered harbours, estuaries and other shallow coastal waters, where they hunt tiny crustaceans. They bob around in sheltered areas such as
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. C ...
s, mangrove stands and
seagrass meadow A seagrass meadow or seagrass bed is an underwater ecosystem formed by seagrasses. Seagrasses are marine (saltwater) plants found in shallow coastal waters and in the brackish waters of estuaries. Seagrasses are flowering plants with stems and ...
s and
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environme ...
. They are camouflaged with murky patterns that blend into kelp and sea grass backgrounds. During social moments or in unusual surroundings, seahorses can turn on bright colours. File:Hippocampus.jpg, Seahorse File:Pseudopleuronectes americanus.jpg, Flatfish File:Cleaning station.JPG, Cleaner wrasses working on a dragon wrasse File:Common triplefin.png, Common triplefin Examples of shallow water pelagic fish, found in both tropical and temperate waters around the world, are grey mullet, sprats and garfish. As pelagic fish, all these fish spend most of their time living in the
water column A water column is a conceptual column of water from the surface of a sea, river or lake to the bottom sediment.Munson, B.H., Axler, R., Hagley C., Host G., Merrick G., Richards C. (2004).Glossary. ''Water on the Web''. University of Minnesota-D ...
away the sea floor. * The grey mullet are medium size fish, typically about long. They are often caught with
seine net Seine fishing (or seine-haul fishing; ) is a method of fishing that employs a surrounding net, called a seine, that hangs vertically in the water with its bottom edge held down by weights and its top edge buoyed by floats. Seine nets can be de ...
s. * The
garfish The garfish (''Belone belone''), also known as the garpike or sea needle, is a pelagic, oceanodromous needlefish found in brackish and marine waters of the Atlantic Ocean and the Mediterranean, Caribbean, Black, and Baltic Seas. Description The ...
is a long, slender fish, looking like a spear, which feeds on seagrass fragments, shrimps and crab larvae. In turn it is preyed on by larger fish and, since it is often near the surface,
cormorant Phalacrocoracidae is a family of approximately 40 species of aquatic birds commonly known as cormorants and shags. Several different classifications of the family have been proposed, but in 2021 the IOC adopted a consensus taxonomy of seven ge ...
s and
gannet Gannets are seabirds comprising the genus ''Morus'' in the family Sulidae, closely related to boobies. Gannets are large white birds with yellowish heads; black-tipped wings; and long bills. Northern gannets are the largest seabirds in the ...
s. File:Mucep u0.gif, Grey mullet File:Belone belone1.jpg,
Garfish The garfish (''Belone belone''), also known as the garpike or sea needle, is a pelagic, oceanodromous needlefish found in brackish and marine waters of the Atlantic Ocean and the Mediterranean, Caribbean, Black, and Baltic Seas. Description The ...


