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A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. Camels have long been domesticated and, as livestock, they provide food ( milk and meat) and textiles (fiber and felt from
hair Hair is a protein filament that grows from follicles found in the dermis. Hair is one of the defining characteristics of mammals. The human body, apart from areas of glabrous skin, is covered in follicles which produce thick terminal and f ...
). Camels are working animals especially suited to their desert habitat and are a vital means of transport for passengers and cargo. There are three surviving species of camel. The one-humped dromedary makes up 94% of the world's camel population, and the two-humped Bactrian camel makes up 6%. The Wild Bactrian camel is a separate species and is now critically endangered. The word ''camel'' is also used informally in a wider sense, where the more correct term is "camelid", to include all seven species of the family
Camelidae Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, ...
: the true camels (the above three species), along with the "New World" camelids: the llama, the alpaca, the guanaco, and the vicuña, which belong to the separate tribe
Lamini Lamini (members are called ''laminoids'') is a tribe of the subfamily Camelinae. It contains one extant genus with four species, all exclusively from South America: llamas, alpacas, vicuñas, and guanacos. The former two are domesticated specie ...
. Camelids originated in North America during the Eocene, with the ancestor of modern camels, '' Paracamelus'', migrating across the Bering land bridge into Asia during the late Miocene, around 6 million years ago.


Taxonomy


Extant species

3 species are extant:


Biology

The average life expectancy of a camel is 40 to 50 years. A full-grown adult dromedary camel stands at the shoulder and at the hump. Bactrian camels can be a foot taller. Camels can run at up to in short bursts and sustain speeds of up to . Bactrian camels weigh and dromedaries . The widening toes on a camel's hoof provide supplemental grip for varying soil sediments. The male dromedary camel has an organ called a dulla in his throat, a large, inflatable sac he extrudes from his mouth when in rut to assert dominance and attract females. It resembles a long, swollen, pink tongue hanging out of the side of his mouth. Camels mate by having both male and female sitting on the ground, with the male mounting from behind. The male usually
ejaculates Ejaculation is the discharge of semen (the ''ejaculate''; normally containing sperm) from the male reproductory tract as a result of an orgasm. It is the final stage and natural objective of male sexual stimulation, and an essential component ...
three or four times within a single mating session. Camelids are the only ungulates to mate in a sitting position.


