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The capybaraAlso called capivara (in Brazil), capiguara (in Bolivia), chigüire, chigüiro, or fercho (in Colombia and Venezuela), carpincho (in Argentina, Paraguay and Uruguay) and ronsoco (in Peru). or greater capybara (''Hydrochoerus hydrochaeris'') is a giant cavy rodent native to South America. It is the largest living rodent and a member of the genus '' Hydrochoerus''. The only other
extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...
member is the lesser capybara (''Hydrochoerus isthmius''). Its close relatives include guinea pigs and rock cavies, and it is more distantly related to the agouti, the
chinchilla Chinchillas are either of two species (''Chinchilla chinchilla'' and ''Chinchilla lanigera'') of crepuscular rodents of the parvorder Caviomorpha. They are slightly larger and more robust than ground squirrels, and are native to the Andes mount ...
, and the nutria. The capybara inhabits savannas and dense forests, and lives near bodies of water. It is a highly social species and can be found in groups as large as 100 individuals, but usually live in groups of 10–20 individuals. The capybara is hunted for its meat and hide and also for grease from its thick fatty skin. It is not considered a threatened species.


Etymology

Its
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
is derived from
Tupi Tupi may refer to: * Tupi people of Brazil * Tupi or Tupian languages, spoken in South America ** Tupi language, an extinct Tupian language spoken by the Tupi people * Tupi oil field off the coast of Brazil * Tupi Paulista, a Brazilian municipalit ...
', a complex agglutination of ' (leaf) + ' (slender) + ' (eat) + ' (a suffix for agent nouns), meaning "one who eats slender leaves", or "grass-eater". The
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
, both ''hydrochoerus'' and ''hydrochaeris'', comes from Greek (' "water") and (' "pig, hog"). * In page 57, Darwin says "The largest gnawing animal in the world, the ''Hydrochærus Capybara'' (the water-hog), is here also common." * See it also i
The Complete Work of Charles Darwin Online
/ref>


Classification and phylogeny

The capybara and the lesser capybara both belong to the subfamily Hydrochoerinae along with the rock cavies. The living capybaras and their
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
relatives were previously classified in their own family Hydrochoeridae. Since 2002, molecular phylogenetic studies have recognized a close relationship between ''Hydrochoerus'' and '' Kerodon'', the rock cavies, supporting placement of both genera in a subfamily of Caviidae. Paleontological classifications previously used Hydrochoeridae for all capybaras, while using Hydrochoerinae for the living genus and its closest fossil relatives, such as '' Neochoerus'', but more recently have adopted the classification of Hydrochoerinae within Caviidae. The taxonomy of fossil hydrochoerines is also in a state of flux. In recent years, the diversity of fossil hydrochoerines has been substantially reduced. This is largely due to the recognition that capybara molar teeth show strong variation in shape over the life of an individual. In one instance, material once referred to four genera and seven species on the basis of differences in molar shape is now thought to represent differently aged individuals of a single species, ''Cardiatherium paranense''. Among fossil species, the name "capybara" can refer to the many species of Hydrochoerinae that are more closely related to the modern ''Hydrochoerus'' than to the "cardiomyine" rodents like ''Cardiomys''. The fossil genera ''Cardiatherium'', ''Phugatherium'', ''Hydrochoeropsis'', and ''Neochoerus'' are all capybaras under that concept.