Coastal pelagic fish


Plankton feeding

At the base of
food chain A food chain is a linear network of links in a food web starting from producer organisms (such as grass or algae which produce their own food via photosynthesis) and ending at an apex predator species (like grizzly bears or killer whales), d ...
s are the
primary producer Primary or primaries may refer to: Arts, entertainment, and media Music Groups and labels * Primary (band), from Australia * Primary (musician), hip hop musician and record producer from South Korea * Primary Music, Israeli record label Work ...
s. In the ocean these primary producers are mainly a type of
plankton Plankton are the diverse collection of organisms found in water (or air) that are unable to propel themselves against a current (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they provide a cr ...
, microscopic
phytoplankton Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater ecosystems. The name comes from the Greek words (), meaning 'plant', and (), meaning 'wanderer' or 'drifter'. P ...
which drift in the
water column A water column is a conceptual column of water from the surface of a sea, river or lake to the bottom sediment.Munson, B.H., Axler, R., Hagley C., Host G., Merrick G., Richards C. (2004).Glossary. ''Water on the Web''. University of Minnesota-D ...
. Phytoplankton need sunlight for
photosynthesis Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored i ...
to power
carbon fixation Biological carbon fixation or сarbon assimilation is the process by which inorganic carbon (particularly in the form of carbon dioxide) is converted to organic compounds by living organisms. The compounds are then used to store energy and as ...
, so they are mainly located in sunlit surface waters. Phytoplankton also need and rapidly use nutrients in the water column.Anderson, G (2003
Coral Reef Formation
''Marine Science''.
The phytoplankton are eaten by
zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
, which in turn are eaten by predatory zooplankton.
Filter feeder Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feedin ...
s then eat the plankton and larger
predatory fish Predatory fish are hypercarnivorous fish that actively prey upon other fish or aquatic animals, with examples including shark, billfish, barracuda, pike/muskellunge, walleye, perch and salmon. Some omnivorous fish, such as the red-bellied pir ...
eat the filter feeders (see diagram at the right).Mann, K.H., Lazier, J.R.N. (2006) ''Dynamics of Marine Ecosystems: Biological-Physical Interactions in the Oceans''. Oxford: Blackwell Publishing Ltd. Most filter-feeding pelagic fish found in coastal waters are small, silvery
forage fish Forage fish, also called prey fish or bait fish, are small pelagic fish which are preyed on by larger predators for food. Predators include other larger fish, seabirds and marine mammals. Typical ocean forage fish feed near the base of the ...
. Forage fish include fishes of the family
Clupeidae Clupeidae is a family of ray-finned fishes, comprising, for instance, the herrings, shads, sardines, hilsa, and menhadens. The clupeoids include many of the most important food fishes in the world, and are also commonly caught for production ...
(
herring Herring are forage fish, mostly belonging to the family of Clupeidae. Herring often move in large schools around fishing banks and near the coast, found particularly in shallow, temperate waters of the North Pacific and North Atlantic Ocea ...
s,
shad The Alosinae, or the shads,Alosinae
sardine "Sardine" and "pilchard" are common names for various species of small, oily forage fish in the herring family Clupeidae. The term "sardine" was first used in English during the early 15th century, a folk etymology says it comes from the ...
s and pilchards, hilsa,
menhaden Menhaden, also known as mossbunker and bunker and "the most important fish in the sea", are forage fish of the genera ''Brevoortia'' and ''Ethmidium'', two genera of marine fish in the family Clupeidae. ''Menhaden'' is a blend of ''poghaden'' ...
and
sprat Sprat is the common name applied to a group of forage fish belonging to the genus '' Sprattus'' in the family Clupeidae. The term also is applied to a number of other small sprat-like forage fish ('' Clupeoides'', '' Clupeonella'', '' Corica'' ...
s), as well as
anchovies An anchovy is a small, common forage fish of the family Engraulidae. Most species are found in marine waters, but several will enter brackish water, and some in South America are restricted to fresh water. More than 140 species are placed in 1 ...
,
capelin The capelin or caplin (''Mallotus villosus'') is a small forage fish of the smelt family found in the North Atlantic, North Pacific and Arctic oceans. In summer, it grazes on dense swarms of plankton at the edge of the ice shelf. Larger capel ...
