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''Camelops''Being occasionally called ''Western Camel'' or ''Yesterday's Camel''. is an extinct genus of
camel A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. C ...
s that lived in North and Central America, ranging from Alaska to Guatemala, from the middle Pliocene to the end of the Pleistocene. It is more closely related to the
Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by the ...
dromedary and bactrian and wild bactrian camels than the New World guanaco, vicuña, alpaca and llama; making it a true camel of the Camelini tribe. Its name is derived from the Ancient Greek (, "camel") and (, "face"), i.e. "camel-face".


Taxonomy and evolution

The genus ''Camelops'' first appeared during the middle Pliocene (about 4.0–3.2 million years ago (Mya) in southern North America and became extinct at the end of the Pleistocene (around 11,000 years ago). Despite the fact that camels are popularly associated with the deserts of Asia and Africa, the family Camelidae, which comprises camels and llamas, originated in North America during the middle Eocene period, at least 44 Mya. Both the camel and horse families originated in the Americas and migrated into Eurasia via the Bering Strait. Modern camels are descended from the extinct genus '' Paracamelus'', which probably crossed the Bering land bridge into Asia between 7.5 and 6.5 Mya. The divergence between ''Paracamelus'' and ''Camelops'' occurred about 11–10 Mya. ''Paracamelus'' would continue to live in North America as the High Arctic camel until the middle Pleistocene. During Pleistocene warm periods, a smaller morph of ''Camelops'' inhabited Alaska and northern Yukon. These specimens date to around 50–45 thousand years ago, and seem to have been extirpated from the area after this time, similar to the contemporaneous mastodon, the
ground sloth Ground sloths are a diverse group of extinct sloths in the mammalian superorder Xenarthra. The term is used to refer to all extinct sloths because of the large size of the earliest forms discovered, compared to existing tree sloths. The Caribbe ...
'' Megalonyx'', and the giant beaver '' Castoroides''. The skull of a ''Camelops'' specimen was found above the
Glenns Ferry Formation The Glenns Ferry Formation is a Pliocene stratigraphic unit in the western United States. Outcrops of the formation in Hagerman Fossil Beds National Monument preserve the remains of seven fish species, five of which are extinct. These include the te ...
in present-day Idaho in a thick layer of coarse gravel known as the Tauna Gravels. Above this layer of gravel is another layer of fine river channel sands, where the skull was found. The age of this fossil is as young as 2 million years old and perhaps even younger, which can be inferred because it is younger than the other fossils found at the Hagerman Fossil Beds National Monument. During the late
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
and early Miocene periods, camels apparently underwent swift evolutionary change, resulting in several genera with different anatomical structures, ranging from those with short limbs, those with gazelle-like bodies, and giraffe-like camels with long legs and long necks. This rich diversity decreased until only a few species, such as ''Camelops hesternus'', remained in North America, before going extinct entirely around 11,000 years ago. By the end of the Pleistocene, with the extinction of '' Paracamelus'' and '' Titanotylopus'', ''Camelops'' was the only true camel remaining in North America and possibly both Americas. ''Camelops'' extinction was part of a larger North American extinction in which native
horses The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...
, mastodons, and other camelids also died out. Possible causal factors for this megafaunal extinction include global climate change and hunting pressure from human beings. The mass extinction coincided roughly with the appearance of people belonging to the big game-hunting Clovis culture, who were prolific hunters with distinct fluted stone tools, which allowed for a spear shaft to be attached to the stone tool. Biochemical analyses have shown that Clovis tools were used in butchering camels.