Ecological and behavioral adaptations

Camels do not directly store water in their humps; they are reservoirs of fatty tissue. When this tissue is metabolized, it yields more than one gram of water for every gram of fat processed. This fat metabolization, while releasing energy, causes water to evaporate from the lungs during respiration (as oxygen is required for the metabolic process): overall, there is a net decrease in water. Camels have a series of physiological adaptations that allow them to withstand long periods of time without any external source of water. The dromedary camel can drink as seldom as once every 10 days even under very hot conditions, and can lose up to 30% of its body mass due to dehydration. Unlike other mammals, camels' red blood cells are oval rather than circular in shape. This facilitates the flow of red blood cells during dehydration and makes them better at withstanding high osmotic variation without rupturing when drinking large amounts of water: a camel can drink of water in three minutes. Camels are able to withstand changes in body temperature and water consumption that would kill most other mammals. Their temperature ranges from at dawn and steadily increases to by sunset, before they cool off at night again. In general, to compare between camels and the other livestock, camels lose only 1.3 liters of fluid intake every day while the other livestock lose 20 to 40 liters per day. Maintaining the brain temperature within certain limits is critical for animals; to assist this, camels have a
rete mirabile A rete mirabile (Latin for "wonderful net"; plural retia mirabilia) is a complex of arteries and veins lying very close to each other, found in some vertebrates, mainly warm-blooded ones. The rete mirabile utilizes countercurrent blood flow within ...
, a complex of arteries and veins lying very close to each other which utilizes countercurrent blood flow to cool blood flowing to the brain. Inside Nature's Giants. Channel 4 (UK) documentary. Transmitted 30 August 2011 Camels rarely sweat, even when ambient temperatures reach . Any sweat that does occur evaporates at the skin level rather than at the surface of their coat; the
heat of vaporization The enthalpy of vaporization (symbol ), also known as the (latent) heat of vaporization or heat of evaporation, is the amount of energy (enthalpy) that must be added to a liquid substance to transform a quantity of that substance into a gas. T ...
therefore comes from body heat rather than ambient heat. Camels can withstand losing 25% of their body weight in water, whereas most other mammals can withstand only about 12–14% dehydration before cardiac failure results from circulatory disturbance. When the camel exhales, water vapor becomes trapped in their nostrils and is reabsorbed into the body as a means to conserve water. Camels eating green herbage can ingest sufficient moisture in milder conditions to maintain their bodies' hydrated state without the need for drinking. The camel's thick coat insulates it from the intense heat radiated from desert sand; a shorn camel must sweat 50% more to avoid overheating. During the summer the coat becomes lighter in color, reflecting light as well as helping avoid sunburn. The camel's long legs help by keeping its body farther from the ground, which can heat up to . Dromedaries have a pad of thick tissue over the sternum called the ''pedestal''. When the animal lies down in a sternal recumbent position, the pedestal raises the body from the hot surface and allows cooling air to pass under the body. Camels' mouths have a thick leathery lining, allowing them to chew thorny desert plants. Long eyelashes and ear hairs, together with nostrils that can close, form a barrier against sand. If sand gets lodged in their eyes, they can dislodge it using their transparent third eyelid (also known as the nictitating membrane). The camels' gait and widened feet help them move without sinking into the sand. The kidneys and intestines of a camel are very efficient at reabsorbing water. Camels' kidneys have a 1:4 cortex to medulla ratio. Thus, the medullary part of a camel's kidney occupies twice as much area as a cow's kidney. Secondly, renal corpuscles have a smaller diameter, which reduces surface area for filtration. These two major anatomical characteristics enable camels to conserve water and limit the volume of urine in extreme desert conditions.Rehan S and AS Qureshi, 2006. Microscopic evaluation of the heart, kidneys and adrenal glands of one-humped camel calves (Camelus dromedarius) using semi automated image analysis system. J Camel Pract Res. 13(2): 123
Camel urine Camel urine is a liquid by-product of metabolism in a camel's anatomy. Urine from Arabian camels has been used in the Arabian Peninsula as prophetic medicine for centuries, being a part of ancient Bedouin practices. After the spread of MERS-CoV ...
comes out as a thick syrup, and camel faeces are so dry that they do not require drying when the Bedouins use them to fuel fires. The camel immune system differs from those of other mammals. Normally, the Y-shaped antibody molecules consist of two heavy (or long) chains along the length of the Y, and two light (or short) chains at each tip of the Y. Camels, in addition to these, also have antibodies made of only two heavy chains, a trait that makes them smaller and more durable. These "heavy-chain-only" antibodies, discovered in 1993, are thought to have developed 50 million years ago, after camelids split from ruminants and pigs. Camels suffer from
surra Surra (from the Marathi ''sūra'', meaning the sound of heavy breathing through nostrils, of imitative origin) is a disease of vertebrate animals. The disease is caused by protozoan trypanosomes, specifically ''Trypanosoma evansi'', of severa ...
caused by ''
Trypanosoma evansi ''Trypanosoma evansi'' is a parasitic species of excavate trypanosome in the genus ''Trypanosoma'' that causes one form of surra in animals. Discovered by Griffith Evans in 1880 at Dera Ismail Khan (British India), it is the first known tryp ...
'' wherever camels are domesticated in the world, and resultantly camels have evolved trypanolytic antibodies as with many mammals. In the future, nanobody/
single-domain antibody A single-domain antibody (sdAb), also known as a nanobody, is an antibody fragment consisting of a single monomeric variable antibody domain. Like a whole antibody, it is able to bind selectively to a specific antigen. With a molecular weight of ...
therapy will surpass natural camel antibodies by reaching locations currently unreachable due to natural antibodies' larger size. Such therapies may also be suitable for other mammals. Tran ''et al.'' 2009 provides a new
reference test A medical test is a medical procedure performed to detect, diagnose, or monitor diseases, disease processes, susceptibility, or to determine a course of treatment. Medical tests such as, physical and visual exams, diagnostic imaging, genetic te ...
for
surra Surra (from the Marathi ''sūra'', meaning the sound of heavy breathing through nostrils, of imitative origin) is a disease of vertebrate animals. The disease is caused by protozoan trypanosomes, specifically ''Trypanosoma evansi'', of severa ...
(''T. evansi'') of camel. They use recombinant (rISG75, an
Invariant Surface Glycoprotein Variant surface glycoprotein (VSG) is a ~60kDa protein which densely packs the cell surface of protozoan parasites belonging to the genus ''Trypanosoma''. This genus is notable for their cell surface proteins. They were first isolated from ''Tryp ...
) and ELISA. The Tran test has high
test specificity ''Sensitivity'' and ''specificity'' mathematically describe the accuracy of a test which reports the presence or absence of a condition. Individuals for which the condition is satisfied are considered "positive" and those for which it is not are ...
and appears likely to work just as well for ''T. evansi'' in other hosts, and for a pan-''
Trypanozoon ''Trypanosoma'' is a genus of Kinetoplastida, kinetoplastids (class Trypanosomatidae), a monophyletic group of unicellular parasite, parasitic flagellate protozoa. Trypanosoma is part of the phylum Sarcomastigophora. The name is derived from the ...
'' test, which would also be useful for '' T. b. brucei'', ''
T. b. gambiense ''Trypanosoma brucei'' is a species of parasitic kinetoplastid belonging to the genus ''Trypanosoma'' that is present in sub-Saharan Africa. Unlike other protozoan parasites that normally infect blood and tissue cells, it is exclusively extracellu ...
'', '' T. b. rhodesiense'', and '' T. equiperdum''.


Genetics

The
karyotype A karyotype is the general appearance of the complete set of metaphase chromosomes in the cells of a species or in an individual organism, mainly including their sizes, numbers, and shapes. Karyotyping is the process by which a karyotype is disce ...
s of different camelid species have been studied earlier by many groups, but no agreement on chromosome nomenclature of camelids has been reached. A 2007 study flow sorted camel chromosomes, building on the fact that camels have 37 pairs of chromosomes (2n=74), and found that the karyotype consisted of one metacentric, three submetacentric, and 32 acrocentric autosomes. The Y is a small metacentric chromosome, while the X is a large metacentric chromosome. The hybrid camel, a hybrid between Bactrian and dromedary camels, has one hump, though it has an indentation deep that divides the front from the back. The hybrid is at the shoulder and tall at the hump. It weighs an average of and can carry around , which is more than either the dromedary or Bactrian can. According to molecular data, the wild Bactrian camel (''C. ferus'') separated from the domestic Bactrian camel (''C. bactrianus'') about 1 million years ago. New World and Old World camelids diverged about 11 million years ago. In spite of this, these species can hybridize and produce viable offspring. The cama is a camel-llama hybrid bred by scientists to see how closely related the parent species are. Scientists
collected semen Semen collection refers to the process of obtaining semen from human males or other animals with the use of various methods, for the purposes of artificial insemination, or medical study (usually in fertility clinics). Semen can be collected via ...
from a camel via an artificial vagina and inseminated a llama after stimulating ovulation with
gonadotrophin Gonadotropins are glycoprotein hormones secreted by gonadotropic cells of the anterior pituitary of vertebrates. This family includes the mammalian hormones follicle-stimulating hormone (FSH) and luteinizing hormone (LH), the placental/chorionic ...
injections. The cama is halfway in size between a camel and a llama and lacks a hump. It has ears intermediate between those of camels and llamas, longer legs than the llama, and partially
cloven hooves A cloven hoof, cleft hoof, divided hoof or split hoof is a hoof split into two toes. This is found on members of the mammalian order Even-toed ungulate, Artiodactyla. Examples of mammals that possess this type of hoof are cattle, deer, pigs, ant ...
. Like the
mule The mule is a domestic equine hybrid between a donkey and a horse. It is the offspring of a male donkey (a jack) and a female horse (a mare). The horse and the donkey are different species, with different numbers of chromosomes; of the two pos ...
, camas are sterile, despite both parents having the same number of chromosomes.


