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''Celtis paniculata'', commonly known as tripewood, silky keltis, silky celtis, native hackberry, native celtis, Investigator tree or whitewood, is a rainforest tree native to parts of
Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms, and also a phytogeographical floristic region in the Paleotropical Kingdom. It has been given different definitions. Th ...
,
Melanesia Melanesia (, ) is a subregion of Oceania in the southwestern Pacific Ocean. It extends from Indonesia's New Guinea in the west to Fiji in the east, and includes the Arafura Sea. The region includes the four independent countries of Fiji, Va ...
and Australia.


Description

The tree grows to tall and trunk width, though usually much smaller. The trunk is mostly cylindrical and somewhat buttressed at the base. The bark is thin, greyish brown and smooth, with pustular vertical lines.


Leaves

Leaves are elliptical in shape with a prominent leaf tip. Leaves are alternate and simple, smooth, not toothed, long with a pointed tip. The leaf stalk is somewhat channelled and flattened on the top, and in length. Both sides of the leaf are of a dull green shade. With a hand lens, translucent glands may be seen in the leaves. The base of the leaf is often oblique, that is of unequal length on either side of the leaf stem. Leaf
venation Venation may refer to: * Venation (botany), the arrangement of veins in leaves * Wing venation, the arrangement of veins in insect wings See also * * Vernation Vernation (from ''vernal'' meaning ''spring'', since that is when leaves spring fort ...
is more evident under the leaf. An identifying feature is the intramarginal leaf vein, which starts at the leaf base and travels on either edge of the leaf, a distance of ¼ to ½ the length of the leaf.


Flowers and fruit

Green flowers in axilliary cymes or panicles appear from December to January. The fruit is a blue to black, round to oval shaped drupe, long. containing a single seed which has veiny ridges. Fruit matures from July to March.


Germination and seedlings

Germination takes between 48 and 100 days. The
cotyledon A cotyledon (; ; ; , gen. (), ) is a significant part of the embryo within the seed of a plant, and is defined as "the embryonic leaf in seed-bearing plants, one or more of which are the first to appear from a germinating seed." The num ...
s are large, about 30 x 25 mm with an apex divided into two acute lobes. The first pairs of juvenile leaves are often toothed. At the tenth leaf stage leaves become oval in shape with a prominent tip.


Taxonomy & Naming

This plant first appeared in scientific literature as Solenostigma paniculatum in 1833, published in the ''Prodromus Florae Norfolkicae'' by the
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
n botanist Stephan Endlicher. The
type specimen In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes th ...
was collected at Norfolk Island by the artist
Ferdinand Bauer Ferdinand Lucas Bauer (20 January 1760 – 17 March 1826) was an Austrian botanical illustrator who travelled on Matthew Flinders' expedition to Australia. Biography Early life and career Bauer was born in Feldsberg in 1760, the youngest son ...
. The specific epithet ''paniculata'' refers to the tufted and branched flowers. In 1848
Jules Émile Planchon Jules Émile Planchon (21 March 1823 – 1 April 1888) was a French botanist born in Ganges, Hérault. Biography After receiving his Doctorate of Science at the University of Montpellier in 1844, he worked for a while at the Royal Botanical G ...
moved the species to the
Celtis ''Celtis'' is a genus of about 60–70 species of deciduous trees, commonly known as hackberries or nettle trees, widespread in warm temperate regions of the Northern Hemisphere. The genus is part of the extended hemp family (Cannabaceae). ...
genus.


Origin of the name "Investigator Tree"

In 1802 the explorer Matthew Flinders discovered and named Sweers Island in the Gulf of Carpentaria during his
circumnavigation Circumnavigation is the complete navigation around an entire island, continent, or astronomical body (e.g. a planet or moon). This article focuses on the circumnavigation of Earth. The first recorded circumnavigation of the Earth was the Mage ...
of Australia in the sloop
HMS Investigator Nine ships of the Royal Navy have borne the name HMS ''Investigator''. Another was planned, but renamed before being launched. The name ''Investigator'' passed on to the Royal Indian Navy and after India's Independence, to its successor the Indian ...
, and he carved the word "Investigator" into a tree on the island, a specimen of ''Celtis paniculata''. The tree became well known amongst later explorers, who also carved the names of their vessels into the tree. When the tree was damaged by a
tropical cyclone A tropical cyclone is a rapidly rotating storm system characterized by a low-pressure center, a closed low-level atmospheric circulation, strong winds, and a spiral arrangement of thunderstorms that produce heavy rain and squalls. Depen ...
in 1887, part of the trunk was removed to the
Queensland Museum The Queensland Museum is the state museum of Queensland, dedicated to natural history, cultural heritage, science and human achievement. The museum currently operates from its headquarters and general museum in South Brisbane with specialist m ...
.


