Chloris Chloris
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The European greenfinch or simply the greenfinch (''Chloris chloris'') is a small
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
bird in the
finch The true finches are small to medium-sized passerine birds in the family Fringillidae. Finches have stout conical bills adapted for eating seeds and nuts and often have colourful plumage. They occupy a great range of habitats where they are usua ...
family Fringillidae. This bird is widespread throughout Europe,
North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...
and
Southwest Asia Western Asia, West Asia, or Southwest Asia, is the westernmost subregion of the larger geographical region of Asia, as defined by some academics, UN bodies and other institutions. It is almost entirely a part of the Middle East, and includes Anat ...
. It is mainly resident, but some northernmost populations
migrate Migration, migratory, or migrate may refer to: Human migration * Human migration, physical movement by humans from one region to another ** International migration, when peoples cross state boundaries and stay in the host state for some minimum le ...
further south. The greenfinch has also been introduced into
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
,
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
,
Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...
, and
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
.


Taxonomy

The greenfinch was described by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in his landmark 1758 10th edition of ''Systema Naturae'' under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Loxia chloris''. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
is from ''khloris'', the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
name for this bird, from ''khloros'', "green". The finch family,
Fringillidae The true finches are small to medium-sized passerine birds in the family Fringillidae. Finches have stout conical bills adapted for eating seeds and nuts and often have colourful plumage. They occupy a great range of habitats where they are usua ...
, is divided into two subfamilies, the
Carduelinae The cardueline finches are a subfamily, Carduelinae, one of three subfamilies of the finch family Fringillidae, the others being the Fringillinae and the Euphoniinae. The Hawaiian honeycreepers are now included in this subfamily. Except for the H ...
, containing around 28
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
with 141
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
and the
Fringillinae The genus ''Fringilla'' is a small group of finches from the Old World, which are the only species in the subfamily Fringillinae. The genus name ''Fringilla'' is Latin for "finch". Taxonomy The genus ''Fringilla'' was introduced in 1758 by the ...
containing a single genus, ''
Fringilla The genus ''Fringilla'' is a small group of finches from the Old World, which are the only species in the subfamily Fringillinae. The genus name ''Fringilla'' is Latin for "finch". Taxonomy The genus ''Fringilla'' was introduced in 1758 by the ...
'', with four species. The finch family are all seed-eaters with stout conical bills. They have similar skull morphologies, nine large
primaries Primary elections, or direct primary are a voting process by which voters can indicate their preference for their party's candidate, or a candidate in general, in an upcoming general election, local election, or by-election. Depending on the c ...
, 12 tail feathers and no
crop A crop is a plant that can be grown and harvested extensively for profit or subsistence. When the plants of the same kind are cultivated at one place on a large scale, it is called a crop. Most crops are cultivated in agriculture or hydroponic ...
. In all species the female bird builds the nest, incubates the eggs and broods the young. Fringilline finches raise their young almost entirely on
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s, while the cardueline finches raise their young on regurgitated seeds. A
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
study published in 2012 found that the greenfinches are not closely related to other members of the genus ''
Carduelis The genus ''Carduelis'' is a group of birds in the finch family Fringillidae. The genus ''Carduelis'' was introduced by the French zoologist Mathurin Jacques Brisson in 1760 by tautonomy based on Carl Linnaeus's specific epithet for the Europe ...
''. They have therefore been placed in the resurrected genus ''
Chloris In Greek mythology, the name Chloris (; Greek Χλωρίς ''Chlōrís'', from χλωρός ''chlōrós'', meaning "greenish-yellow", "pale green", "pale", "pallid", or "fresh") appears in a variety of contexts. Some clearly refer to different char ...
'' that had originally been introduced by the French naturalist
Georges Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuvier ...
in 1800, with the European greenfinch as the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
.