Evolution

The earliest known camel, called ''
Protylopus ''Protylopus'' is an extinct genus of camel that lived during middle to late Eocene some 50-40 million years ago in North America. Along with being the oldest camel known, it was also the smallest, reaching a length of , and probably weighing a ...
'', lived in North America 40 to 50 million years ago (during the Eocene). It was about the size of a rabbit and lived in the open woodlands of what is now South Dakota. By 35 million years ago, the ''
Poebrotherium ''Poebrotherium'' ( ) is an extinct genus of camelid, endemic to North America. They lived from the Eocene to Miocene epochs, 46.3—13.6 mya, existing for approximately . Discovery and history ''Poebrotherium'' was first named by scientist J ...
'' was the size of a goat and had many more traits similar to camels and llamas. The hoofed ''
Stenomylus ''Stenomylus'' is an extinct genus of miniature camelid native to North America that died out around 30 million years ago. Its name is derived from the Greek (, "narrow") and (, "molar"). ''Stenomylus'' was extremely diminutive compared to ...
'', which walked on the tips of its toes, also existed around this time, and the long-necked '' Aepycamelus'' evolved in the Miocene. The ancestor of modern camels, '' Paracamelus'', migrated into Eurasia from North America via
Beringia Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip ...
during the late Miocene, between 7.5 and 6.5 million years ago. During the Pleistocene, around 3 to 1 million years ago, the North American Camelidae spread to South America as part of the
Great American Interchange The Great American Biotic Interchange (commonly abbreviated as GABI), also known as the Great American Interchange and the Great American Faunal Interchange, was an important late Cenozoic paleozoogeographic biotic interchange event in which lan ...
via the newly formed
Isthmus of Panama The Isthmus of Panama ( es, Istmo de Panamá), also historically known as the Isthmus of Darien (), is the narrow strip of land that lies between the Caribbean Sea and the Pacific Ocean, linking North and South America. It contains the country ...
, where they gave rise to
guanacos The guanaco (; ''Lama guanicoe'') is a camelid native to South America, closely related to the llama. Guanacos are one of two wild South American camelids, the other being the vicuña, which lives at higher elevations. Etymology The guanaco g ...
and related animals. Populations of ''Paracamelus'' continued to exist in the North American Arctic into the
Early Pleistocene The Early Pleistocene is an unofficial sub-epoch in the international geologic timescale in chronostratigraphy, being the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently estimated to span the time ...
. This creature is estimated to have stood around tall. The Bactrian camel diverged from the dromedary about 1 million years ago, according to the fossil record. The last camel native to North America was ''
Camelops hesternus ''Camelops''Being occasionally called ''Western Camel'' or ''Yesterday's Camel''. is an extinct genus of camels that lived in North and Central America, ranging from Alaska to Guatemala, from the middle Pliocene to the end of the Pleistocene. It ...
'', which vanished along with horses, short-faced bears, mammoths and mastodons,
ground sloth Ground sloths are a diverse group of extinct sloths in the mammalian superorder Xenarthra. The term is used to refer to all extinct sloths because of the large size of the earliest forms discovered, compared to existing tree sloths. The Caribbe ...
s,
sabertooth cat Machairodontinae is an extinct subfamily of carnivoran mammals of the family Felidae (true cats). They were found in Asia, Africa, North America, South America, and Europe from the Miocene to the Pleistocene, living from about 16 million ...
s, and many other megafauna, coinciding with the migration of humans from Asia at the end of the Pleistocene, around 15–11,000 years ago. File:Stenomylus.jpg, alt=A drawing of two early camels,
Stenomylus ''Stenomylus'' is an extinct genus of miniature camelid native to North America that died out around 30 million years ago. Its name is derived from the Greek (, "narrow") and (, "molar"). ''Stenomylus'' was extremely diminutive compared to ...
illustration File:NMNH-USNMV16601Stenomylus.tif,
Stenomylus ''Stenomylus'' is an extinct genus of miniature camelid native to North America that died out around 30 million years ago. Its name is derived from the Greek (, "narrow") and (, "molar"). ''Stenomylus'' was extremely diminutive compared to ...
skeleton File:NMNH-USNMV15917Poebrotherium.jpg,
Poebrotherium ''Poebrotherium'' ( ) is an extinct genus of camelid, endemic to North America. They lived from the Eocene to Miocene epochs, 46.3—13.6 mya, existing for approximately . Discovery and history ''Poebrotherium'' was first named by scientist J ...
skeleton File:NMNH-USNM244271 2.jpg, Procamelus skull File:Camelops hesternus.jpg, alt=, ''
Camelops hesternus ''Camelops''Being occasionally called ''Western Camel'' or ''Yesterday's Camel''. is an extinct genus of camels that lived in North and Central America, ranging from Alaska to Guatemala, from the middle Pliocene to the end of the Pleistocene. It ...
'', the last true camel native to North America


Domestication

Like horses, camels originated in North America and eventually spread across
Beringia Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip ...
to Asia. They survived in the Old World, and eventually humans domesticated them and spread them globally. Along with many other megafauna in North America, the original wild camels were wiped out during the spread of the first indigenous peoples of the Americas from Asia into North America, 10 to 12,000 years ago; although fossils have never been associated with definitive evidence of hunting. Most camels surviving today are domesticated. Although feral populations exist in
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
, India and Kazakhstan, wild camels survive only in the wild Bactrian camel population of the
Gobi Desert The Gobi Desert (Chinese: 戈壁 (沙漠), Mongolian: Говь (ᠭᠣᠪᠢ)) () is a large desert or brushland region in East Asia, and is the sixth largest desert in the world. Geography The Gobi measures from southwest to northeast an ...
.