Distribution and habitat

''C. paniculatta'' has a wide natural distribution in tropical Asia and the Pacific, including
Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and ea ...
, the
Moluccas The Maluku Islands (; Indonesian: ''Kepulauan Maluku'') or the Moluccas () are an archipelago in the east of Indonesia. Tectonically they are located on the Halmahera Plate within the Molucca Sea Collision Zone. Geographically they are located ...
, the Lesser Sunda Islands, the
Caroline Islands The Caroline Islands (or the Carolines) are a widely scattered archipelago of tiny islands in the western Pacific Ocean, to the north of New Guinea. Politically, they are divided between the Federated States of Micronesia (FSM) in the ce ...
,
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torr ...
, the
Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capit ...
,
Vanuatu Vanuatu ( or ; ), officially the Republic of Vanuatu (french: link=no, République de Vanuatu; bi, Ripablik blong Vanuatu), is an island country located in the South Pacific Ocean. The archipelago, which is of volcanic origin, is east of no ...
, New Caledonia, and the
Northern Territory The Northern Territory (commonly abbreviated as NT; formally the Northern Territory of Australia) is an Australian territory in the central and central northern regions of Australia. The Northern Territory shares its borders with Western Aust ...
and
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , establishe ...
. In Australia it occurs from
Kiama Kiama () is a coastal town 120 kilometres south of Sydney in the Illawarra. One of the main tourist attractions is the Kiama Blowhole. Kiama features several popular surfing beaches and caravan parks, and numerous alfresco cafes and restaurants ...
(34° S) in
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
to
Cape York Peninsula Cape York Peninsula is a large peninsula located in Far North Queensland, Australia. It is the largest unspoiled wilderness in northern Australia.Mittermeier, R.E. et al. (2002). Wilderness: Earth’s last wild places. Mexico City: Agrupació ...
, as well as Norfolk Island. It will grow in a variety of forest types including sandy beach forests, rainforests, monsoon forests and
sclerophyll forests Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaf, leaves, short Internode (botany), internodes (the distance between leaves along the stem) and leaf orientation which is paral ...
, and on a variety of soil types including soils derived from
basalt Basalt (; ) is an aphanitic (fine-grained) extrusive igneous rock formed from the rapid cooling of low-viscosity lava rich in magnesium and iron (mafic lava) exposed at or very near the surface of a rocky planet or moon. More than 90 ...
,
granite Granite () is a coarse-grained ( phaneritic) intrusive igneous rock composed mostly of quartz, alkali feldspar, and plagioclase. It forms from magma with a high content of silica and alkali metal oxides that slowly cools and solidifies under ...
and
limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...
. It is often seen in coastal areas, but in northern parts of Queensland it can be found up to an altitude of .


Ecology & Uses

Celtis paniculata is a food tree for the blue beak, common aeroplane, tailed emperor and other butterflies. Many rainforest pigeons and other birds feed on the fruit. An endangered population of the beetle Menippus darcyi feeds on Celtis paniculata. The tree produces a useful general purpose timber, with a specific gravity of 0.70.


Cultivation

Removal of the flesh from the seed is not advised when sowing.


Gallery

File:Investigator Tree Celtis paniculata Bass Point.jpg , Growing at Bass Point, Australia File:Image_of_the_Investigator_tree_on_Sweers_Island_in_1871.tiff , The original " Investigator Tree" on Sweers Island in 1871 File:Celtis paniculata seedling.jpg ,
Cotyledon A cotyledon (; ; ; , gen. (), ) is a significant part of the embryo within the seed of a plant, and is defined as "the embryonic leaf in seed-bearing plants, one or more of which are the first to appear from a germinating seed." The num ...
s File:Celtis paniculata leaf scan.jpg , Mature leaves


References

{{Taxonbar, from=Q2682213 paniculata Rosales of Australia Trees of Australia Flora of Queensland Flora of New South Wales Flora of Norfolk Island Flora of Malesia Flora of the Caroline Islands Flora of New Guinea Flora of the Solomon Islands (archipelago) Flora of Vanuatu Flora of New Caledonia Taxa named by Stephan Endlicher Plants described in 1833