and
halfbeak Hemiramphidae is a family of fishes that are commonly called halfbeaks, spipe fish or spipefish. They are a geographically widespread and numerically abundant family of epipelagic fish inhabiting warm waters around the world. The halfbeaks a ...
s. They use schooling strategies to avoid predators, and different schools of forage fish often associate with each other in open coastal waters. Forage fish feed near the base of the food chain on
plankton Plankton are the diverse collection of organisms found in water (or air) that are unable to propel themselves against a current (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they provide a cr ...
and fry (recently hatched fish), often by
filter feeding Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feedi ...
. In turn, they are preyed on by larger predatory fish,
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same envir ...
s and
marine mammal Marine mammals are aquatic mammals that rely on the ocean and other marine ecosystems for their existence. They include animals such as Pinniped, seals, Cetacea, whales, Sirenia, manatees, sea otters and polar bears. They are an informal group, ...
s. Worldwide, there are five major coastal currents associated with upwelling areas: the
Canary Current The Canary Current is a wind-driven surface current that is part of the North Atlantic Gyre. This eastern boundary current branches south from the North Atlantic Current and flows southwest about as far as Senegal where it turns west and later j ...
(off
Northwest Africa The Maghreb (; ar, الْمَغْرِب, al-Maghrib, lit=the west), also known as the Arab Maghreb ( ar, المغرب العربي) and Northwest Africa, is the western part of North Africa and the Arab world. The region includes Algeria, ...
), the Benguela Current (off
southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number ...
), the
California Current The California Current is a cold water Pacific Ocean current that moves southward along the western coast of North America, beginning off southern British Columbia and ending off southern Baja California Sur. It is considered an Eastern boundar ...
(off
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the ...
and
Oregon Oregon () is a state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington, while the Snake River delineates much of its eastern boundary with Idah ...
), the
Humboldt Current The Humboldt Current, also called the Peru Current, is a cold, low-salinity ocean current that flows north along the western coast of South America.Montecino, Vivian, and Carina B. Lange. "The Humboldt Current System: Ecosystem components and p ...
(off
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
and
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the eas ...
), and the
Somali Current The Somali Current is a warm ocean boundary current that runs along the coast of Somalia and Oman in the Western Indian Ocean and is analogous to the Gulf Stream in the Atlantic Ocean. This current is heavily influenced by the monsoons and is th ...
(off
Western India Western India is a loosely defined region of India consisting of its western part. The Ministry of Home Affairs in its Western Zonal Council Administrative division includes the states of Goa, Gujarat, and Maharashtra along with the Uni ...
). All of these currents support major fisheries. Many forage fish are important commercial species, and the schools can be targeted by spotter planes. The fish are caught by purse seiners—fishing boats that use nets to enclose the fish—and can be overfished. File:Anchovy closeup.jpg, Schooling
anchovies An anchovy is a small, common forage fish of the family Engraulidae. Most species are found in marine waters, but several will enter brackish water, and some in South America are restricted to fresh water. More than 140 species are placed in 1 ...
File:Clupea sprattus.jpg,
European sprat The European sprat (''Sprattus sprattus''), also known as bristling, brisling, garvie, garvock, Russian sardine, russlet, skipper or whitebait, is a species of small marine fish in the herring family Clupeidae. Found in European waters, it has si ...
File:Herring2.jpg,
Atlantic herring Atlantic herring (''Clupea harengus'') is a herring in the family Clupeidae. It is one of the most abundant fish species in the world. Atlantic herrings can be found on both sides of the Atlantic Ocean, congregating in large schools. They can ...
File:Heringsschwarm.gif, Underwater video loop of a school of herrings migrating to their spawning grounds in the
Baltic Sea The Baltic Sea is an arm of the Atlantic Ocean that is enclosed by Denmark, Estonia, Finland, Germany, Latvia, Lithuania, Poland, Russia, Sweden and the North and Central European Plain. The sea stretches from 53°N to 66°N latitude and fr ...


