HOME

TheInfoList



OR:

The Cambrian Period ( ; sometimes symbolized ) was the first
geological period The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochronol ...
of the
Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...
Era, and of the
Phanerozoic Eon The Phanerozoic Eon is the current geologic eon in the geologic time scale, and the one during which abundant animal and plant life has existed. It covers 538.8 million years to the present, and it began with the Cambrian Period, when anima ...
. The Cambrian lasted 53.4 million years from the end of the preceding
Ediacaran The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and th ...
Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by
Adam Sedgwick Adam Sedgwick (; 22 March 1785 – 27 January 1873) was a British geologist and Anglican priest, one of the founders of modern geology. He proposed the Cambrian and Devonian period of the geological timescale. Based on work which he did on W ...
, who named it after
Cambria Cambria is a name for Wales, being the Latinised form of the Welsh name for the country, . The term was not in use during the Roman period (when Wales had not come into existence as a distinct entity). It emerged later, in the medieval period, a ...
, the Latin name for 'Cymru' (
Wales Wales ( cy, Cymru ) is a Countries of the United Kingdom, country that is part of the United Kingdom. It is bordered by England to the Wales–England border, east, the Irish Sea to the north and west, the Celtic Sea to the south west and the ...
), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with
Roderick Murchison Sir Roderick Impey Murchison, 1st Baronet, (19 February 1792 – 22 October 1871) was a Scotland, Scottish geologist who served as director-general of the British Geological Survey from 1855 until his death in 1871. He is noted for investigat ...
, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Cambrian biology surpasses that of some later periods. The Cambrian marked a profound change in
life on Earth Life on Earth may refer to: Science * Life * Earliest known life forms * Evolutionary history of life ** Abiogenesis Film and television * ''Life on Earth'' (film) (''La Vie Sur Terre''), a 1998 Malian film * ''Life on Earth'' (TV series), a 197 ...
: prior to the Cambrian, the majority of living organisms on the whole were small,
unicellular A unicellular organism, also known as a single-celled organism, is an organism that consists of a single cell, unlike a multicellular organism that consists of multiple cells. Organisms fall into two general categories: prokaryotic organisms and ...
and simple (
Ediacaran The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and th ...
fauna Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is '' funga''. Flora, fauna, funga and other forms of life are collectively referred to as '' biota''. Zoo ...
being notable exceptions). Complex,
multicellular organism A multicellular organism is an organism that consists of more than one cell, in contrast to unicellular organism. All species of animals, land plants and most fungi are multicellular, as are many algae, whereas a few organisms are partially uni- ...
s gradually became more common in the millions of years immediately preceding the Cambrian, but it was not until this period that mineralized – hence readily fossilized – organisms became common. The rapid diversification of lifeforms in the Cambrian, known as the
Cambrian explosion The Cambrian explosion, Cambrian radiation, Cambrian diversification, or the Biological Big Bang refers to an interval of time approximately in the Cambrian Period when practically all major animal phyla started appearing in the fossil recor ...
, produced the first representatives of all modern animal
phyla Phyla, the plural of ''phylum'', may refer to: * Phylum, a biological taxon between Kingdom and Class * by analogy, in linguistics, a large division of possibly related languages, or a major language family which is not subordinate to another Phyl ...
. Phylogenetic analysis has supported the view that before the Cambrian radiation, in the
Cryogenian The Cryogenian (from grc, κρύος, krýos, meaning "cold" and , romanized: , meaning "birth") is a geologic period that lasted from . It forms the second geologic period of the Neoproterozoic Era, preceded by the Tonian Period and followed by ...
or
Tonian The Tonian (from grc, τόνος, tónos, meaning "stretch") is the first geologic period of the Neoproterozoic era (geology), Era. It lasted from to Mya (million years ago). Instead of being based on stratigraphy, these dates are defined by t ...
, animals (metazoans) evolved monophyletically from a single common ancestor: flagellated colonial
protists A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the excl ...
similar to modern choanoflagellates. Although diverse life forms prospered in the oceans, the land is thought to have been comparatively barren – with nothing more complex than a microbial soil crust and a few molluscs and arthropods (albeit not terrestrial) that emerged to browse on the microbial biofilm. By the end of the Cambrian, myriapods, arachnids, and hexapods started adapting to the land, along with the first plants. Most of the
continent A continent is any of several large landmasses. Generally identified by convention rather than any strict criteria, up to seven geographical regions are commonly regarded as continents. Ordered from largest in area to smallest, these seven ...
s were probably dry and rocky due to a lack of vegetation. Shallow seas flanked the margins of several continents created during the breakup of the supercontinent
Pannotia Pannotia (from Greek: '' pan-'', "all", '' -nótos'', "south"; meaning "all southern land"), also known as the Vendian supercontinent, Greater Gondwana, and the Pan-African supercontinent, was a relatively short-lived Neoproterozoic supercontinent ...
. The seas were relatively warm, and polar ice was absent for much of the period.


