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Cassin's auklet (''Ptychoramphus aleuticus'') is a small, chunky
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same enviro ...
that ranges widely in the
North Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
. It is the only species placed in the genus ''Ptychoramphus''. It nests in small burrows and because of its presence on well studied
island An island (or isle) is an isolated piece of habitat that is surrounded by a dramatically different habitat, such as water. Very small islands such as emergent land features on atolls can be called islets, skerries, cays or keys. An island ...
s in
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...
and off
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
it is one of the better known
auk An auk or alcid is a bird of the family Alcidae in the order Charadriiformes. The alcid family includes the murres, guillemots, auklets, puffins, and murrelets. The word "auk" is derived from Icelandic ''álka'', from Old Norse ''alka'' (a ...
s. It is named for the American ornithologist
John Cassin John Cassin (September 6, 1813 – January 10, 1869) was an American ornithologist from Pennsylvania. He worked as curator and Vice President at the Philadelphia Academy of Natural Sciences and focused on the systemic classification of the Acade ...
. Cassin's auklet is a small (25 cm, 200 g) nondescript auk. Its
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
is generally dark above and pale below, with a small white mark above the eye. Its bill is overall dark with a pale spot, and its feet are blue. Unlike many other auks, Cassin's auklet lacks dramatic
breed A breed is a specific group of domestic animals having homogeneous appearance (phenotype), homogeneous behavior, and/or other characteristics that distinguish it from other organisms of the same species. In literature, there exist several slig ...
ing plumage, remaining the same over most of the year. At sea it is usually identified by its flight, which is described as looking like a flying
tennis ball A tennis ball is a ball designed for the sport of tennis. Tennis balls are fluorescent yellow in organised competitions, but in recreational play can be virtually any color. Tennis balls are covered in a fibrous felt which modifies their aerodyna ...
. Cassin's auklet ranges from midway up the Baja California peninsula to
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
's
Aleutian Islands The Aleutian Islands (; ; ale, Unangam Tanangin,”Land of the Aleuts", possibly from Chukchi language, Chukchi ''aliat'', "island"), also called the Aleut Islands or Aleutic Islands and known before 1867 as the Catherine Archipelago, are a cha ...
, off
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. It nests on offshore islands, with the main population stronghold being
Triangle Island Triangle Island is a small island, approximately in area,Anne Vallee Triangle Island Ecological Reserve Purpose Statement Ministry of Environmen/ref> located off the northwestern tip of Vancouver Island, British Columbia, Canada. It is situated ap ...
off
Vancouver Island Vancouver Island is an island in the northeastern Pacific Ocean and part of the Canadian Provinces and territories of Canada, province of British Columbia. The island is in length, in width at its widest point, and in total area, while are o ...
's
Cape Scott Cape Scott is a cape at the western side of the terminus of Dennistoun Glacier on the northern coast of Victoria Land in Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the ...
, where the population is estimated to be around 55,0000 pairs. It is not known to be migratory. However, northern birds may move farther south during the winter. Two
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised: *''P. a. aleuticus'' – (
Pallas Pallas may refer to: Astronomy * 2 Pallas asteroid ** Pallas family, a group of asteroids that includes 2 Pallas * Pallas (crater), a crater on Earth's moon Mythology * Pallas (Giant), a son of Uranus and Gaia, killed and flayed by Athena * Pa ...
, 1811)
:
Aleutian Islands The Aleutian Islands (; ; ale, Unangam Tanangin,”Land of the Aleuts", possibly from Chukchi language, Chukchi ''aliat'', "island"), also called the Aleut Islands or Aleutic Islands and known before 1867 as the Catherine Archipelago, are a cha ...
& south
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
to northern
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
.
Kurile Islands The Kuril Islands or Kurile Islands (; rus, Кури́льские острова́, r=Kuril'skiye ostrova, p=kʊˈrʲilʲskʲɪjə ɐstrɐˈva; Japanese: or ) are a volcanic archipelago currently administered as part of Sakhalin Oblast in the ...
. *''P. a. australis'' – van Rossem, 1939: southern
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
. An extinct relative or predecessor species, ''Ptychoramphus tenuis'' L. H. Miller et Bowman, 1958, is known from the Late
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58San Diego Formation The San Diego Formation is a geologic formation in southwestern San Diego County in southern California (United States), and northwestern Baja California (México). Geology It is a coastal transitional marine and non-marine pebble and cobble ...
in
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
.