Predatory

Predatory pelagic fishes found on continental shelves worldwide in both tropical and temperate waters include porgies,
barracuda A barracuda, or cuda for short, is a large, predatory, ray-finned fish known for its fearsome appearance and ferocious behaviour. The barracuda is a saltwater fish of the genus ''Sphyraena'', the only genus in the family Sphyraenidae, which was ...
, amberjacks and cutlassfishes. They tend to be larger fish, and are carnivorous, feeding on the smaller, silvery
forage fish Forage fish, also called prey fish or bait fish, are small pelagic fish which are preyed on by larger predators for food. Predators include other larger fish, seabirds and marine mammals. Typical ocean forage fish feed near the base of the ...
that eat plankton (see section above). Some species also feed on crabs and other invertebrates, foraged from the sea floor. *
Mackerel Mackerel is a common name applied to a number of different species of pelagic fish, mostly from the family Scombridae. They are found in both temperate and tropical seas, mostly living along the coast or offshore in the oceanic environment. ...
scom* * Porgies sometimes called sea breams, are any of about 100 species belonging to the family Sparidae. Porgies usually have high backs and a single dorsal fin, like snapper or grunt fishes (grunts are named for the sound they make grinding their teeth). They are bottom feeding pelagic fishes, with small mouths equipped with strong teeth adapted for handling small fishes and invertebrates with hard shells. Most do not exceed a size of about , but some may grow to four times that length. They often school, and will migrate between reefs. Larger fish enter estuaries and harbours. *
Barracuda A barracuda, or cuda for short, is a large, predatory, ray-finned fish known for its fearsome appearance and ferocious behaviour. The barracuda is a saltwater fish of the genus ''Sphyraena'', the only genus in the family Sphyraenidae, which was ...
have long slender bodies typically about long. They have a wicked set of teeth and are ferocious predators. They feed on
crustaceans Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean gr ...
,
cephalopods A cephalopod is any member of the molluscan Taxonomic rank, class Cephalopoda (Greek language, Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral symm ...
and small fish like
anchovy An anchovy is a small, common forage fish of the family Engraulidae. Most species are found in marine waters, but several will enter brackish water, and some in South America are restricted to fresh water. More than 140 species are placed in 1 ...
and pilchard. Barracouta often hunt in schools near the bottom or midwater, and sometimes even near the surface at night. File:Calamus bajonado.jpg, The jolthead porgy is a porgi gruntPollnac, R.B. 1998 Aspects of the human ecology of the coral reefs of Discovery Bay. p. 141-199. In R.B. Pollnac (ed.) Rapid assessment of management parameters for coral reefs. Coastal Resources Center, University of Rhode Island. CMR 2205, ICLARM 1445. File:Great Barracuda off the Netherland Antilles.jpg, Great barracuda and jacks File:Seriolalalandii.jpg, The
yellowtail amberjack The yellowtail amberjack, yellowtail kingfish, hiramasa or great amberjack (''Seriola lalandi'') is a large fish found in the Southern Ocean. Although previously thought to be found in all oceans and seas, recent genetic analysis restricts ''S. ...
, pound for pound, is one of the hardest fighting fish in the ocean. File:Atlantic_cutlassfish.jpg, The
largehead hairtail The largehead hairtail (''Trichiurus lepturus'') or beltfish is a member of the cutlassfish family, Trichiuridae. This common to abundant species is found in tropical and temperate oceans throughout the world. The taxonomy is not fully resolved ...
is a cutlassfish
* Cutlassfishes are a group of about 40 species belonging to the family Trichiuridae. They are ocean fish which regularly stray into coastal waters around the world. Fish of this family are long, slender, and generally steely blue or silver in colour, giving rise to their name. They have reduced or absent
pelvic The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The ...
and
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as s ...
s, giving them an eel-like appearance, and large fang-like teeth. * Jacks, amberjacks, pompanos, horse mackerel, scads, leatherjackets and
trevally The Carangidae are a family of ray-finned fish which includes the jacks, pompanos, jack mackerels, runners, and scads. It is the largest of the six families included within the order Carangiformes. Some authorities classify it as the only family ...
are fish of the family ''
Carangidae The Carangidae are a family of ray-finned fish which includes the jacks, pompanos, jack mackerels, runners, and scads. It is the largest of the six families included within the order Carangiformes. Some authorities classify it as the only family ...
''. Found in most coastal waters, they are fast predatory fishes that hunt in the waters above
reef A reef is a ridge or shoal of rock, coral or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic component, abiotic processes—deposition (geology), deposition of ...
s and in the open sea; some dig in the sea floor for invertebrates (some can also filter feed, such as the white trevally). The largest fish in the family, the
giant trevally The giant trevally (''Caranx ignobilis''), also known as the lowly trevally, barrier trevally, ronin jack, giant kingfish or ''ulua'', is a species of large marine fish classified in the jack family, Carangidae. The giant trevally is distributed ...
, grows up to 1.7 m in length; most fish in the family reach a maximum length of 25–100 cm. The family contains many important commercial and game fish, notably the Pacific jack mackerel and the other jack mackerels in the genus '' Trachurus''. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
of this genus is the Atlantic horse mackerel. Jack mackerels are an important inshore commercial species. ** Amberjacks are a group of nine species belonging to the genus ''Seriola'' within the family Carangidae. Mainly open water fish, they can follow small
forage fish Forage fish, also called prey fish or bait fish, are small pelagic fish which are preyed on by larger predators for food. Predators include other larger fish, seabirds and marine mammals. Typical ocean forage fish feed near the base of the ...
into estuaries and enclosed waters, where they will also hunt for crustaceans. Amberjacks are fast swimming and aggressive predators that often hunt in schools around offshore reefs. The
yellowtail amberjack The yellowtail amberjack, yellowtail kingfish, hiramasa or great amberjack (''Seriola lalandi'') is a large fish found in the Southern Ocean. Although previously thought to be found in all oceans and seas, recent genetic analysis restricts ''S. ...
can reach 1.8 m in length and weigh 60 kilograms.