Stratigraphy

The Cambrian Period followed the
Ediacaran The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and th ...
Period and was followed by the Ordovician Period. The base of the Cambrian lies atop a complex assemblage of trace fossils known as the '' Treptichnus pedum'' assemblage.A. Knoll, M. Walter, G. Narbonne, and N. Christie-Blick (2004)
The Ediacaran Period: A New Addition to the Geologic Time Scale.
Submitted on Behalf of the Terminal Proterozoic Subcommission of the International Commission on Stratigraphy.
The use of ''Treptichnus pedum'', a reference ichnofossil to mark the lower boundary of the Cambrian, is problematic because very similar trace fossils belonging to the Treptichnids group are found well below ''T. pedum'' in
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...
,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
and
Newfoundland Newfoundland and Labrador (; french: Terre-Neuve-et-Labrador; frequently abbreviated as NL) is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region ...
, and possibly in the western USA. The stratigraphic range of ''T. pedum'' overlaps the range of the
Ediacaran The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and th ...
fossils in Namibia, and probably in Spain.M.A. Fedonkin, B.S. Sokolov, M.A. Semikhatov, N.M.Chumakov (2007).
Vendian versus Ediacaran: priorities, contents, prospectives.
" In: edited by M. A. Semikhatov
The Rise and Fall of the Vendian (Ediacaran) Biota. Origin of the Modern Biosphere. Transactions of the International Conference on the IGCP Project 493, August 20–31, 2007, Moscow.
" Moscow: GEOS.
A. Ragozina, D. Dorjnamjaa, A. Krayushkin, E. Serezhnikova (2008).

". 33 Intern. Geol. Congr. 6–14 August 2008, Oslo, Norway. Abstracts. Section HPF 07 Rise and fall of the Ediacaran (Vendian) biota. p. 183.


Subdivisions

The Cambrian is divided into four
epochs In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided by ...
( series) and ten
ages Ages may refer to: *Advanced glycation end-products, known as AGEs *Ages, Kentucky, census-designated place, United States * ''Ages'' (album) by German electronic musician Edgar Froese *The geologic time scale, a system of chronological measuremen ...
(
stages Stage or stages may refer to: Acting * Stage (theatre), a space for the performance of theatrical productions * Theatre, a branch of the performing arts, often referred to as "the stage" * ''The Stage'', a weekly British theatre newspaper * S ...
). Currently only three series and six stages are named and have a GSSP (an internationally agreed-upon stratigraphic reference point). Because the international stratigraphic subdivision is not yet complete, many local subdivisions are still widely used. In some of these subdivisions the Cambrian is divided into three epochs with locally differing names – the Early Cambrian (Caerfai or Waucoban, mya),
Middle Cambrian Middle or The Middle may refer to: * Centre (geometry), the point equally distant from the outer limits. Places * Middle (sheading), a subdivision of the Isle of Man * Middle Bay (disambiguation) * Middle Brook (disambiguation) * Middle Creek (di ...
(St Davids or Albertan, mya) and
Late Cambrian Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, ...
( mya; also known as Merioneth or Croixan).
Trilobite zone Trilobites are used as index fossils to subdivide the Cambrian period. Assemblages of trilobites define trilobite zones. The ''Olenellus''-zone has traditionally marked the top of the Lower Cambrian, and is followed by the '' Eokochaspis'' zone. ...
s allow biostratigraphic correlation in the Cambrian. Rocks of these epochs are referred to as belonging to the Lower, Middle, or Upper Cambrian. Each of the local series is divided into several stages. The Cambrian is divided into several regional
faunal stages In chronostratigraphy, a stage is a succession of rock strata laid down in a single age on the geologic timescale, which usually represents millions of years of deposition. A given stage of rock and the corresponding age of time will by conventi ...
of which the Russian-Kazakhian system is most used in international parlance: *Most Russian paleontologists define the lower boundary of the Cambrian at the base of the Tommotian Stage, characterized by diversification and global distribution of organisms with mineral skeletons and the appearance of the first Archaeocyath bioherms.