History

When humans first domesticated camels is disputed. Dromedaries may have first been domesticated by humans in Somalia or
South Arabia South Arabia () is a historical region that consists of the southern region of the Arabian Peninsula in Western Asia, mainly centered in what is now the Republic of Yemen, yet it has also historically included Najran, Jizan, Al-Bahah, and 'Asi ...
sometime during the
3rd millennium BC The 3rd millennium BC spanned the years 3000 through 2001 BC. This period of time corresponds to the Early to Middle Bronze Age, characterized by the early empires in the Ancient Near East. In Ancient Egypt, the Early Dynastic Period is followe ...
, the Bactrian in central Asia around 2,500 BC, as at Shar-i Sokhta (also known as the Burnt City), Iran. Martin Heide's 2010 work on the domestication of the camel tentatively concludes that humans had domesticated the Bactrian camel by at least the middle of the third millennium somewhere east of the Zagros Mountains, with the practice then moving into Mesopotamia. Heide suggests that mentions of camels "in the patriarchal narratives may refer, at least in some places, to the Bactrian camel", while noting that the camel is not mentioned in relationship to Canaan. Recent excavations in the Timna Valley by Lidar Sapir-Hen and Erez Ben-Yosef discovered what may be the earliest domestic camel bones yet found in Israel or even outside the
Arabian Peninsula The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plate ...
, dating to around 930 BC. This garnered considerable media coverage, as it is strong evidence that the stories of Abraham, Jacob, Esau, and Joseph were written after this time. The existence of camels in Mesopotamia—but not in the eastern Mediterranean lands—is not a new idea. The historian Richard Bulliet did not think that the occasional mention of camels in the Bible meant that the domestic camels were common in the Holy Land at that time. The archaeologist
William F. Albright William Foxwell Albright (May 24, 1891– September 19, 1971) was an American archaeologist, biblical scholar, philologist, and expert on ceramics. He is considered "one of the twentieth century's most influential American biblical scholars." ...
, writing even earlier, saw camels in the Bible as an
anachronism An anachronism (from the Ancient Greek, Greek , 'against' and , 'time') is a chronology, chronological inconsistency in some arrangement, especially a juxtaposition of people, events, objects, language terms and customs from different time per ...
. The official report by Sapir-Hen and Ben-Joseph notes:
The introduction of the dromedary camel (Camelus dromedarius) as a pack animal to the southern Levant ... substantially facilitated trade across the vast deserts of Arabia, promoting both economic and social change (e.g., Kohler 1984; Borowski 1998: 112–116; Jasmin 2005). This ... has generated extensive discussion regarding the date of the earliest domestic camel in the southern Levant (and beyond) (e.g., Albright 1949: 207; Epstein 1971: 558–584; Bulliet 1975; Zarins 1989; Köhler-Rollefson 1993; Uerpmann and Uerpmann 2002; Jasmin 2005; 2006; Heide 2010; Rosen and Saidel 2010; Grigson 2012). Most scholars today agree that the dromedary was exploited as a pack animal sometime in the early Iron Age (not before the 12th century C
and concludes:
Current data from copper smelting sites of the Aravah Valley enable us to pinpoint the introduction of domestic camels to the southern Levant more precisely based on stratigraphic contexts associated with an extensive suite of
radiocarbon dates Radiocarbon dating (also referred to as carbon dating or carbon-14 dating) is a method for determining the age of an object containing organic material by using the properties of radiocarbon, a radioactive isotope of carbon. The method was dev ...
. The data indicate that this event occurred not earlier than the last third of the 10th century Cand most probably during this time. The coincidence of this event with a major reorganization of the copper industry of the region—attributed to the results of the campaign of Pharaoh
Shoshenq I Hedjkheperre Setepenre Shoshenq I (Egyptian ''ššnq''; reigned c. 943–922 BC)—also known as Shashank or Sheshonk or Sheshonq Ifor discussion of the spelling, see Shoshenq—was a pharaoh of ancient Egypt and the founder of the Twenty-secon ...
—raises the possibility that the two were connected, and that camels were introduced as part of the efforts to improve efficiency by facilitating trade.
File:Camel cart.JPG, alt= A camel harnessed to a cart loaded with branches and twigs, A camel serving as a
draft animal A working animal is an animal, usually domesticated, that is kept by humans and trained to perform tasks instead of being slaughtered to harvest animal products. Some are used for their physical strength (e.g. oxen and draft horses) or for t ...
in Pakistan (2009) File:A camel with its rider playing kettle drums..jpg, alt= A painting of a man sitting on a camel and playing the drums, A camel in a ceremonial procession, its rider playing kettledrums, Mughal Empire (c. 1840) File:Negev camel petroglyph.jpg, Petroglyph of a camel, Negev, southern Israel (prior to c. 5300 BC) File:Bartholomeus Breenbergh 002.jpg, ''Joseph Sells Grain'' by
Bartholomeus Breenbergh Bartholomeus Breenbergh (before 13 November 1598 – after 3 October 1657) was a Dutch Golden Age painter of Italian and Italianate landscapes, in Rome (1619-1630) and Amsterdam (1630-1657). Biography Little is known of his early life. In his ...
(1655), showing camel with rider at left


Textiles

Desert tribes and Mongolian nomads use camel hair for tents, yurts, clothing, bedding and accessories. Camels have outer guard hairs and soft inner down, and the fibers are sorted by color and age of the animal. The guard hairs can be felted for use as waterproof coats for the herdsmen, while the softer hair is used for premium goods. The fiber can be spun for use in weaving or made into yarns for hand knitting or crochet. Pure camel hair is recorded as being used for western garments from the 17th century onwards, and from the 19th century a mixture of wool and camel hair was used.