Description

left, Taxidermy specimen of a capybara The capybara has a heavy,
barrel A barrel or cask is a hollow cylindrical container with a bulging center, longer than it is wide. They are traditionally made of wooden staves and bound by wooden or metal hoops. The word vat is often used for large containers for liquids, ...
-shaped body and short head, with reddish-brown fur on the upper part of its body that turns yellowish-brown underneath. Its sweat glands can be found in the surface of the hairy portions of its skin, an unusual trait among rodents. The animal lacks down hair, and its guard hair differs little from over hair.Adult capybaras grow to in length, stand tall at the withers, and typically weigh , with an average in the Venezuelan
llanos The Llanos (Spanish ''Los Llanos'', "The Plains"; ) is a vast tropical grassland plain situated to the east of the Andes in Colombia and Venezuela, in northwestern South America. It is an ecoregion of the tropical and subtropical grasslands, sav ...
of .Capybara
, Arkive
Females are slightly heavier than males. The top recorded weights are for a wild female from Brazil and for a wild male from Uruguay. Also, an 81 kg individual was reported in São Paulo in 2001 or 2002. The dental formula is . Capybaras have slightly webbed feet and vestigial tails. Their hind legs are slightly longer than their forelegs; they have three toes on their rear feet and four toes on their front feet. Their muzzles are blunt, with nostrils, and the eyes and ears are near the top of their heads. Its
karyotype A karyotype is the general appearance of the complete set of metaphase chromosomes in the cells of a species or in an individual organism, mainly including their sizes, numbers, and shapes. Karyotyping is the process by which a karyotype is disce ...
has 2n = 66 and FN = 102, meaning it has 66 chromosomes with a total of 102 arms.


Ecology

Capybaras are semiaquatic mammals found throughout all countries of South America except Chile. They live in densely forested areas near bodies of water, such as lakes, rivers,
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s, ponds, and marshes, as well as flooded
savannah A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the Canopy (forest), canopy does not close. The open canopy allows sufficient light to rea ...
and along rivers in the tropical rainforest. They are superb swimmers and can hold their breath underwater for up to five minutes at a time. Capybara have flourished in cattle ranches. They roam in home
range Range may refer to: Geography * Range (geographic), a chain of hills or mountains; a somewhat linear, complex mountainous or hilly area (cordillera, sierra) ** Mountain range, a group of mountains bordered by lowlands * Range, a term used to i ...
s averaging 10 hectares (25 acres) in high-density populations. Many escapees from captivity can also be found in similar watery habitats around the world. Sightings are fairly common in Florida, although a breeding population has not yet been confirmed. In 2011, one specimen was spotted on the
Central Coast of California The Central Coast is an area of California, roughly spanning the coastal region between Point Mugu and Monterey Bay. It lies northwest of Los Angeles County and south of San Mateo and Santa Clara counties, and includes the rugged, undeveloped ...
. These escaped populations occur in areas where prehistoric capybaras inhabited;
late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division of ...
capybaras inhabited Florida and ''
Hydrochoerus hesperotiganites ''Hydrochoerus hesperotiganites'' is an extinct species of capybara that lived in San Diego County, California, during the Rancholabrean stage of the Pleistocene (between 130,000 and 80,000 years ago). It is currently the only known capybara of th ...
'' in California and '' Hydrochoerus gaylordi'' in
Grenada Grenada ( ; Grenadian Creole French: ) is an island country in the West Indies in the Caribbean Sea at the southern end of the Grenadines island chain. Grenada consists of the island of Grenada itself, two smaller islands, Carriacou and Pe ...
, and feral capybaras in North America may actually fill the ecological niche of the Pleistocene species.