Demersal pelagic fish

Fish that live on or in close association with the sea floor are called
demersal fish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They o ...
. This section discusses the coastal demersal fish that live on the continental shelf, but are living further from the coast and in deeper water than the nearshore fish discussed above. Demersal fish are white fish. Unlike
oily fish Oily fish are fish species with oil (fats) in soft tissues and in the coelomic cavity around the gut. Their fillets may contain up to 30% oil, although this figure varies both within and between species. Examples of oily fish include smal ...
, white fish contain
oils An oil is any nonpolar chemical substance that is composed primarily of hydrocarbons and is hydrophobic (does not mix with water) & lipophilic (mixes with other oils). Oils are usually flammable and surface active. Most oils are unsaturat ...
only in their
liver The liver is a major organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the synthesis of proteins and biochemicals necessary for digestion and growth. In humans, it ...
, rather than in the
gut Gut or guts may refer to: Anatomy * Abdomen or belly, the region of a vertebrate between the chest and pelvis * Abdominal obesity or "a gut", a large deposit of belly fat * Gastrointestinal tract or gut, the system of digestive organs * Ins ...
, and can therefore be gutted as soon as they are caught, on board the ship. White fish has dry and white flesh. They can be divided into benthopelagic fish (mostly "round" fish) which live ''near'' the sea bed, such as cod, and benthic fish (
flatfish A flatfish is a member of the ray-finned demersal fish order Pleuronectiformes, also called the Heterosomata, sometimes classified as a suborder of Perciformes. In many species, both eyes lie on one side of the head, one or the other migratin ...
) such as
plaice Plaice is a common name for a group of flatfish that comprises four species: the European, American, Alaskan and scale-eye plaice. Commercially, the most important plaice is the European. The principal commercial flatfish in Europe, it is ...
which live ''on'' the sea bed. Benthic fish tend to be "flat", so they can lie on the bottom. * Cod-like fishes are a number of round benthopelagic species belonging to the order Gadiformes, such as
Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the "Old World" of Africa, Europe an ...
and Pacific cod, morid cod,
haddock The haddock (''Melanogrammus aeglefinus'') is a saltwater ray-finned fish from the family Gadidae, the true cods. It is the only species in the monotypic genus ''Melanogrammus''. It is found in the North Atlantic Ocean and associated seas w ...
and
pollock Pollock or pollack (pronounced ) is the common name used for either of the two species of North Atlantic marine fish in the genus ''Pollachius''. '' Pollachius pollachius'' is referred to as pollock in North America, Ireland and the United Ki ...
, including the highly commercial
Alaska pollock The Alaska pollock or walleye pollock (''Gadus chalcogrammus'') is a marine fish species of the cod genus '' Gadus'' and family Gadidae. It is a semi- pelagic schooling fish widely distributed in the North Pacific, with largest concentratio ...
. Cod-like fishes are often found in large schools over sandy or muddy bottoms. They have a barbel (fleshy filament) on their lower jaw which they use to detect prey buried in the sand or mud. Some migrate to warm water in winter to spawn. * John Dory are fishes of the genus ''Zeus''. They have a widespread distribution and are typically found near the seabed in depths from . The John Dory grows to a maximum length of . Although it is a benthopelagic fish, its body is flat and it can hardly be seen from the front because it is so thin. It is a poor swimmer with long spines on the dorsal fin. It has a large dark eyespot on the side of its body which is used to confuse prey, which are scooped up in its big mouth. Large eyes at the front of the head provide it with bifocal vision and depth perception, which are important for predators. The John Dory usually gets its food by stalking it then shooting out a tube in its mouth to capture its prey. It eats forage fish, and occasionally
squid True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fitting ...
and
cuttlefish Cuttlefish or cuttles are marine molluscs of the order Sepiida. They belong to the class Cephalopoda which also includes squid, octopuses, and nautiluses. Cuttlefish have a unique internal shell, the cuttlebone, which is used for control ...
. In turn, they are preyed on by sharks, like the dusky shark, and other large bony fish. They are normally solitary. *
Turbot The turbot (''Scophthalmus maximus'') is a relatively large species of flatfish in the family Scophthalmidae. It is a demersal fish native to marine or brackish waters of the Northeast Atlantic, Baltic Sea and the Mediterranean Sea. It is an ...
and brill are benthic flatfish, resembling flounder and sole, but found in deeper offshore waters on the continental shelf. They are brownish-green, with dark blotches on the turbot and mottling on the brill. They are fished by coastal trawlers. File:Boreogadus saida.jpg, The polar cod is found further north than any other fish species. It frequents river mouths and feeds on
plankton Plankton are the diverse collection of organisms found in water (or air) that are unable to propel themselves against a current (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they provide a cr ...
and
krill Krill are small crustaceans of the order Euphausiacea, and are found in all the world's oceans. The name "krill" comes from the Norwegian word ', meaning "small fry of fish", which is also often attributed to species of fish. Krill are consid ...
. It is preyed on by narwhals, belugas,
ringed seal The ringed seal (''Pusa hispida'') is an earless seal inhabiting the Arctic and sub-Arctic regions. The ringed seal is a relatively small seal, rarely greater than 1.5 m in length, with a distinctive patterning of dark spots surrounded by light ...