Military uses

By at least 1200 BC the first camel saddles had appeared, and
Bactrian camels The Bactrian camel (''Camelus bactrianus''), also known as the Mongolian camel or domestic Bactrian camel, is a large even-toed ungulate native to the steppes of Central Asia. It has two humps on its back, in contrast to the single-humped drome ...
could be ridden. The first saddle was positioned to the back of the camel, and control of the Bactrian camel was exercised by means of a stick. However, between 500 and 100 BC, Bactrian camels came into military use. New saddles, which were inflexible and bent, were put over the humps and divided the rider's weight over the animal. In the seventh century BC the military Arabian saddle evolved, which again improved the saddle design slightly. Military forces have used camel cavalries in wars throughout Africa, the Middle East, and into the modern-day Border Security Force (BSF) of India (though as of July 2012, the BSF planned the replacement of camels with
ATVs ATV may refer to: Broadcasting * Amateur television *Analog television Television stations and companies * Ràdio i Televisió d'Andorra * ATV (Armenia) * ATV (Aruba), NBC affiliate * ATV (Australian TV station), Melbourne * ATV (Austria) * ATV ...
). The first documented use of camel cavalries occurred in the Battle of Qarqar in 853 BC. Armies have also used camels as freight animals instead of horses and mules. The
East Roman Empire The Byzantine Empire, also referred to as the Eastern Roman Empire or Byzantium, was the continuation of the Roman Empire primarily in its eastern provinces during Late Antiquity and the Middle Ages, when its capital city was Constantinopl ...
used auxiliary forces known as ''
dromedarii Dromedarii were camel-riding auxiliary forces recruited in the desert provinces of the Late Roman Empire in Syria. They were developed to take the place of horses, where horses were not common. They were also successful against enemy horses, as ...
'', whom the Romans recruited in desert provinces. The camels were used mostly in combat because of their ability to scare off horses at close range (horses are afraid of the camels' scent), a quality famously employed by the Achaemenid Persians when fighting
Lydia Lydia (Lydian language, Lydian: ‎𐤮𐤱𐤠𐤭𐤣𐤠, ''Śfarda''; Aramaic: ''Lydia''; el, Λυδία, ''Lȳdíā''; tr, Lidya) was an Iron Age Monarchy, kingdom of western Asia Minor located generally east of ancient Ionia in the mod ...
in the
Battle of Thymbra The Battle of Thymbra was the decisive battle in the war between Croesus of the Lydian Kingdom and Cyrus the Great of the Achaemenid Empire. Cyrus, after he had pursued Croesus into Lydia after the drawn Battle of Pteria, met the remains of Croe ...
(547 BC).


19th and 20th centuries

The United States Army established the U.S. Camel Corps, stationed in California, in the 19th century. One may still see stables at the
Benicia Arsenal The Benicia Arsenal (1851–1964) and Benicia Barracks (1852–66) were part of a large military reservation located next to Suisun Bay in Benicia, California. For over 100 years, the arsenal was the primary US Army Ordnance facility for the West ...
in
Benicia, California Benicia ( , ) is a waterside city in Solano County, California, located in the North Bay region of the San Francisco Bay Area. It served as the capital of California for nearly thirteen months from 1853 to 1854. The population was 26,997 at the ...
, where they nowadays serve as the Benicia Historical Museum. Though the experimental use of camels was seen as a success ( John B. Floyd, Secretary of War in 1858, recommended that funds be allocated towards obtaining a thousand more camels), the outbreak of the American Civil War in 1861 saw the end of the Camel Corps: Texas became part of the Confederacy, and most of the camels were left to wander away into the desert. France created a ''
méhariste Méhariste is a French word that roughly translates to camel cavalry. The word is most commonly used as a designation of military units. French camel corps Origins France created a corps of méhariste camel companies (''Compagnies Méharistes ...
'' camel corps in 1912 as part of the
Armée d'Afrique The Army of Africa (french: Armée d’Afrique ) was an unofficial but commonly used term for those portions of the French Army stationed in French North Africa (Morocco, Algeria and Tunisia) from 1830 until the end of the Algerian War in 1962, ...
in the Sahara in order to exercise greater control over the camel-riding Tuareg and Arab insurgents, as previous efforts to defeat them on foot had failed. The
Free French Camel Corps The Free French Camel Corps (french: Corps de Méharistes Français Libres) was a méhariste camel cavalry unit of the Free French forces, founded by among others Captain Edouard Dieffenbach, under command of General Charles de Gaulle during Wor ...
fought during World War II, and camel-mounted units remained in service until the end of French rule over Algeria in 1962. In 1916, the British created the Imperial Camel Corps. It was originally used to fight the
Senussi The Senusiyya, Senussi or Sanusi ( ar, السنوسية ''as-Sanūssiyya'') are a Muslim political-religious tariqa (Sufi order) and clan in colonial Libya and the Sudan region founded in Mecca in 1837 by the Grand Senussi ( ar, السنوسي ...
, but was later used in the Sinai and Palestine Campaign in World War I. The Imperial Camel Corps comprised infantrymen mounted on camels for movement across desert, though they dismounted at battle sites and fought on foot. After July 1918, the Corps began to become run down, receiving no new reinforcements, and was formally disbanded in 1919. In World War I, the British Army also created the
Egyptian Camel Transport Corps The Egyptian Camel Transport Corps (known as the CTC, Camel Corps or Camel Transport) were a group of Egyptian camel drivers who supported the British Army in Egypt during the First World War's Sinai and Palestine Campaign. The work done by the 17 ...
, which consisted of a group of Egyptian camel drivers and their camels. The Corps supported British war operations in
Sinai Sinai commonly refers to: * Sinai Peninsula, Egypt * Mount Sinai, a mountain in the Sinai Peninsula, Egypt * Biblical Mount Sinai, the site in the Bible where Moses received the Law of God Sinai may also refer to: * Sinai, South Dakota, a place ...
, Palestine, and Syria by transporting supplies to the troops. The Somaliland Camel Corps was created by colonial authorities in
British Somaliland British Somaliland, officially the Somaliland Protectorate ( so, Dhulka Maxmiyada Soomaalida ee Biritishka), was a British Empire, British protectorate in present-day Somaliland. During its existence, the territory was bordered by Italian Soma ...
in 1912; it was disbanded in 1944. Bactrian camels were used by Romanian forces during World War II in the Caucasian region. At the same period the Soviet units operating around Astrakhan in 1942 adopted local camels as draft animals due to shortage of trucks and horses, and kept them even after moving out of the area. Despite severe losses, some of these camels ended up as far west as to Berlin itself. The Bikaner Camel Corps of British India fought alongside the
British Indian Army The British Indian Army, commonly referred to as the Indian Army, was the main military of the British Raj before its dissolution in 1947. It was responsible for the defence of the British Indian Empire, including the princely states, which co ...
in World Wars I and II. The ''
Tropas Nómadas The (Nomad Troops) were an auxiliary regiment to the colonial army in Spanish Sahara (today Western Sahara), from the 1930s until the end of the Spanish presence in the territory in 1975. Composed of Sahrawi tribesmen, the Tropas Nómadas were equ ...
'' (Nomad Troops) were an auxiliary regiment of Sahrawi tribesmen serving in the colonial army in Spanish Sahara (today Western Sahara). Operational from the 1930s until the end of the Spanish presence in the territory in 1975, the ''Tropas Nómadas'' were equipped with small arms and led by Spanish officers. The unit guarded outposts and sometimes conducted patrols on camelback.