Dating the Cambrian

The International Commission on Stratigraphy lists the Cambrian Period as beginning at and ending at . The lower boundary of the Cambrian was originally held to represent the first appearance of complex life, represented by
trilobite Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the At ...
s. The recognition of
small shelly fossils The small shelly fauna, small shelly fossils (SSF), or early skeletal fossils (ESF) are mineralized fossils, many only a few millimetres long, with a nearly continuous record from the latest stages of the Ediacaran to the end of the Early Camb ...
before the first trilobites, and Ediacara biota substantially earlier, led to calls for a more precisely defined base to the Cambrian Period. Despite the long recognition of its distinction from younger Ordovician rocks and older
Precambrian The Precambrian (or Pre-Cambrian, sometimes abbreviated pꞒ, or Cryptozoic) is the earliest part of Earth's history, set before the current Phanerozoic Eon. The Precambrian is so named because it preceded the Cambrian, the first period of the ...
rocks, it was not until 1994 that the Cambrian system/period was internationally ratified. After decades of careful consideration, a continuous sedimentary sequence at Fortune Head,
Newfoundland Newfoundland and Labrador (; french: Terre-Neuve-et-Labrador; frequently abbreviated as NL) is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region ...
was settled upon as a formal base of the Cambrian Period, which was to be correlated worldwide by the earliest appearance of '' Treptichnus pedum''. Discovery of this fossil a few metres below the GSSP led to the refinement of this statement, and it is the ''T. pedum'' ichnofossil assemblage that is now formally used to correlate the base of the Cambrian. This formal designation allowed radiometric dates to be obtained from samples across the globe that corresponded to the base of the Cambrian. Early dates of quickly gained favour, though the methods used to obtain this number are now considered to be unsuitable and inaccurate. A more precise date using modern radiometric dating yield a date of . The ash horizon in Oman from which this date was recovered corresponds to a marked fall in the abundance of carbon-13 that correlates to equivalent excursions elsewhere in the world, and to the disappearance of distinctive Ediacaran fossils (''Namacalathus'', ''Cloudina''). Nevertheless, there are arguments that the dated horizon in Oman does not correspond to the Ediacaran-Cambrian boundary, but represents a facies change from marine to evaporite-dominated strata – which would mean that dates from other sections, ranging from 544 or 542 Ma, are more suitable.