s and
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same envir ...
s. File:MacGillivray, William John Dory.jpg, The John Dory is so thin it can hardly be seen from the front. The large eyespot on the side of its body confuses its prey. File:Psetta_maxima_Luc_Viatour.jpg, The
turbot The turbot (''Scophthalmus maximus'') is a relatively large species of flatfish in the family Scophthalmidae. It is a demersal fish native to marine or brackish waters of the Northeast Atlantic, Baltic Sea and the Mediterranean Sea. It is an ...
is a large left-eyed
flatfish A flatfish is a member of the ray-finned demersal fish order Pleuronectiformes, also called the Heterosomata, sometimes classified as a suborder of Perciformes. In many species, both eyes lie on one side of the head, one or the other migratin ...
usually found not too far from shore in sandy shallow waters. It is a prized
food fish Food is any substance consumed by an organism for nutritional support. Food is usually of plant, animal, or fungal origin, and contains essential nutrients, such as carbohydrates, fats, proteins, vitamins, or minerals. The substance is inges ...
. File:Helidonichthys_spinosus.jpg, The red gurnard is a mail-cheeked fish found to depths of around 180 metres. They often grunt when captured as air is expelled from its air bladder.
* Mail-cheeked fishes belong to a group of about 30 species in the order Scorpaeniformes. Mail-cheeked fishes are named after a plate of bone that runs across each cheek. They are widespread in all the oceans of the world. Mail-cheeked fishes are
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other ...
, mostly feeding on crustaceans, such as crabs and shrimp, and on smaller fish. Most species live on the sea bottom in relatively shallow waters, although species are known from mid and deep water, from the mid-water, and even from fresh water. They typically have spiny heads, and rounded pectoral and
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as s ...
s. Most species are less than in length, but the full size range of the order varies from the velvetfishes, which can be just long as adults, to the
Lingcod The lingcod or ling cod (''Ophiodon elongatus''), also known as the buffalo cod or cultus cod, is a fish of the greenling family Hexagrammidae. It is the only extant member of the genus ''Ophiodon. ''A slightly larger, extinct species, ''Ophio ...
, which can reach in length. ** Red gurnard are mail-cheeked fish. They use their large pectoral fins to rest on the bottom and to detect food. Stargazers are about 50 species of fishes, belonging to the family Uranoscopidae, and found worldwide in shallow waters. Stargazers are
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a ...
; they have two large
poison Poison is a chemical substance that has a detrimental effect to life. The term is used in a wide range of scientific fields and industries, where it is often specifically defined. It may also be applied colloquially or figuratively, with a broa ...
spines situated behind the opercle and above the
pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
s. They can also deliver
electric shock Electrical injury is a physiological reaction caused by electric current passing through the body. The injury depends on the density of the current, tissue resistance and duration of contact. Very small currents may be imperceptible or produce a ...
s. They are
ambush predator Ambush predators or sit-and-wait predators are carnivorous animals that capture or trap prey via stealth, luring or by (typically instinctive) strategies utilizing an element of surprise. Unlike pursuit predators, who chase to capture prey u ...
s with eyes on top of their heads (thus the name). Stargazers also have a large upward-facing mouth in a large head. They bury themselves in sand with only their eyes showing, and leap upwards to ambush fish and invertebrates overhead. Some species have a worm-shaped lure growing out of the floor of the mouth, which they wiggle to attract prey's attention. Lengths range from 18 cm up to 90 cm, for the giant stargazer ''Kathetostoma giganteum''. Stargazers are a delicacy in some cultures. The venom is destroyed when it is cooked, and stargazers are sold in some
fish market A fish market is a marketplace for selling fish and fish products. It can be dedicated to wholesale trade between fishermen and fish merchants, or to the sale of seafood to individual consumers, or to both. Retail fish markets, a type of wet m ...
s with their electric organ removed. They have been called "the meanest things in creation" and the "worst pet on earth".Grady, Denis
Venom Runs Thick in Fish Families, Researchers Learn
''
The New York Times ''The New York Times'' (''the Times'', ''NYT'', or the Gray Lady) is a daily newspaper based in New York City with a worldwide readership reported in 2020 to comprise a declining 840,000 paid print subscribers, and a growing 6 million paid ...
'' 22 August 2006.
* Sandperches are a family, Pinguipedidae, containing 63 species of fishes in the order
Perciformes Perciformes (), also called the Percomorpha or Acanthopteri, is an order or superorder of ray-finned fish. If considered a single order, they are the most numerous order of vertebrates, containing about 41% of all bony fish. Perciformes means ...
. They are
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
carnivores, feeding on small fish and invertebrates. Examples are the redbanded weever, yellow weaver and blue cod. They are often caught in pots like crayfish. * Medusa fishes are a family Centrolophidae of 31 species of
perciform Perciformes (), also called the Percomorpha or Acanthopteri, is an order or superorder of ray-finned fish. If considered a single order, they are the most numerous order of vertebrates, containing about 41% of all bony fish. Perciformes means " ...
