Cirrhitus
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''Cirrhitus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or h ...
, hawkfishes from the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Cirrhitidae Cirrhitidae, the hawkfishes, are a family of marine perciform ray-finned fishes found in tropical seas and which are associated with coral reefs. Taxonomy The Cirrhitidae were first recognised as a family by the Scots-born Australian naturali ...
. The species in this genus are found on
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
reef A reef is a ridge or shoal of rock, coral or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic processes— deposition of sand, wave erosion planing down rock o ...
s worldwide.


Taxonomy

''Cirrhites'' was first formally described as a genus in 1803 by the French naturalist Bernard Germain de Lacépède, Lacépède created it as a
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...
genus for his newly described species from Mauritius, ''Cirrhitus maculatus''. However, it was later shown that Lacépède's ''C. maculatus'' was synonymous with ''Labrus pinnulatus'' described in
manuscript A manuscript (abbreviated MS for singular and MSS for plural) was, traditionally, any document written by hand – or, once practical typewriters became available, typewritten – as opposed to mechanically printed or reproduced in ...
by the German naturalist and explorer
Johann Reinhold Forster Johann Reinhold Forster (22 October 1729 – 9 December 1798) was a German Reformed (Calvinist) pastor and naturalist of partially Scottish descent who made contributions to the early ornithology of Europe and North America. He is best known ...
from
Tahiti Tahiti (; Tahitian ; ; previously also known as Otaheite) is the largest island of the Windward group of the Society Islands in French Polynesia. It is located in the central part of the Pacific Ocean and the nearest major landmass is Austra ...
. Forster's was the basis of the description published in 1801 by
Johann Gottlob Schneider Johann Gottlob Theaenus Schneider (18 January 1750 – 12 January 1822) was a German classicist and naturalist. Biography Schneider was born at Collm in Saxony. In 1774, on the recommendation of Christian Gottlob Heine, he became secretary to ...
in his and
Marcus Elieser Bloch Marcus Elieser Bloch (1723–1799) was a German physician and naturalist who is best known for his contribution to ichthyology through his multi-volume catalog of plates illustrating the fishes of the world. Brought up in a Hebrew-speaking Jewish ...
's ''Systema Ichthyologiae'', although ''
Catalog of Fishes Catalog of Fishes is a comprehensive on-line database and reference work on the scientific names of fish species and genera. It is global in its scope and is hosted by the California Academy of Sciences. It has been compiled and is continuously u ...
'' attributes the name to Forster. The name of this genus is dereived from ''cirrhus'' meaning a "lock of hair" or a "barbel", Lacépède did not explain what he feature the name alludes to. It may be alluding to the unbranched pectoral fin rays which Lacépède termed as “''barbillons''”, which means "barbels" in his
description Description is the pattern of narrative development that aims to make vivid a place, object, character, or group. Description is one of four rhetorical modes (also known as ''modes of discourse''), along with exposition, argumentation, and narra ...
of the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...
of the genus '' C. maculatus'', and which he thought to be “false” pectoral fins. Another possibility is that the name refers to cirri extending from the tips of the spines in the
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...
spines, although Lacépède did not mention this feature.


Species

The currently recognized species in this genus are: * ''
Cirrhitus albopunctatus ''Cirrhitus'' is a genus of marine ray-finned fish, hawkfishes from the family Cirrhitidae. The species in this genus are found on tropical reefs worldwide. Taxonomy ''Cirrhites'' was first formally described as a genus in 1803 by the French ...
'' L. P. Schultz, 1950 * ''
Cirrhitus atlanticus ''Cirrhitus'' is a genus of marine ray-finned fish, hawkfishes from the family Cirrhitidae. The species in this genus are found on tropical reefs worldwide. Taxonomy ''Cirrhites'' was first formally described as a genus in 1803 by the French ...
'' Osório, 1893 (West African hawkfish) * '' Cirrhitus pinnulatus'' ( J. R. Forster, 1801) (stocky hawkfish) * ''
Cirrhitus rivulatus The giant hawkfish (''Cirrhitus rivulatus''), also known as the hieroglyphic hawkfish, is a species of marine ray-finned fish, a hawkfish belonging to the family Cirrhitidae. It is a marine fish and the largest of the hawkfish family with maximu ...
''
Valenciennes Valenciennes (, also , , ; nl, label=also Dutch, Valencijn; pcd, Valincyinnes or ; la, Valentianae) is a commune in the Nord department, Hauts-de-France, France. It lies on the Scheldt () river. Although the city and region experienced a ...
, 1846
(giant hawkfish) The American ichthyologist
Leonard Peter Schultz Leonard Peter Schultz (1901–1986) was an American ichthyologist. Biography Schultz was born in 1901, at Albion, Michigan. He received education on ichthyology at Albion College, in which he got his bachelor's degree, in 1924. In 1926, he got ...
recognised three species from the widespread species ''C. pinnulatus'', ''C. spilotoceps'' from the
Red Sea The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; ...
, ''C. pinnulatus'' from the wider Indo-Pacific region except for
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only state ...
and ''C. maculosus'' from Hawaii and the Johnston Atoll.
John Ernest Randall John Ernest "Jack" Randall (May 22, 1924 – April 26, 2020) was an American ichthyologist and a leading authority on coral reef fishes. Randall described over 800 species and authored 11 books and over 900 scientific papers and popular articles. ...
in his 1963 review of the family Cirrhitidae did not recognise these species but treated them as subspecies. ''Catalog of Fishes'' recognises ''C. spilotoceps'' as a valid species and treats ''C. maculosus'' as a subspecies of ''C. pinnulatus'', while FishBase treats these names as synonyms of ''C. pinnulatus''.


Characteristics

''Cirrhites'' hawkfishes have a deep body which is not highly laterally compressed. They have large heads with a blunt snout and have a fringe of cirri on the posterior margin of the anterior nostril. They have a moderately large mouth with two types of teeth, an outer row of canines and an inner row of villiform teeth. There are also teeth on the centre and sides of the roof of the mouth. The upper margin of the preoperculum either has small serrations or it smooth and the gill cover has 2 flattened spines. They have a continuous dorsal fin which has 10 spines, the membranes between the spines having deep notches and the tip of each spine has a large tuft of cirri, and 11-12 soft rays with a small incision separating the spiny portion from the soft rayed portion. The anal fin contains 3 spines and 5-7 soft rays. The caudal fin is trubcate. The lower 7 pectoral fin rays are robust, with deeply incised membranes and these are notabtly longer than other pectoral fin rays. The upper 1 and lower 7 pectoral fin rays are unbranched. The pelvic fin has a single spine and 5 soft rays and has its origin behind the base of the pectoral fin. The scales are smooth, the intraorbital space lacks scales and there are fewer than 12 irregular scale rows, made up of small scales, on the cheeks. There are 41-49 scales on the lateral line. The published sizes vary from a standard length of for ''C. atlanticus'' to a
total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...
of for ''C. rivulatus''.


Distribution and habitat

''Cirrhites'' hawkfishes are found around the tropical seas of the world, except for the western Atlantic Ocean. They prefer hard substrates such as coral reefs, rocky reefs and rocky bottoms.


References

{{Taxonbar, from=Q2674969 Cirrhitidae