Paleogeography

Plate reconstructions suggest a global supercontinent,
Pannotia Pannotia (from Greek: '' pan-'', "all", '' -nótos'', "south"; meaning "all southern land"), also known as the Vendian supercontinent, Greater Gondwana, and the Pan-African supercontinent, was a relatively short-lived Neoproterozoic supercontinent ...
, was in the process of breaking up early in the Cambrian, with
Laurentia Laurentia or the North American Craton is a large continental craton that forms the ancient geological core of North America. Many times in its past, Laurentia has been a separate continent, as it is now in the form of North America, although ...
(North America), Baltica, and
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
having separated from the main supercontinent of
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
to form isolated land masses. Most continental land was clustered in the Southern Hemisphere at this time, but was drifting north. Large, high-velocity rotational movement of Gondwana appears to have occurred in the Early Cambrian. With a lack of sea ice – the great glaciers of the Marinoan
Snowball Earth The Snowball Earth hypothesis proposes that, during one or more of Earth's Greenhouse and icehouse Earth, icehouse Climate, climates, the Earth's surface, planet's surface became entirely or nearly entirely Freezing, frozen. It is believed that ...
were long melted – the sea level was high, which led to large areas of the continents being flooded in warm, shallow seas ideal for sea life. The sea levels fluctuated somewhat, suggesting there were "ice ages", associated with pulses of expansion and contraction of a south polar
ice cap In glaciology, an ice cap is a mass of ice that covers less than of land area (usually covering a highland area). Larger ice masses covering more than are termed ice sheets. Description Ice caps are not constrained by topographical features ...
. In
Baltoscandia Baltoscandian Confederation or Baltoscandia is a geopolitical concept of a Baltic–Scandinavian ( Nordic) union comprising Denmark, Estonia, Finland, Iceland, Latvia, Lithuania, Norway, and Sweden. The idea was proposed by a Swedish Professor ...
a Lower Cambrian
transgression Transgression may refer to: Legal, religious and social *Sin, a violation of God's Ten Commandments or other elements of God's moral law *Crime, legal transgression, usually created by a violation of social or economic boundary **In civil law ju ...
transformed large swathes of the
Sub-Cambrian peneplain The sub-Cambrian peneplain is an ancient, extremely flat, erosion surface (peneplain) that has been exhumed and exposed by erosion from under Cambrian strata over large swathes of Fennoscandia. Eastward, where this peneplain dips below Cambrian an ...
into an
epicontinental sea An inland sea (also known as an epeiric sea or an epicontinental sea) is a continental body of water which is very large and is either completely surrounded by dry land or connected to an ocean by a river, strait, or "arm of the sea". An inland s ...
.


Climate

The Earth was generally very warm during the early Cambrian; its climate was comparable to the hot greenhouse of the Late Cretaceous and Early Palaeogene, as evidenced by a maximum in continental weathering rates over the last 900 million years and the presence of tropical, lateritic palaeosols at high palaeolatitudes during this time. Nonetheless, glaciers likely existed during the earliest Cambrian at high and possibly even at middle palaeolatitudes, possibly due to the ancient continent of
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
covering the
South Pole The South Pole, also known as the Geographic South Pole, Terrestrial South Pole or 90th Parallel South, is one of the two points where Earth's axis of rotation intersects its surface. It is the southernmost point on Earth and lies antipod ...
and cutting off polar ocean currents. However, other authors believe these very early, pretrilobitic glacial deposits may not even be of Cambrian age at all but instead date back to the Neoproterozoic, an era characterised by numerous severe
icehouse Icehouse or ice house may refer to: * Ice house (building), a building where ice is stored * Ice shanty, a shelter for ice fishing also known as an ''Icehouse'' * Ice skating rink, a facility for ice skating. * Ice hockey arena, an area where ice ...
periods. Although molecular clock estimates suggest terrestrial plants may have first emerged during the Middle or Late Cambrian, the consequent large-scale removal of the
greenhouse gas A greenhouse gas (GHG or GhG) is a gas that Absorption (electromagnetic radiation), absorbs and Emission (electromagnetic radiation), emits radiant energy within the thermal infrared range, causing the greenhouse effect. The primary greenhouse ...
CO2 from the atmosphere through sequestration did not begin until the Ordovician. It became even warmer towards the end of the period, and sea levels rose dramatically. This warming trend continued into the Early Ordovician, which was characterised by an extremely hot global climate.


Flora

The Cambrian flora was little different from the Ediacaran. The principal taxa were the marine macroalgae '' Fuxianospira'', '' Sinocylindra'', and '' Marpolia''. No calcareous macroalgae are known from the period. No land plant (
embryophyte The Embryophyta (), or land plants, are the most familiar group of green plants that comprise vegetation on Earth. Embryophytes () have a common ancestor with green algae, having emerged within the Phragmoplastophyta clade of green algae as sist ...
) fossils are known from the Cambrian. However, biofilms and microbial mats were well developed on Cambrian tidal flats and beaches 500 mya., and microbes forming microbial Earth ecosystems, comparable with modern soil crust of desert regions, contributing to soil formation.