21st century competition

At the King Abdulaziz Camel Festival, in Saudi Arabia, thousands of camels are paraded and are judged on their lips and humps. The festival also features camel racing and camel milk tasting and has combined prize money of $57m (£40m). In 2018, 12 camels were disqualified from the beauty contest after it was discovered their owners had tried to improve their camel's good looks with injections of botox, into the animals' lips, noses and jaws. In 2021 over 40 camels were disqualified for acts of tampering and deception in beautifying camels.


Food uses


Dairy

Camel milk is a
staple food A staple food, food staple, or simply a staple, is a food that is eaten often and in such quantities that it constitutes a dominant portion of a standard diet for a given person or group of people, supplying a large fraction of energy needs and ...
of desert nomad tribes and is sometimes considered a meal itself; a nomad can live on only camel milk for almost a month. Camel milk can readily be made into yogurt, but can only be made into butter if it is soured first, churned, and a clarifying agent is then added. Until recently, camel milk could not be made into camel cheese because
rennet Rennet () is a complex set of enzymes produced in the stomachs of ruminant mammals. Chymosin, its key component, is a protease enzyme that curdles the casein in milk. In addition to chymosin, rennet contains other enzymes, such as pepsin and a ...
was unable to coagulate the milk proteins to allow the collection of curds. Developing less wasteful uses of the milk, the FAO commissioned Professor J.P. Ramet of the
École Nationale Supérieure d'Agronomie et des Industries Alimentaires École may refer to: * an elementary school in the French educational stages normally followed by secondary education establishments (collège and lycée) * École (river), a tributary of the Seine flowing in région Île-de-France * École, Sav ...
, who was able to produce curdling by the addition of calcium phosphate and vegetable rennet in the 1990s. The cheese produced from this process has low levels of cholesterol and is easy to digest, even for the lactose intolerant. Camel milk can also be made into ice cream.


Meat

They provide food in the form of meat and milk. Approximately 3.3 million camels and camelids are slaughtered each year for meat worldwide. A camel carcass can provide a substantial amount of meat. The male dromedary carcass can weigh , while the carcass of a male Bactrian can weigh up to . The carcass of a female dromedary weighs less than the male, ranging between . The brisket, ribs and loin are among the preferred parts, and the hump is considered a delicacy. The hump contains "white and sickly fat", which can be used to make the ''khli'' (preserved meat) of mutton, beef, or camel. On the other hand, camel milk and meat are rich in protein, vitamins, glycogen, and other nutrients making them essential in the diet of many people. From chemical composition to meat quality, the dromedary camel is the preferred breed for meat production. It does well even in arid areas due to its unusual physiological behaviors and characteristics, which include tolerance to extreme temperatures, radiation from the sun, water paucity, rugged landscape and low vegetation. Camel meat is reported to taste like coarse beef, but older camels can prove to be very tough, although camel meat becomes tenderer the more it is cooked. Camel is one of the animals that can be ritually slaughtered and divided into three portions (one for the home, one for extended family/social networks, and one for those who cannot afford to slaughter an animal themselves) for the qurban of Eid al-Adha. The
Abu Dhabi Abu Dhabi (, ; ar, أَبُو ظَبْيٍ ' ) is the capital and second-most populous city (after Dubai) of the United Arab Emirates. It is also the capital of the Emirate of Abu Dhabi and the centre of the Abu Dhabi Metropolitan Area. ...
Officers' Club serves a camel burger mixed with beef or lamb fat in order to improve the texture and taste. In Karachi, Pakistan, some restaurants prepare nihari from camel meat. Specialist camel butchers provide expert cuts, with the hump considered the most popular. Camel meat has been eaten for centuries. It has been recorded by ancient Greek writers as an available dish at banquets in ancient Persia, usually roasted whole. The Roman emperor Heliogabalus enjoyed camel's heel. Camel meat is mainly eaten in certain regions, including
Eritrea Eritrea ( ; ti, ኤርትራ, Ertra, ; ar, إرتريا, ʾIritriyā), officially the State of Eritrea, is a country in the Horn of Africa region of Eastern Africa, with its capital and largest city at Asmara. It is bordered by Ethiopia ...
, Somalia, Djibouti, Saudi Arabia, Egypt,
Syria Syria ( ar, سُورِيَا or سُورِيَة, translit=Sūriyā), officially the Syrian Arab Republic ( ar, الجمهورية العربية السورية, al-Jumhūrīyah al-ʻArabīyah as-Sūrīyah), is a Western Asian country loc ...
, Libya,
Sudan Sudan ( or ; ar, السودان, as-Sūdān, officially the Republic of the Sudan ( ar, جمهورية السودان, link=no, Jumhūriyyat as-Sūdān), is a country in Northeast Africa. It shares borders with the Central African Republic t ...
, Ethiopia, Kazakhstan, and other arid regions where alternative forms of protein may be limited or where camel meat has had a long cultural history. Camel blood is also consumable, as is the case among pastoralists in northern Kenya, where camel blood is drunk with milk and acts as a key source of iron, vitamin D, salts and minerals. A 2005 report issued jointly by the Saudi
Ministry of Health Ministry of Health may refer to: Note: Italics indicate now-defunct ministries. * Ministry of Health (Argentina) * Ministry of Health (Armenia) * Australia: ** Ministry of Health (New South Wales) * Ministry of Health (The Bahamas) * Ministry of ...
and the United States Centers for Disease Control and Prevention details four cases of human
bubonic plague Bubonic plague is one of three types of plague caused by the plague bacterium (''Yersinia pestis''). One to seven days after exposure to the bacteria, flu-like symptoms develop. These symptoms include fever, headaches, and vomiting, as well a ...
resulting from the ingestion of raw camel liver.