Subspecies

There are 10 recognised
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
. Greenfinch (Chloris chloris aurantiiventris) Extremadura.jpg, Male ''C. c. aurantiiventris'' Extremadura, Spain European greenfinch (Chloris chloris voousi) male.jpg, Male ''C. c. voousi'', Morocco European greenfinch (Chloris chloris voousi) female.jpg, Female ''C. c. voousi'', Morocco


Description

The European greenfinch is long with a wingspan of . It is similar in size and shape to a
house sparrow The house sparrow (''Passer domesticus'') is a bird of the sparrow family Passeridae, found in most parts of the world. It is a small bird that has a typical length of and a mass of . Females and young birds are coloured pale brown and grey, a ...
, but is mainly green, with yellow in the wings and tail. The female and young birds are duller and have brown tones on the back. The bill is thick and conical. The song contains a lot of trilling twitters interspersed with wheezes, and the male has a "butterfly" display flight. Male greenfinch birds exhibit higher degrees of fluctuating asymmetry. The development of bones of males may be more easily disrupted than that of females.


Behaviour and ecology

Woodland edges, farmland hedges and gardens with relatively thick vegetation are favoured for breeding. It nests in trees or bushes, laying 3 to 6
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...
s. This species can form large flocks outside the breeding season, sometimes mixing with other finches and buntings. They feed largely on seeds, but also take berries.


Reproduction

Breeding season occurs in spring, starting in the second half of March, until June, with fledging young in early July. Incubation lasts about 13–14 days, by the female. The male feeds her at the nest during this period. Chicks are covered with thick, long, greyish-white down at hatching. They are fed on insect larvae by both adults during the first days, and later, by a frequently regurgitated yellowish paste made of seeds. They leave the nest about 13 days later, but they are not able to fly. Usually, they fledge 16–18 days after hatching. This species produces two or three broods per year. In
Australasia Australasia is a region that comprises Australia, New Zealand and some neighbouring islands in the Pacific Ocean. The term is used in a number of different contexts, including geopolitically, physiogeographically, philologically, and ecologica ...
, the European greenfinch's breeding season is from October to March.


Predators and parasites

The
protozoa Protozoa (singular: protozoan or protozoon; alternative plural: protozoans) are a group of single-celled eukaryotes, either free-living or parasitic, that feed on organic matter such as other microorganisms or organic tissues and debris. Histo ...
l parasite ''
Trichomonas gallinae ''Trichomonas gallinae'' is a cosmopolitan parasite of birds including finches, pigeons, doves, turkeys, chickens, parrots, raptors (hawks, golden eagle, etc.). The condition in birds of prey is called frounce.Kennedy, Murry J. "Trichomoniasis ...
'' was known to infect pigeons and raptors, but, beginning in Great Britain in 2005, carcasses of dead European greenfinches and
common chaffinch The common chaffinch or simply the chaffinch (''Fringilla coelebs'') is a common and widespread small passerine bird in the finch family. The male is brightly coloured with a blue-grey cap and rust-red underparts. The female is more subdued in ...
es were found to be infected with the parasite. The disease spread and in 2008, infected carcasses were found in Norway, Sweden and Finland and a year later in Germany. The spread of the disease is believed to have been mediated by common chaffinches, as large numbers of the birds breed in northern Europe and winter in Great Britain. In Great Britain, the number of infected carcasses recovered each year declined after a peak in 2006. There was a reduction in the number of European greenfinches from around 4.3 million to around 2.8 million, but no significant decline in the overall number of common chaffinches. A similar pattern occurred in Finland where, after the arrival of the disease in 2008, there was a reduction in the number of European greenfinches but only a small change in the number of common chaffinches.


In literature

The English poet
William Wordsworth William Wordsworth (7 April 177023 April 1850) was an English Romantic poet who, with Samuel Taylor Coleridge, helped to launch the Romantic Age in English literature with their joint publication ''Lyrical Ballads'' (1798). Wordsworth's ' ...
wrote a poem about this species entitled ''The Green Linnet'' in 1803.Wordsworth, William "The Poetical Works of William Wordsworth". Copyright 1847, 1858 Edward Moxon, Dover Street, London. pp. 118-119.Poetry Foundatio
''The Green Linnet''
/ref>


References


Sources

* * * *


External links


Audio recordings from Xeno-canto

Videos and photos from the Internet Bird Collection

Ageing and sexing by Javier Blasco-Zumeta & Gerd-Michael Heinze


{{Taxonbar , from=Q27075785 Chloris (bird) Finches Birds of Europe Birds described in 1758 Taxa named by Carl Linnaeus