Diet and predation

Capybaras are
herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
s, grazing mainly on grasses and
aquatic plants Aquatic plants are plants that have adapted to living in aquatic environments (saltwater or freshwater). They are also referred to as hydrophytes or macrophytes to distinguish them from algae and other microphytes. A macrophyte is a plant that ...
, as well as fruit and tree bark. They are very selective feeders and feed on the leaves of one species and disregard other species surrounding it. They eat a greater variety of plants during the dry season, as fewer plants are available. While they eat grass during the wet season, they have to switch to more abundant reeds during the dry season. Plants that capybaras eat during the summer lose their nutritional value in the winter, so they are not consumed at that time. The capybara's jaw hinge is not perpendicular, so they chew food by grinding back-and-forth rather than side-to-side. Capybaras are autocoprophagous, meaning they eat their own
feces Feces ( or faeces), known colloquially and in slang as poo and poop, are the solid or semi-solid remains of food that was not digested in the small intestine, and has been broken down by bacteria in the large intestine. Feces contain a relati ...
as a source of bacterial
gut flora Gut microbiota, gut microbiome, or gut flora, are the microorganisms, including bacteria, archaea, fungi, and viruses that live in the digestive tracts of animals. The gastrointestinal metagenome is the aggregate of all the genomes of the gut mi ...
, to help
digest Digest may refer to: Biology *Digestion of food *Restriction digest Literature and publications *''The Digest'', formerly the English and Empire Digest *Digest size magazine format * ''Digest'' (Roman law), also known as ''Pandects'', a digest ...
the cellulose in the grass that forms their normal diet, and to extract the maximum protein and vitamins from their food. They also regurgitate food to masticate again, similar to cud-chewing by cattle. As is the case with other rodents, the front teeth of capybaras grow continually to compensate for the constant wear from eating grasses; their cheek teeth also grow continuously. Like its relative the guinea pig, the capybara does not have the capacity to synthesize vitamin C, and capybaras not supplemented with vitamin C in captivity have been reported to develop gum disease as a sign of scurvy. They can have a lifespan of 8–10 years, but tend to live less than four years in the wild due to predation from jaguars, pumas,
ocelot The ocelot (''Leopardus pardalis'') is a medium-sized spotted wild cat that reaches at the shoulders and weighs between on average. It was first described by Carl Linnaeus in 1758. Two subspecies are recognized. It is native to the southwes ...
s,
eagle Eagle is the common name for many large birds of prey of the family Accipitridae. Eagles belong to several groups of genera, some of which are closely related. Most of the 68 species of eagle are from Eurasia and Africa. Outside this area, just ...
s, and caimans. The capybara is also the preferred prey of the
green anaconda The green anaconda (''Eunectes murinus''), also known as the giant Emerald anaconda, common anaconda, common water boa or sucuri, is a boa species found in South America. It is the heaviest and one of the longest known extant snake species. Lik ...
.


Social organization

Capybaras are known to be gregarious. While they sometimes live solitarily, they are more commonly found in groups of around 10–20 individuals, with two to four adult males, four to seven adult females, and the remainder juveniles. Capybara groups can consist of as many as 50 or 100 individuals during the dry season when the animals gather around available water sources. Males establish social bonds, dominance, or general group consensus. They can make dog-like barks when threatened or when females are herding young. Capybaras have two types of scent glands: a morrillo, located on the snout, and anal glands. Both sexes have these glands, but males have much larger morrillos and use their anal glands more frequently. The anal glands of males are also lined with detachable hairs. A crystalline form of scent secretion is coated on these hairs and is released when in contact with objects such as plants. These hairs have a longer-lasting scent mark and are tasted by other capybaras. Capybaras scent-mark by rubbing their morrillos on objects, or by walking over scrub and marking it with their anal glands. Capybaras can spread their scent further by urinating; however, females usually mark without urinating and scent-mark less frequently than males overall. Females mark more often during the wet season when they are in estrus. In addition to objects, males also scent-mark females.


Reproduction

When in estrus, the female's scent changes subtly and nearby males begin pursuit. In addition, a female alerts males she is in estrus by whistling through her nose. During mating, the female has the advantage and mating choice. Capybaras mate only in water, and if a female does not want to mate with a certain male, she either submerges or leaves the water. Dominant males are highly protective of the females, but they usually cannot prevent some of the subordinates from copulating. The larger the group, the harder it is for the male to watch all the females. Dominant males secure significantly more matings than each subordinate, but subordinate males, as a class, are responsible for more matings than each dominant male. The lifespan of the capybara's sperm is longer than that of other rodents. Capybara
gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregna ...
is 130–150 days, and produces a litter of four young on average, but may produce between one and eight in a single litter. Birth is on land and the female rejoins the group within a few hours of delivering the newborn capybaras, which join the group as soon as they are mobile. Within a week, the young can eat grass, but continue to suckle—from any female in the group—until weaned around 16 weeks. The young form a group within the main group. Alloparenting has been observed in this species. Breeding peaks between April and May in Venezuela and between October and November in
Mato Grosso Mato Grosso ( – lit. "Thick Bush") is one of the states of Brazil, the third largest by area, located in the Central-West region. The state has 1.66% of the Brazilian population and is responsible for 1.9% of the Brazilian GDP. Neighboring ...
, Brazil.