Australia

Camel meat is also occasionally found in Australian cuisine: for example, a camel lasagna is available in
Alice Springs Alice Springs ( aer, Mparntwe) is the third-largest town in the Northern Territory of Australia. Known as Stuart until 31 August 1933, the name Alice Springs was given by surveyor William Whitfield Mills after Alice, Lady Todd (''née'' Al ...
. Australia has exported camel meat, primarily to the Middle East but also to Europe and the US, for many years. The meat is very popular among East African Australians, such as Somalis, and other Australians have also been buying it. The feral nature of the animals means they produce a different type of meat to farmed camels in other parts of the world, and it is sought after because it is disease-free, and a unique genetic group. Demand is outstripping supply, and governments are being urged not to cull the camels, but redirect the cost of the cull into developing the market. Australia has seven camel dairies, which produce milk, cheese and skincare products in addition to meat.


Religion


Islam

Muslims consider camel meat '' halal'' ( ar, حلال, 'allowed'). However, according to some
Islamic schools of thought Islamic schools and branches have different understandings of Islam. There are many different sects or denominations, schools of Islamic jurisprudence, and schools of Islamic theology, or ''ʿaqīdah'' (creed). Within Islamic groups themselves ...
, a state of impurity is brought on by the consumption of it. Consequently, these schools hold that Muslims must perform ''
wudhu Wuḍūʾ ( ar, الوضوء ' ) is the Islamic procedure for cleansing parts of the body, a type of ritual purification, or ablution. The 4 Fardh (Mandatory) acts of ''Wudu'' consists of washing the face, arms, then wiping the head and the feet ...
'' (ablution) before the next time they pray after eating camel meat. Also, some Islamic schools of thought consider it '' haram'' ( ar, حرام, 'forbidden') for a Muslim to perform ''
Salat (, plural , romanized: or Old Arabic ͡sˤaˈloːh, ( or Old Arabic ͡sˤaˈloːtʰin construct state) ), also known as ( fa, نماز) and also spelled , are prayers performed by Muslims. Facing the , the direction of the Kaaba wit ...
'' in places where camels lie, as it is said to be a dwelling place of the ''
Shaytan ' (; ''devils'' or ''demons''), singular: (شَيْطَان) are evil spirits in Islam, inciting humans (and jinn) to sin by "whispering" (وَسْوَسَة, “waswasah”) to their hearts (قَلْب ''qalb''). Folklore suggests that they a ...
'' ( ar, شيطان, ' Devil'). According to Abu Yusuf (d.798), the urine of camel may be used for medical treatment if necessary, but according to
Abū Ḥanīfa Nuʿmān ibn Thābit ibn Zūṭā ibn Marzubān ( ar, نعمان بن ثابت بن زوطا بن مرزبان; –767), commonly known by his '' kunya'' Abū Ḥanīfa ( ar, أبو حنيفة), or reverently as Imam Abū Ḥanīfa by Sunni Musl ...
h, the drinking of camel urine is discouraged. The Islamic texts contain several stories featuring camels. In the story of the people of Thamud, the Prophet Salih miraculously brings forth a '' naqat'' ( ar, ناقة, '
milch-camel Camel milk has supported nomad and pastoral cultures since the domestication of camels millennia ago. Herders may for periods survive solely on the milk when taking the camels on long distances to graze in desert and arid environments, especially i ...
') out of a rock. After the Prophet Muhammad migrated from Mecca to Medina, he allowed his she-camel to roam there; the location where the camel stopped to rest determined the location where he would build his house in Medina.


Judaism

According to Jewish tradition, camel meat and milk are not
kosher (also or , ) is a set of dietary laws dealing with the foods that Jewish people are permitted to eat and how those foods must be prepared according to Jewish law. Food that may be consumed is deemed kosher ( in English, yi, כּשר), fro ...
. Camels possess only one of the two kosher criteria; although they chew their cud, they do not possess
cloven hooves A cloven hoof, cleft hoof, divided hoof or split hoof is a hoof split into two toes. This is found on members of the mammalian order Even-toed ungulate, Artiodactyla. Examples of mammals that possess this type of hoof are cattle, deer, pigs, ant ...
: "But these you shall not eat among those that bring up the cud and those that have a cloven hoof: the camel, because it brings up its cud, but does not have a ompletelycloven hoof; it is unclean for you."