fishes. They are found in temperate and tropical waters throughout the world, usually feeding on fish, crustaceans and small squid near rough sea floors on continental shelf and slope. Examples are barrelfish, southern driftfish, imperial blackfish, the
Japanese Japanese may refer to: * Something from or related to Japan, an island country in East Asia * Japanese language, spoken mainly in Japan * Japanese people, the ethnic group that identifies with Japan through ancestry or culture ** Japanese diaspor ...
and pelagic butterfish, the
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 List of islands of New Zealand, smaller islands. It is the ...
and
Tasmanian ruffe The Tasmanian ruffe, ''Tubbia tasmanica'', is a medusafish of the family Centrolophidae, found in temperate waters in the Indian and southwest Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic division ...
, and the
common Common may refer to: Places * Common, a townland in County Tyrone, Northern Ireland * Boston Common, a central public park in Boston, Massachusetts * Cambridge Common, common land area in Cambridge, Massachusetts * Clapham Common, originally ...
,
silver Silver is a chemical element with the Symbol (chemistry), symbol Ag (from the Latin ', derived from the Proto-Indo-European wikt:Reconstruction:Proto-Indo-European/h₂erǵ-, ''h₂erǵ'': "shiny" or "white") and atomic number 47. A soft, whi ...
and white warehou. The young of some species associate with
jellyfish Jellyfish and sea jellies are the informal common names given to the medusa-phase of certain gelatinous members of the subphylum Medusozoa, a major part of the phylum Cnidaria. Jellyfish are mainly free-swimming marine animals with umbrella- ...
, which provides them with protection from predators and opportunities to scavenge the remains of the jellyfish's meals. The young of other species associate with large masses of floating kelp. File:Parapercis hexophtalma 1.jpg, A speckled
sand perch The sandperches are a family, Pinguipedidae, of fishes in the percomorph order Trachiniformes. Sandperches are benthic fish which normally occur over sand or rubble substrates in shallow seas. They are found off the coasts of South America, ...
perched on coral sand File:Centrolophus niger.jpg, The rudderfish (''
Centrolophus niger The black ruff (''Centrolophus niger'') is a medusafish, the only member of the genus ''Centrolophus''. It is a bathypelagic fish found in all tropical and temperate oceans at depths of . Its length is typically up to , but it may reach . Ot ...
'') is a medusa fish File:Epinephelus malabaricus.jpg, Malabar grouper File:Polyprion americanus.2 - Aquarium Finisterrae.JPG,
Atlantic wreckfish The Atlantic wreckfish, (''Polyprion americanus''), also known as the stone bass or bass grouper, is a marine, bathydemersal, and oceanodromous ray-finned fish in the family Polyprionidae. It has a worldwide, if disjunct, distribution in the At ...
inhabit caves and wrecks. They are good game fish, reaching a maximum reported length of and weight of .
*
Grouper Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes. Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is ...
are fish belonging to a number of
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial ...
in the subfamily Epinephelinae of the family
Serranidae The Serranidae are a large family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, in some c ...
, in the order Perciformes. Species of grouper include the
black Black is a color which results from the absence or complete absorption of visible light. It is an achromatic color, without hue, like white and grey. It is often used symbolically or figuratively to represent darkness. Black and white ha ...
,
comet A comet is an icy, small Solar System body that, when passing close to the Sun, warms and begins to release gases, a process that is called outgassing. This produces a visible atmosphere or Coma (cometary), coma, and sometimes also a Comet ta ...
, gag,
giant In folklore, giants (from Ancient Greek: '' gigas'', cognate giga-) are beings of human-like appearance, but are at times prodigious in size and strength or bear an otherwise notable appearance. The word ''giant'' is first attested in 1297 fr ...
,
Goliath Goliath ( ) ''Goləyāṯ''; ar, جُليات ''Ǧulyāt'' (Christian term) or (Quranic term). is a character in the Book of Samuel, described as a Philistine giant defeated by the young David in single combat. The story signified King Saul's ...
, Nassau, saddletail,
tiger The tiger (''Panthera tigris'') is the largest living cat species and a member of the genus '' Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily preys on ...
,
Warsaw Warsaw ( pl, Warszawa, ), officially the Capital City of Warsaw,, abbreviation: ''m.st. Warszawa'' is the capital and largest city of Poland. The metropolis stands on the River Vistula in east-central Poland, and its population is official ...
,
white White is the lightness, lightest color and is achromatic (having no hue). It is the color of objects such as snow, chalk, and milk, and is the opposite of black. White objects fully diffuse reflection, reflect and scattering, scatter all the ...
and yellowfin grouper. Typical lengths are 80–120 centimetres. They inhabit depths from reefs near the surface down to over 400 metres. They feed on just about any moving animal they encounter. Groper are important inshore commercial fish, usually caught with gill nets (in earlier times longlines were used). * Wreckfish are a family Polyprionidae of perciform fishes, found on the floor of the continental shelf and slope where they inhabit caves and shipwrecks (thus their common name). The
Atlantic wreckfish The Atlantic wreckfish, (''Polyprion americanus''), also known as the stone bass or bass grouper, is a marine, bathydemersal, and oceanodromous ray-finned fish in the family Polyprionidae. It has a worldwide, if disjunct, distribution in the At ...
is at depths between . They are largely a solitary fish, though juveniles school below floating objects. Their diet includes large ocean cephalopods, crustaceans, and other bottom-dwelling fishes.