Activities

Though quite agile on land, capybaras are equally at home in the water. They are excellent swimmers, and can remain completely submerged for up to five minutes, an ability they use to evade predators. Capybaras can sleep in water, keeping only their noses out. As temperatures increase during the day, they
wallow Wallowing in animals is comfort behaviour during which an animal rolls about or lies in mud, water or snow. Some definitions include rolling about in dust, however, in ethology this is usually referred to as dust bathing. Wallowing is often comb ...
in water and then graze during the late afternoon and early evening. They also spend time wallowing in mud. They rest around midnight and then continue to graze before dawn.


Conservation and human interaction

Capybaras are not considered a threatened species; their population is stable throughout most of their South American range, though in some areas hunting has reduced their numbers. Capybaras are hunted for their meat and
pelts Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket ...
in some areas, and otherwise killed by humans who see their grazing as competition for livestock. In some areas, they are
farmed Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled people t ...
, which has the effect of ensuring the wetland
habitats In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
are protected. Their survival is aided by their ability to breed rapidly. Capybaras have adapted well to urbanization in South America. They can be found in many areas in zoos and
park A park is an area of natural, semi-natural or planted space set aside for human enjoyment and recreation or for the protection of wildlife or natural habitats. Urban parks are urban green space, green spaces set aside for recreation inside t ...
s, and may live for 12 years in
captivity Captivity, or being held captive, is a state wherein humans or other animals are confined to a particular space and prevented from leaving or moving freely. An example in humans is imprisonment. Prisoners of war are usually held in captivity by a ...
, more than double their wild lifespan. Capybaras are docile and usually allow humans to pet and hand-feed them, but physical contact is normally discouraged, as their ticks can be vectors to Rocky Mountain spotted fever. The European Association of Zoos and Aquaria asked
Drusillas Park Drusillas Park is a small zoo near to Alfriston, in East Sussex, UK. Its exhibits are targeted towards children between 2 and 10 years old. It attracts between 350,000 and 370,000 visitors per year and until December 2021 was home to the fir ...
in Alfriston,
Sussex Sussex (), from the Old English (), is a historic county in South East England that was formerly an independent medieval Anglo-Saxon kingdom. It is bounded to the west by Hampshire, north by Surrey, northeast by Kent, south by the English ...
, England, to keep the studbook for capybaras, to monitor captive populations in Europe. The studbook includes information about all births, deaths and movements of capybaras, as well as how they are related. Capybaras are farmed for meat and skins in South America. The meat is considered unsuitable to eat in some areas, while in other areas it is considered an important source of protein. In parts of South America, especially in Venezuela, capybara meat is popular during
Lent Lent ( la, Quadragesima, 'Fortieth') is a solemn religious observance in the liturgical calendar commemorating the 40 days Jesus spent fasting in the desert and enduring temptation by Satan, according to the Gospels of Matthew, Mark and Luke ...
and Holy Week as the Catholic Church previously issued special dispensation to allow it to be eaten while other meats are generally forbidden. After several attempts a 1784 Papal bull was obtained that allowed the consumption of capybara during Lent. There is widespread perception in Venezuela that consumption of capybaras is exclusive to rural people. Although it is illegal in some states, capybaras are occasionally kept as pets in the United States. The image of a capybara features on the 2-peso coin of Uruguay. In
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
, following the lead of
Izu Shaboten Zoo Izu Shaboten Zoo (Japanese language, Japanese: , Hepburn romanization, Hepburn: ) is a zoological park and botanical garden in Itō, Shizuoka, Itō, Shizuoka Prefecture, Shizuoka, Japan. Founded in 1959, the zoo is currently notable for its exhib ...
in 1982, multiple establishments or zoos in Japan that raise capybaras have adopted the practice of having them relax in
onsen In Japan, are the country's hot springs and the bathing facilities and traditional inns around them. As a volcanically active country, Japan has many onsens scattered throughout all of its major islands. There are approximately 25,000 hot ...
during the winter. They are seen as an attraction by Japanese people. Capybaras became big in Japan due to the popular cartoon character . In August 2021, Argentine and international media reported that capybaras had been causing serious problems for residents of Nordelta, an affluent
gated community A gated community (or walled community) is a form of residential community or housing estate containing strictly controlled entrances for pedestrians, bicycles, and automobiles, and often characterized by a closed perimeter of walls and fences ...
north of Buenos Aires built atop wetland habitat. This inspired Argentine and worldwide leftists to jokingly adopt the capybara as a symbol of
class struggle Class conflict, also referred to as class struggle and class warfare, is the political tension and economic antagonism that exists in society because of socio-economic competition among the social classes or between rich and poor. The forms ...
and communism. Brazilian Lyme-like
borreliosis Lyme disease, also known as Lyme borreliosis, is a vector-borne disease caused by the ''Borrelia'' bacterium, which is spread by ticks in the genus ''Ixodes''. The most common sign of infection is an expanding red rash, known as erythema migran ...
likely involves capybaras as reservoirs and '' Amblyomma'' and ''
Rhipicephalus ''Rhipicephalus'' is a genus of ticks in the family Ixodidae, the hard ticks, consisting of about 74 or 75 species. Most are native to tropical Africa.Olwoch, J. M., et al. (2007)Climate change and the genus ''Rhipicephalus'' (Acari: Ixodidae) in ...
'' ticks as vectors.


