Chotoy Spinetail
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The chotoy spinetail (''Schoeniophylax phryganophilus'') is a species of
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
in the Furnariinae subfamily of the ovenbird
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Furnariidae. It is found in
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
, Bolivia,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
,
Paraguay Paraguay (; ), officially the Republic of Paraguay ( es, República del Paraguay, links=no; gn, Tavakuairetã Paraguái, links=si), is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to th ...
, and
Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...
.


Taxonomy and systematics

The chotoy spinetail was originally described in genus '' Sylvia''. Later authors placed it in ''
Synallaxis ''Synallaxis'' is a genus of birds in the ovenbird family, Furnariidae. It is one of the most diverse genera in the family and is composed of small birds that inhabit dense undergrowth across tropical and subtropical habitats in the Neotropical ...
'' but by the early 2010s it was recognized in its own genus ''Schoeniophylax''. It and the white-bellied spinetail (''Mazaria propinqua'') are
sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
.Remsen, Jr., J. V. (2020). Chotoy Spinetail (''Schoeniophylax phryganophilus''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.chospi2.01 retrieved November 20, 2023Derryberry, E. P., S. Claramunt, G. Derryberry, R. T. Chesser, J. Cracraft, A. Aleixo, J. Pérez-Emán, J. V. Remsen, Jr., and R. T. Brumfield. (2011). Lineage diversification and morphological evolution in a large-scale continental radiation: the Neotropical ovenbirds and woodcreepers (Aves: Furnariidae). Evolution 65(10):2973–2986. https://doi.org/10.1111/j.1558-5646.2011.01374.x A high resolution version of the phylogenetic tree in Figure 1 is available from the first author's websit
here
The chotoy spinetail has two subspecies, the
nominate Nomination is part of the process of selecting a candidate for either election to a public office, or the bestowing of an honor or award. A collection of nominees narrowed from the full list of candidates is a short list. Political office In the ...
''S. p. phryganophilus'' (
Vieillot Louis Pierre Vieillot (10 May 1748, Yvetot – 24 August 1830, Sotteville-lès-Rouen) was a French ornithologist. Vieillot is the author of the first scientific descriptions and Linnaean names of a number of birds, including species he collect ...
, 1817) and ''S. p. petersi'' (
Pinto Pinto is a Portuguese language, Portuguese, Spanish language, Spanish, Sephardi Jews, Jewish (Sephardic), and Italian language, Italian surname. It is a high-frequency surname in all List of countries and territories where Portuguese is an officia ...
, 1949).


Description

The chotoy spinetail is long and weighs . It is a large spinetail with a unique throat pattern. The sexes have the same plumage. Adults of the nominate subspecies have a white supercilium and light brown ear
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are s ...
. Their forehead is dark brown, their crown chestnut with faint brown streaks, their back sandy brown with obvious dark brown streaks, and their lower back, rump, and uppertail
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are s ...
light brown. Their wings are mostly light brown with a chestnut shoulder. Their tail is long and light brown with blackish feather shafts. Their chin and upper throat are bright yellow; the lower throat is a crisp black square surrounded by white. Their upper breast is apricot, their lower breast buffy white, their belly whitish, their flanks brownish buff, and their undertail coverts rufescent buff. Their iris is red to brown, their
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. T ...
blackish to dark gray, their
mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...
gray to bluish (sometimes with a dark tip), and their legs and feet light gray to brownish. Juveniles have less sharp streaking on their uppersides than adults, a brown crown, a whitish chin, and an indistinct black throat patch. Subspecies ''S. p. petersi'' has plumage like the nominate's but is smaller.


Distribution and habitat

The nominate subspecies of the chotoy spinetail is by far the more widespread of the two. It is found in eastern Bolivia, Mato Grosso do Sul and Rio Grande do Sul in Brazil, most of Paraguay, northeastern Argentina as far south as northern Buenos Aires Province, and essentially all of Uruguay. Subspecies ''S. p. petersi'' is found disjunctly in northeastern Brazil's states of
Minas Gerais Minas Gerais () is a state in Southeastern Brazil. It ranks as the second most populous, the third by gross domestic product (GDP), and the fourth largest by area in the country. The state's capital and largest city, Belo Horizonte (literally ...
,
Bahia Bahia ( , , ; meaning "bay") is one of the 26 states of Brazil, located in the Northeast Region of the country. It is the fourth-largest Brazilian state by population (after São Paulo, Minas Gerais, and Rio de Janeiro) and the 5th-largest b ...
, and far separated in Piauí. The species inhabits a variety of open to semi-open landscapes including treed savanna;
gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...
, thickets, and scrub along watercourses; monte woodlands; and the edges of marshes. In elevation it ranges from near sea level to .


Behavior


Movement

The chotoy spinetail is a year-round resident throughout its range.


Feeding

The chotoy spinetail feeds on
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chiti ...
s. It typically forages singly or in pairs, gleaning prey from small branches and foliage within about of the ground.


Breeding

The chotoy spinetail breeds in the austral spring and summer, roughly late September to January. It is thought to be monogamous. Its nest is a large ball of thorny sticks with an entrance tube on the side; the tube and inner chamber are lined with softer plant material. It is typically placed in a bush or small tree up to about above the ground but occasionally much higher. The clutch size is three to six eggs but usually four or five. The incubation period is 15 to 16 days and fledging occurs 13 to 14 days after hatch.


Vocalization

The chotoy spinetail's song is "a distinctive series of low-pitched “cho” notes" that sound like they come from a larger bird. It has been rendered as "rrrrtooo chicheetitichichicheecheychachochoochew". Its call is a "hurried 'sreepsreeptrrit' ".


Status

The IUCN has assessed the chotoy spinetail as being of Least Concern. It has a large range and an unknown population size that is believed to be stable. No immediate threats have been identified. It is considered fairly common to common and appears "fairly adaptable, occurring in secondary habitats and scrub".


References


External links


Chotoy spinetail photo gallery
VIREO
Photo-Medium Res
chandra.as.utexas.edu–''"Birds of Brazil"''
Photo-High Res
Tropical Birding {{Taxonbar, from=Q428748 Furnariidae Birds of Uruguay Birds of Paraguay Birds of the Pantanal Birds of Bolivia Birds of Argentina Birds of Brazil Birds described in 1817 Taxa named by Louis Pierre Vieillot Taxonomy articles created by Polbot