See also

* Coral reef fish *
Demersal fish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They o ...
*
Forage fish Forage fish, also called prey fish or bait fish, are small pelagic fish which are preyed on by larger predators for food. Predators include other larger fish, seabirds and marine mammals. Typical ocean forage fish feed near the base of the ...
*
Marine habitats Marine habitats are habitats that support marine life. Marine life depends in some way on the saltwater that is in the sea (the term ''marine'' comes from the Latin ''mare'', meaning sea or ocean). A habitat is an ecological or environmenta ...
* Moorish idol *
Monkfish Members of the genus ''Lophius'', also sometimes called monkfish, fishing-frogs, frog-fish, and sea-devils, are various species of lophiid anglerfishes found in the Atlantic and Indian Oceans. ''Lophius'' is known as the "monk" or "monkfish" ...
*
NaGISA NaGISA (Natural Geography in Shore Areas or Natural Geography of In-Shore Areas) is an international collaborative effort aimed at inventorying, cataloguing, and monitoring biodiversity of the in-shore area. So named for the Japanese word "nagisa ...
*
Pelagic fish Pelagic fish live in the pelagic zone of ocean or lake waters—being neither close to the bottom nor near the shore—in contrast with demersal fish that do live on or near the bottom, and reef fish that are associated with coral re ...
*
Shoaling and schooling In biology, any group of fish that stay together for social reasons are shoaling, and if the group is swimming in the same direction in a coordinated manner, they are schooling. In common usage, the terms are sometimes used rather loosely. A ...
* Weever


Notes


References

* Elliott, Michael and Hemingway, Krystal (2002
''Fishes in estuaries''
John Wiley & Sons. . * Humann, Paul and DeLoach, Ned (2008
''Coastal Fish Identification: California to Alaska''
New World Publications Incorporated. . * Moyle, PB and Cech, JJ (2004
''Fishes, An Introduction to Ichthyology''
5th Ed, Benjamin Cummings. * Hinrichsen D (1999
''Coastal Waters of the World: Trends, Threats, and Strategies''
Island Press. * Mann KH (2000
''Ecology of coastal waters: with implications for management''
(Vol 8 of ''Studies in ecology'') Wiley-Blackwell.


External links

* Walrond, Carl
"Coastal fish"
Te Ara, the Encyclopedia of New Zealand. Updated 23 November 2009.



Retrieved 20 February 2010.
Estuarine Fish Kills in Morth Carolina
Retrieved 20 February 2010.

{{Fishery science topics Marine fish