Popularity and meme culture

In the early 2020s, capybaras became a growing figure of meme culture due to many factors, including the disturbances in Nordelta which led to them being comically postulated as figures of class struggle. Also, a common meme format includes capybaras in various situations with the song " After Party" by
Don Toliver Caleb Zackery "Don" Toliver ( ; born June 12, 1994) is an American rapper, singer, and songwriter. He released his first major label mixtape, ''Donny Womack'', in August 2018. He is known for his singles "No Idea (song), No Idea" and "After Party ...
, leading to a tremendous growth in popularity. Due to a lyric in Toliver's song, capybaras are also associated with the phrase "Ok I pull up". File:Capybara Ueno Zoo 2009.ogv, Captive capybaras resting at
Ueno Zoo The is a zoo, managed by the Tokyo Metropolitan Government, and located in Taitō, Tokyo, Japan. It is Japan's oldest zoo, opened on March 20, 1882. It is served by Ueno Station, Keisei Ueno Station and Nezu Station, with convenient access fro ...
File:Capybara Izu Shaboten 001.jpg, Capybaras in a bath at
Izu Shaboten Zoo Izu Shaboten Zoo (Japanese language, Japanese: , Hepburn romanization, Hepburn: ) is a zoological park and botanical garden in Itō, Shizuoka, Itō, Shizuoka Prefecture, Shizuoka, Japan. Founded in 1959, the zoo is currently notable for its exhib ...
in Japan


See also

* ''
Josephoartigasia monesi ''Josephoartigasia'' is an extinct genus of enormous dinomyid rodent from the Early Pliocene to Early Pleistocene of Uruguay. The only living member of Dinomyidae is the pacarana. ''Josephoartigasia'' is named after Uruguayan national hero J ...
'', an extinct species identified as the largest known rodent ever * Kurloff cell, a type of cell found in capybaras and guinea pigs * ''
Capybara Walking Animal locomotion, in ethology, is any of a variety of methods that animals use to move from one place to another. Some modes of locomotion are (initially) self-propelled, e.g., running, swimming, jumping, flying, hopping, soaring and gliding. Th ...
'', a historical animal locomotion film by Eadweard Muybridge


Notes


References


External links

*
Animal Diversity Web ''Hydrochoerus hydrochaeris''

Capybara information
{{Taxonbar, from=Q131538 Mammals described in 1766 Cavies Herbivorous mammals Mammals of Argentina Mammals of Bolivia Mammals of Brazil Mammals of Colombia Mammals of Ecuador Mammals of French Guiana Mammals of Guyana Mammals of Paraguay Mammals of Peru Mammals of Uruguay Mammals of Venezuela Articles containing video clips Semiaquatic mammals Rodents of South America Taxa named by Carl Linnaeus 2020s in Internet culture