Description

Because soft tissues are generally not preserved in the fossil record, it is not certain if ''Camelops'' possessed hump(s), like modern camels, or lacked ones, like modern Camelidaes of South America ( guanacos and vicuñas). One-humped camels are now known to have evolved from two-humped camels, but two-humps, as an evolutionary outcome, likely associated with arctic climates and two-humped camels presumably evolved into one-humped camels in warmer regions in Eurasia, while ''Camelops'' first appeared in southern North America and lived among both warmer and colder regions of the continent until early Holocene. ''C. hesternus'' had legs 20% longer than that of the dromedary, and was about tall at the shoulder and weighed about .Anthony J. Stuart, 2021, ''Vanished Giants: The Lost World of the Ice Age'', "6.17 Yesterday's Camel: Camelops Hesternus", p.99, University of Chicago Press


Palaeobiology

The species ranged widely from Alaska in the north to Oregon and California in the west, southernmost Baja California Peninsula and Mexico and
Guatemala Guatemala ( ; ), officially the Republic of Guatemala ( es, República de Guatemala, links=no), is a country in Central America. It is bordered to the north and west by Mexico; to the northeast by Belize and the Caribbean; to the east by H ...
in the south, and Tennessee in the east, and a notable number of fossils have been excavated among central North America such as at Colorado and California. Plant remains found in the teeth of the Rancho La Brea ''C. hesternus'' fossils further reveal that rather than being limited to grazing, this species likely ate mixed species of plants, including coarse shrubs growing in coastal southern California. ''Camelops'' probably could travel long distances, similar to modern camel species. Whether or not ''Camelops'' could survive for long periods without water, as with extant camels, is still unknown; this may have been an adaptation that occurred much later, after camelids migrated to Asia and Africa. The creosote bush from the US-Mexican border has been speculated to be part of ''Camelopss diet, based on the fact that no local living ungulate eats it, yet it was readily consumed by the Arabian camels of the experimental United States Camel Corps in the mid-19th century.


Extinction

The last species of ''Camelops'' are hypothesized to have disappeared as a result of the Blitzkrieg model. This model presents the hypothesis that ''Camelops'', along with other North American
megafauna In terrestrial zoology, the megafauna (from Greek μέγας ''megas'' "large" and New Latin ''fauna'' "animal life") comprises the large or giant animals of an area, habitat, or geological period, extinct and/or extant. The most common threshold ...
, disappeared as new cultures of experienced and efficient hunters moved southeastward across the continent. The result of this migration and expansion of human populations was a significant reduction in range for the megafauna. Of the many ''Camelops'' specimens recovered in North America, only a small number demonstrate modification through human actions. Some specimens have been interpreted as having been killed by humans based on the presence of spirally fractured bone fragments. None of the reported ''Camelops'' sites has been associated with stone tools, however, which would be an indicator of possible human use. At many of these ''Camelops'' sites, no fossils have been found of carcasses that were evidently processed, but rather small fragments and pieces of remains. Researchers originally thought that ''Camelops'' species were in fact hunted and butchered by early humans in North America because of these reasons: the fragmenting of bones into shapes that look like tools, damage or weathering of the “working” edge of said tools, having attributes that were similar to the making of chopping tools, and scarred fragments from possible chopping tools. Further examination showed, though, that these assumptions were misguided, and that while humans did coexist and associate with ''Camelops'', human use has yet to be completely proven as the sole cause of extinction.


See also

*'' Aepycamelus'' *''
Eulamaops ''Eulamaops'' is an extinct genus of camelid, endemic to South America during the Pleistocene (Lujanian, 781,000—12,000 years ago), existing about . Fossil remains of ''Eulamaops'' have been found in the Luján Formation in Argentina A ...
'' *'' Oxydactylus'' *'' Poebrotherium'' *'' Procamelus'' *'' Protylopus'' *''
Stenomylus ''Stenomylus'' is an extinct genus of miniature camelid native to North America that died out around 30 million years ago. Its name is derived from the Greek (, "narrow") and (, "molar"). ''Stenomylus'' was extremely diminutive compared to ...
'' * Syrian camel, an extinct species that reached at least tall at the shoulder * Pleistocene megafauna * Snowmastodon Project


References

{{Taxonbar, from=Q133439 Prehistoric camelids Prehistoric even-toed ungulate genera Pliocene even-toed ungulates Pleistocene even-toed ungulates Piacenzian first appearances Holocene extinctions Pliocene mammals of North America Pleistocene mammals of North America Taxa named by Joseph Leidy Fossil taxa described in 1854