Behaviour

Cassin's auklet nests in burrows on small islands, and in the southern area of its range may be found in the breeding colony year round. It either digs holes in the soil or uses natural cracks and crevices to nest in, also readily using man-made structures. Pairs will show a strong loyalty towards each other and to a nesting site for many years. Both the parents incubate the single white egg, returning to swap shifts at night (usually after 2300 h) to avoid being taken by predators such as the
western gull The western gull (''Larus occidentalis'') is a large white-headed gull that lives on the west coast of North America. It was previously considered conspecific with the yellow-footed gull (''Larus livens'') of the Gulf of California. The western g ...
or
peregrine falcon The peregrine falcon (''Falco peregrinus''), also known as the peregrine, and historically as the duck hawk in North America, is a Cosmopolitan distribution, cosmopolitan bird of prey (Bird of prey, raptor) in the family (biology), family Falco ...
. They also depart from the colony before dawn. The egg is incubated for 40 days, the small chick is then fed nightly for 35 days by both parents, who regurgitate partially digested food (euphausiids and other small crustaceans) carried in a special
gular pouch Gular skin (throat skin), in ornithology, is an area of featherless skin on birds that joins the lower mandible of the beak (or ''bill'') to the bird's neck. Other vertebrate taxa may have a comparable anatomical structure that is referred to as e ...
, often referred to in the literature as a sublingual pouch. The chick
fledge Fledging is the stage in a flying animal's life between hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnerable c ...
s alone and makes its way to the sea. Cassin's auklet is unusual amongst seabirds in occasionally laying a second clutch after a successful first clutch (it is the only northern hemisphere seabird to do so). Most individuals in a cohort begin breeding at age 3 (27%), and by 8 years >95% of a given cohort has recruited. Mean age of recruitment is 3.6 years. Minimum annual breeding propensity is 0.83, apparent local survival is 0.76, juvenile survival (ages 0 to 2) is 0.15. At sea Cassin's auklets feeds offshore, in clear often
pelagic The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or wa ...
water, often associating with
bathymetric Bathymetry (; ) is the study of underwater depth of ocean floors (''seabed topography''), lake floors, or river floors. In other words, bathymetry is the underwater equivalent to hypsometry or topography. The first recorded evidence of water d ...
landmarks such as underwater canyons and
upwelling Upwelling is an oceanographic phenomenon that involves wind-driven motion of dense, cooler, and usually nutrient-rich water from deep water towards the ocean surface. It replaces the warmer and usually nutrient-depleted surface water. The nutr ...
s. Numbers at sea may be grossly underestimated because the bird moves away from ships at a distance of more than a kilometer. Recently their distribution around Triangle Island has been determined by telemetry. It feeds by diving underwater beating its wings for propulsion, hunting down large
zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
, especially
krill Krill are small crustaceans of the order Euphausiacea, and are found in all the world's oceans. The name "krill" comes from the Norwegian Norwegian, Norwayan, or Norsk may refer to: *Something of, from, or related to Norway, a country in n ...
. It can dive to 30 m below the surface, and by some estimates 80 m.


Conservation

Cassin's auklet is listed as Near Threatened, and some populations (principally the
Farallon Islands The Farallon Islands, or Farallones (from the Spanish ''farallón'' meaning "pillar" or "sea cliff"), are a group of islands and sea stacks in the Gulf of the Farallones, off the coast of San Francisco, California, United States. The island ...
population) have suffered steep declines. Threats to the auklet include introduced carnivores (particularly in Alaska),
oil spill An oil spill is the release of a liquid petroleum hydrocarbon into the environment, especially the marine ecosystem, due to human activity, and is a form of pollution. The term is usually given to marine oil spills, where oil is released into th ...
s and changes in
sea surface temperature Sea surface temperature (SST), or ocean surface temperature, is the ocean temperature close to the surface. The exact meaning of ''surface'' varies according to the measurement method used, but it is between and below the sea surface. Air mass ...
(caused by
El Niño El Niño (; ; ) is the warm phase of the El Niño–Southern Oscillation (ENSO) and is associated with a band of warm ocean water that develops in the central and east-central equatorial Pacific (approximately between the International Date L ...
events). Cassin's auklet is protected under the
Migratory Bird Treaty Act of 1918 The Migratory Bird Treaty Act of 1918 (MBTA), codified at (although §709 is omitted), is a United States federal law, first enacted in 1918 to implement the convention for the protection of migratory birds between the United States and Canada . ...
. Annual variation in ocean climate synchronously affects multiple demographic parameters including survival, breeding propensity, breeding success, and recruitment, a situation conducive to rapid population declines due to climate change.


2014 summer mass death in US

In the last few months of 2014, the carcasses of thousands of Cassin's auklets washed ashore from Northern California up to the north coast of Washington State. The Coastal Observation and Seabird Survey Team estimated a toll between 50,000 and 100,000 deaths that year. Scientists were uncertain about the cause of these deaths. While viruses, bacteria, and oil spills were ruled out the cause appeared to be starvation. Scientists found little evidence of food in the stomachs of the carcasses. Some scientists from the California's Farallon Institute believe these mass deaths could be related to the unusual North Pacific warmth which is pushing marine food chains and could affect other species of zooplankton, krill and fish that normally develop in cold waters and the birds that consume them including the Cassin's auklets. The event happened in the late summer when the auklet chicks began to fledge.


References


Further reading

* *Manuwal, D. A. and A. C. Thoresen. 1993. Cassin's Auklet (''Ptychoramphus aleuticus''). ''In'' The Birds of North America, No. 50 (A. Poole and F. Gill, Eds.). Philadelphia: The Academy of Natural Sciences; Washington, D.C.: The American Ornithologists' Union.


External links


Cassin's auklet images
from tsuru-bird.net by Monte M. Taylor

by VIREO {{taxonbar, from=Q738204 Alcinae Birds of the Aleutian Islands Native birds of Western Canada Native birds of Alaska Birds described in 1811 Taxa named by John Cassin