Candidula Spadae
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''Xerogyra spadae'' is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of air-breathing
land snail A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial gastropod mollusks that have shells (those without shells are known as ...
, a terrestrial
pulmonate Pulmonata or pulmonates, is an informal group (previously an order, and before that a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group includ ...
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
in the family
Geomitridae Geomitridae is a Taxonomy (biology), taxonomic Family (biology), family of small to medium-sized air-breathing land snails, Terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea.MolluscaBase eds. (2021). M ...
. MolluscaBase eds. (2022). MolluscaBase. Xerogyra spadae (Calcara, 1845). Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=1308889 on 2022-07-28 The specific name ''spadae'', "Spada's",
Pietro Calcara Pietro is an Italian masculine given name. Notable people with the name include: People * Pietro I Candiano (c. 842–887), briefly the 16th Doge of Venice * Pietro Tribuno (died 912), 17th Doge of Venice, from 887 to his death * Pietro II Can ...
commemorated
Giovanni Jacopo Spada Giovanni Jacopo Spada (1680 – 1774) was an Italian naturalist and pioneering geologist, born in Verona, who served for many years as archipresbyter at Graz in Styria. He is known for the revised and expanded catalogue of his collection that he pu ...
(1680–1774), an Italian naturalist and pioneering geologist of Verona.


Distribution

''Candidula spadae'' is native to Central Italy. The species in sparsely distributed. It is found in the central
Apennines The Apennines or Apennine Mountains (; grc-gre, links=no, Ἀπέννινα ὄρη or Ἀπέννινον ὄρος; la, Appenninus or  – a singular with plural meaning;''Apenninus'' (Greek or ) has the form of an adjective, which wou ...
. However, the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
asserts that the species is also found in extension to
Gargano Gargano (, Gargano Apulian Italo-Romance arˈgæːnə is a historical and geographical sub-region in the province of Foggia, Apulia, southeast Italy, consisting of a wide isolated mountain massif made of highland and several peaks and forming ...
. It is found in the Sibillini Mountains, including
Monte Vettore Monte Vettore (from Latin ''Vector'', "carrier", "leader") is a mountain of the Umbro-marchigiano Apennine Mountains in Italy. It is the highest peak of the Sibillini massif. It is located in Ascoli piceno, Marche, Italy. Geography The south ...
, and then southward to the
Monti Reatini Monti Reatini is a mountain range in the central Apennines, Italy.The highest peak is the Monte Terminillo Monte Terminillo is a massif in the Monti Reatini, part of the Abruzzi Apennine range in central Italy. It is located some 20 km ...
, the
Gran Sasso d'Italia Gran Sasso d'Italia (; ) is a massif in the Apennine Mountains of Italy. Its highest peak, Corno Grande (2,912 metres), is the highest mountain in the Apennines, and the second-highest mountain in Italy outside the Alps. The mountain lies wit ...
, and the
Monti della Meta The Monti della Meta are a massif of central Italy located around the junction point of the boundaries between the regions of Lazio, Abruzzo and Molise. The major of three massifs of the Parco Nazionale d'Abruzzo, Lazio e Molise, they take their ...
.


Description

The shell of ''Xerogyra spadae'' is pale white in color. Often, there are narrow light brownish color bands on the shell. The shell is almost smooth. The
whorls A whorl ( or ) is an individual circle, oval, volution or equivalent in a whorled pattern, which consists of a spiral or multiple concentric objects (including circles, ovals and arcs). Whorls in nature File:Photograph and axial plane floral d ...
are, to some extent, convex in shape. The number of whorls range between 5.5 and 6. The last two whorls increase more rapidly. The last whorl is weakly angulated. It does not descend near the main opening of the shell, the
aperture In optics, an aperture is a hole or an opening through which light travels. More specifically, the aperture and focal length of an optical system determine the cone angle of a bundle of rays that come to a focus in the image plane. An opt ...
. The aperture is clearly visible and large. The color inside the aperture is white or light yellowish. There is a lip inside the aperture which is white in color. The umbilicus is initially narrow, but at the last
whorl A whorl ( or ) is an individual circle, oval, volution or equivalent in a whorled pattern, which consists of a spiral or multiple concentric objects (including circles, ovals and arcs). Whorls in nature File:Photograph and axial plane floral ...
it increases to one-fourth of the shell diameter. The width of the shell is 10–16 mm. The height of the shell is 6.5–8.5 mm.


