Carnian Extinctions

The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followed by the Norian. Its boundaries are not characterized by major extinctions or biotic turnovers, but a climatic event (known as the Carnian pluvial episode characterized by substantial rainfall) occurred during the Carnian and seems to be associated with important extinctions or biotic radiations.

Stratigraphic definitions

The Carnian was named in 1869 by Mojsisovics. It is unclear if it was named after the Carnic Alps or after the Austrian region of Carinthia (''Kärnten'' in German) or after the Carnia historical region in northwestern Italy. The name, however, was first used referring to a part of the Hallstatt Limestone cropping out in Austria.

The base of the Carnian Stage is defined as the place in the stratigraphic record where the ammonite species ''Daxatina canadensis'' first appears. The global reference profile for the base is located at the ''Stuores-Wiesen'' near Badia in the Val Badia in the region of South Tyrol, Italy.

The top of the Carnian (the base of the Norian) is at the bases of the ammonite biozones of '' Klamathites macrolobatus'' or '' Stikinoceras kerri'' and the conodont biozones of '' Metapolygnathus communisti'' or '' Metapolygnathus primitius''.

Subdivisions

There is no established, standard usage for the Carnian subdivisions, thus, while in some regional stratigraphies a two-substage subdivision is common:
* Julian
* Tuvalian
others prefer a three-substage organization of the stage as follows:
* Cordevolian
* Julian
* Tuvalian

Biostratigraphy

In the Tethys domain, the Carnian Stage contains six ammonite biozones:
* zone of '' Anatropites spinosus''
* zone of '' Tropites subbullatus''
* zone of '' Tropites dilleri''
* zone of '' Austrotrachyceras austriacum''
* zone of ''Trachyceras''

The Otischalkian land vertebrate faunochron corresponds to the early late Carnian, while the Adamanian land vertebrate faunochron corresponds to the latest Carnian.

Paleogeography and climate

The paleogeography of the Carnian was basically the same as for the rest of the Triassic. Most continents were merged into the supercontinent Pangaea, and there was a single global ocean, Panthalassa. The global ocean had a western branch at tropical latitudes called Paleo-Tethys. The sediments of Paleo-Tethys now crop out in southeastern Europe, in the Middle East, in the Himalayas, and up to the island of Timor.
The extreme land-sea distribution led to "mega-monsoons", i.e., an atmospheric monsoon regime more intense than the present one.

As for most of the Mesozoic, there were no ice caps. Climate was mostly arid in the tropics, but an episode of wet tropical climate is documented at least in the Paleo-Tethys. This putative climatic event is called the " Carnian Pluvial Event", its age being between latest early Carnian (Julian) and the beginning of late Carnian (Tuvalian).

Carnian life

In the marine realm, the Carnian saw the first abundant occurrences of calcareous nanoplankton, a morphological group including the coccolithophores.

Invertebrates

There are a few invertebrates which are typical and characteristic of the Carnian. Among molluscs, the ammonoid genus ''Trachyceras'' is exclusive to the lower Carnian (i.e., Julian of the two-substages subdivision, see above). The family Tropitidae and the genus ''Tropites'' appear at the base of the upper Carnian (Tuvalian). The bivalve genus ''Halobia'', a bottom-dweller of deep sea environments, differentiated from ''Daonella'' at the beginning of this age. Scleractinian coral reefs, i.e., reefs with corals of the modern type, became relatively common for the first time in the Carnian.

Vertebrates

The earliest unequivocal dinosaurs, such as those from the Ischigualato Formation (e.g. ''Herrerasaurus'' and ''Eoraptor'') and those from the Santa Maria Formation (e.g. '' Staurikosaurus'' and ''Buriolestes'') originated during the Carnian, around 230 Ma.

In this stage the archosaurs became the dominant faunas in the world, evolving into groups such as the phytosaurs, rhynchosaurs, aetosaurs, and rauisuchians. The first dinosaurs (and the pterosaur Carniadactylus) also appeared in this stage, and though at the time they were small and insignificant, they diversified rapidly and would dominate the fauna for the rest of the Mesozoic. On the other hand, the therapsids, which included the ancestors of mammals, decreased in both size and diversity, and would remain relatively small until the extinction of the dinosaurs.

Conodonts were present in Triassic marine sediments. '' Paragondolella polygnathiformis'' appeared at the base of the Carnian Stage, and is considered a characteristic species. A partial list of Carnian vertebrates is given below. Many Carnian vertebrates are found in Santa Maria Formation rocks of the Paleorrota geopark.

Classic localities and Lagerstätten

The lower Carnian fauna of the San Cassiano Formation (Dolomites, northern Italy) has been studied since the 19th century. Fossiliferous localities are many, and are distributed mostly in the surroundings of Cortina d'Ampezzo and in the high Badia Valley, near the village of San Cassiano, after which the formation was named. This fauna is extremely diverse, including ammonoids, gastropods, bivalves, echinoderms, calcareous sponge, corals, brachiopods, and a variety of less common fossils. A collection of this fauna is exposed in the "Museo delle Regole", a museum in Cortina d'Ampezzo.

The Ischigualasto Formation of the Ischigualasto-Villa Unión Basin in northwestern Argentina yielded a very important vertebrate association, including the oldest dinosaurian assemblage.

The Lagerstätte of the Madygen Formation in Kyrgyzstan has provided over 20,000 fossil insects, vertebrates and flora.

References

Bibliography

* ; 2005: ''The Global boundary Stratotype Section and Point (GSSP) of the Ladinian Stage (Middle Triassic) at Bagolino (Southern Alps, Northern Italy) and its implications for the Triassic time scale'', Episodes 28(4), pp. 233–244.
* ; 1999: ''The Prati di Stuores/Stuores Wiesen section (Dolomites, Italy): a candidate Global Stratotype section and Point for the base of the Carnian stage'', Rivista Italiana di Paleontologia e Stratigrafia 105, pp. 37–78.
* ; 2006: ''High-precision U-Pb zircon age from the Triassic of Italy: Implications for the Triassic time scale and the Carnian origin of calcareous nannoplankton and dinosaurs'', Geology 34, p. 1009–1012.
* ; 2004: ''A Geologic Time Scale 2004'', Cambridge University Press.
* ; 2012: ''The Geologic Time Scale 2012'', Elsevier.
* ; 1993: ''Adelobasileus from the upper Triassic of west Texas: the oldest mammal'', J. Vert. Paleont. 13, pp. 309–334.
* ; 2012: ''The Global Boundary Stratotype Section and Point (GSSP) of the Carnian stage (Late Triassic) at Prati di Stuores/Stuores Wiesen section (Southern Alps, NE Italy)'', Episodes 35, pp. 414–430.

External links

GeoWhen Database – Carnian

Upper Triassic timescale
at the website of the subcommission for stratigraphic information of the ICS
* Norges Network of offshore records of geology and stratigraphy: ''Stratigraphic charts for the Triassic''

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Palaeos Mesozoic: Carnian Age

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Carnian,
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Geological ages
Triassic geochronology