Cultural depictions

What may be the oldest carvings of camels were discovered in 2018 in Saudi Arabia. They were analysed by researchers from several scientific disciplines and, in 2021, were estimated to be 7,000 to 8,000 years old. The dating of
rock art In archaeology, rock art is human-made markings placed on natural surfaces, typically vertical stone surfaces. A high proportion of surviving historic and prehistoric rock art is found in caves or partly enclosed rock shelters; this type also ...
is made difficult by the lack of organic material in the carvings that may be tested, so the researchers attempting to date them tested animal bones found associated with the carvings, assessed erosion patterns, and analysed tool marks in order to determine a correct date for the creation of the sculptures. This Neolithic dating would make the carvings significantly older than Stonehenge (5,000 years old) and the Egyptian pyramids at Giza (4,500 years old) and it predates estimates for the domestication of camels. File:Shadda (detail), Karabagh region, southwest Caucasus.jpeg, Shadda (cover,detail), Karabagh region, southwest Caucasus, early 19th century File:Vessel in the Form of a Recumbent Camel with Jugs, 2015.65.15.jpg, Vessel in the form of a recumbent camel with jugs, 250 BC – 224 AD,
Brooklyn Museum The Brooklyn Museum is an art museum located in the New York City borough of Brooklyn. At , the museum is New York City's second largest and contains an art collection with around 1.5 million objects. Located near the Prospect Heights, Crown H ...
File:Brooklyn Museum - Maru Ragini (Dhola and Maru riding on a Camel).jpg, ''Maru Ragini'' (''Dhola and Maru Riding on a Camel)'', c. 1750,
Brooklyn Museum The Brooklyn Museum is an art museum located in the New York City borough of Brooklyn. At , the museum is New York City's second largest and contains an art collection with around 1.5 million objects. Located near the Prospect Heights, Crown H ...
File:Brooklyn Museum - The Magi Journeying (Les rois mages en voyage) - James Tissot - overall.jpg, ''The Magi Journeying'' (''Les rois mages en voyage'')—James Tissot, c. 1886,
Brooklyn Museum The Brooklyn Museum is an art museum located in the New York City borough of Brooklyn. At , the museum is New York City's second largest and contains an art collection with around 1.5 million objects. Located near the Prospect Heights, Crown H ...
File:KiplingCamel3.gif, ''How the Camel Got His Hump'' (From Rudyard Kipling's ''
Just So Stories ''Just So Stories for Little Children'' is a 1902 collection of origin stories by the British author Rudyard Kipling. Considered a classic of children's literature, the book is among Kipling's best known works. Kipling began working on the ...
'')


Distribution and numbers

There are approximately 14 million camels alive , with 90% being dromedaries. Dromedaries alive today are domesticated animals (mostly living in the
Horn of Africa The Horn of Africa (HoA), also known as the Somali Peninsula, is a large peninsula and geopolitical region in East Africa.Robert Stock, ''Africa South of the Sahara, Second Edition: A Geographical Interpretation'', (The Guilford Press; 2004), ...
, the
Sahel The Sahel (; ar, ساحل ' , "coast, shore") is a region in North Africa. It is defined as the ecoclimatic and biogeographic realm of transition between the Sahara to the north and the Sudanian savanna to the south. Having a hot semi-arid c ...
, Maghreb, Middle East and South Asia). The Horn region alone has the largest concentration of camels in the world, where the dromedaries constitute an important part of local nomadic life. They provide nomadic people in Somalia and Ethiopia with milk, food, and transportation.Plain text version.
Over one million dromedary camels are estimated to be feral in Australia, descended from those introduced as a method of transport in the 19th and early 20th centuries. This population is growing about 8% per year; it was estimated at around 700,000 in 2008. Representatives of the Australian government have culled more than 100,000 of the animals in part because the camels use too much of the limited resources needed by sheep farmers. A small population of introduced camels, dromedaries and Bactrians, wandered through
Southwestern United States The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural region of the United States that generally includes Arizona, New Mexico, and adjacent portions of California, Colorado, Ne ...
after having been imported in the 19th century as part of the U.S. Camel Corps experiment. When the project ended, they were used as draft animals in mines and escaped or were released. Twenty-five U.S. camels were bought and exported to Canada during the Cariboo Gold Rush. The Bactrian camel is, , reduced to an estimated 1.4 million animals, most of which are domesticated. The Wild Bactrian camel is a separate species and is the only truly wild (as opposed to feral) camel in the world. The wild camels are critically endangered and number approximately 1400, inhabiting the Gobi and Taklamakan Deserts in China and Mongolia.


See also

* Afghan cameleers in Australia * Australian feral camel *
Camel howdah A howdah, or houdah (Hindi: हौदा ''haudā''), derived from the Arabic (hawdaj), which means "bed carried by a camel", also known as ''hathi howdah'' (''hāthī haudā'', हाथी हौदा), is a carriage which is positioned on ...
*
Camel milk Camel milk has supported nomad and pastoral cultures since the domestication of camels millennia ago. Herders may for periods survive solely on the milk when taking the camels on long distances to graze in desert and arid environments, especially i ...
* Camel racing * Camel train (caravan) *
Camel urine Camel urine is a liquid by-product of metabolism in a camel's anatomy. Urine from Arabian camels has been used in the Arabian Peninsula as prophetic medicine for centuries, being a part of ancient Bedouin practices. After the spread of MERS-CoV ...
* Camel wrestling * Camelops * '' Camelus moreli'' * Dromedary *
List of animals with humps This is a list of animals that have a naturally occurring hump or humps as a part of their anatomy. Humps may evolve, as a store of fat, as a heat control mechanism, as a development of muscular strength, as a form of display to other animals o ...
* Xerocole


Citations


General and cited references


Camels and Camel Milk. Report Issued by Food and Agriculture Organization of the United Nations. (1982)
* * * *


Further reading

*


External links




Six Green Reasons to Drink Camel's Milk



The Camel as a pet

"Could Emirati camels hold the key to treating venomous snake bites?"
{{Authority control African cuisine Arab cuisine Camelids Domesticated animals Halal meat Livestock Middle Eastern cuisine Taxa named by Carl Linnaeus