Habitat

''Xerogyra spadae'' is a terrestrial species. It lives on pebbly
grassland A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...
in the
alpine tundra Alpine tundra is a type of natural region or biome that does not contain trees because it is at high elevation, with an associated alpine climate, harsh climate. As the latitude of a location approaches the poles, the threshold elevation for alp ...
and
subalpine zone Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
up to the summit region.


Population

According to the IUCN, the size of the population, despite being not properly known, is showing a decreasing trend. The population declined remarkably in the decades 1990s and 2000s. This population decline was caused by an overall dryness in Central Italy's climate.


Interaction with humans and conservation status

''Xerogyra spadae'' faces risk from the tourism industry and
grazing In agriculture, grazing is a method of animal husbandry whereby domestic livestock are allowed outdoors to roam around and consume wild vegetations in order to convert the otherwise indigestible (by human gut) cellulose within grass and other ...
activity. Activities associated with tourism, such as
skiing Skiing is the use of skis to glide on snow. Variations of purpose include basic transport, a recreational activity, or a competitive winter sport. Many types of competitive skiing events are recognized by the International Olympic Committee (IO ...
,
mountain biking Mountain biking is a sport of riding bicycles off-road, often over rough terrain, usually using specially designed mountain bikes. Mountain bikes share similarities with other bikes but incorporate features designed to enhance durability and pe ...
,
hiking Hiking is a long, vigorous walk, usually on trails or footpaths in the countryside. Walking for pleasure developed in Europe during the eighteenth century.AMATO, JOSEPH A. "Mind over Foot: Romantic Walking and Rambling." In ''On Foot: A Histor ...
, as well as
rock climbing Rock climbing is a sport in which participants climb up, across, or down natural rock formations. The goal is to reach the summit of a formation or the endpoint of a usually pre-defined route without falling. Rock climbing is a physically and ...
and
mountaineering Mountaineering or alpinism, is a set of outdoor activities that involves ascending tall mountains. Mountaineering-related activities include traditional outdoor climbing, skiing, and traversing via ferratas. Indoor climbing, sport climbing, a ...
result in degradation of this snail's habitat quality. With the intensity of land-use in the
Apennines The Apennines or Apennine Mountains (; grc-gre, links=no, Ἀπέννινα ὄρη or Ἀπέννινον ὄρος; la, Appenninus or  – a singular with plural meaning;''Apenninus'' (Greek or ) has the form of an adjective, which wou ...
described by the IUCN as "stable or even increasing", grazing activity such as pasturing also results in change of the natural vegetation with adverse effects.


References


External links


Pfeiffer L. (1847-1848). Monographia Heliceorum viventium. Sistens descriptiones systematicas et criticas omnium huius familiae generum et specierum hodie cognitarum. Volumen primum. (1): 1–160

feiffer, L. (1853). Monographia heliceorum viventium. Sistens descriptiones systematicas et criticas omnium huius familiae generum et specierum hodie cognitarum. Volumen tertium [Vol. 3
Lipsiae: Brockhaus. i-viii + 711 pp]
Calcara, P. (1845). Cenno sui molluschi viventi e fossili della Sicilia da servire da supplimento ed insieme di critiche osservazioni all'opera di R.A. Philippi. Stamperia Reale, Palermo, 65 pp., pl. 1-4
{{Taxonbar, from=Q65064615 Endemic fauna of Italy Gastropods described in 1845 spadae