Oceanic life

The Cambrian explosion was a period of rapid multicellular growth. Most animal life during the Cambrian was aquatic.
Trilobite Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the At ...
s were once assumed to be the dominant life form at that time, but this has proven to be incorrect.
Arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s were by far the most dominant animals in the ocean, but trilobites were only a minor part of the total arthropod diversity. What made them so apparently abundant was their heavy armor reinforced by calcium carbonate (CaCO3), which fossilized far more easily than the fragile
chitin Chitin ( C8 H13 O5 N)n ( ) is a long-chain polymer of ''N''-acetylglucosamine, an amide derivative of glucose. Chitin is probably the second most abundant polysaccharide in nature (behind only cellulose); an estimated 1 billion tons of chit ...
ous exoskeletons of other arthropods, leaving numerous preserved remains. The period marked a steep change in the diversity and composition of Earth's biosphere. The
Ediacaran biota The Ediacaran (; formerly Vendian) biota is a Taxonomy (biology), taxonomic period classification that consists of all life forms that were present on Earth during the Ediacaran Period (). These were composed of enigmatic tubular and frond-sh ...
suffered a mass extinction at the start of the Cambrian Period, which corresponded with an increase in the abundance and complexity of burrowing behaviour. This behaviour had a profound and irreversible effect on the substrate which transformed the
seabed The seabed (also known as the seafloor, sea floor, ocean floor, and ocean bottom) is the bottom of the ocean. All floors of the ocean are known as 'seabeds'. The structure of the seabed of the global ocean is governed by plate tectonics. Most of ...
ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syste ...
s. Before the Cambrian, the sea floor was covered by microbial mats. By the end of the Cambrian, burrowing animals had destroyed the mats in many areas through
bioturbation Bioturbation is defined as the reworking of soils and sediments by animals or plants. It includes burrowing, ingestion, and defecation of sediment grains. Bioturbating activities have a profound effect on the environment and are thought to be a pr ...
. As a consequence, many of those organisms that were dependent on the mats became extinct, while the other species adapted to the changed environment that now offered new ecological niches. Around the same time there was a seemingly rapid appearance of representatives of all the mineralized
phyla Phyla, the plural of ''phylum'', may refer to: * Phylum, a biological taxon between Kingdom and Class * by analogy, in linguistics, a large division of possibly related languages, or a major language family which is not subordinate to another Phyl ...
, including the
Bryozoa Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about long, they have a special feeding structure called a l ...
, which were once thought to have only appeared in the Lower Ordovician. However, many of those phyla were represented only by stem-group forms; and since mineralized phyla generally have a benthic origin, they may not be a good proxy for (more abundant) non-mineralized phyla. While the early Cambrian showed such diversification that it has been named the Cambrian Explosion, this changed later in the period, when there occurred a sharp drop in biodiversity. About 515 million years ago, the number of species going extinct exceeded the number of new species appearing. Five million years later, the number of genera had dropped from an earlier peak of about 600 to just 450. Also, the
speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within ...
rate in many groups was reduced to between a fifth and a third of previous levels. 500 million years ago, oxygen levels fell dramatically in the oceans, leading to
hypoxia Hypoxia means a lower than normal level of oxygen, and may refer to: Reduced or insufficient oxygen * Hypoxia (environmental), abnormally low oxygen content of the specific environment * Hypoxia (medical), abnormally low level of oxygen in the tis ...
, while the level of poisonous
hydrogen sulfide Hydrogen sulfide is a chemical compound with the formula . It is a colorless chalcogen-hydride gas, and is poisonous, corrosive, and flammable, with trace amounts in ambient atmosphere having a characteristic foul odor of rotten eggs. The unde ...
simultaneously increased, causing another extinction. The later half of Cambrian was surprisingly barren and showed evidence of several rapid extinction events; the stromatolites which had been replaced by reef building sponges known as Archaeocyatha, returned once more as the archaeocyathids became extinct. This declining trend did not change until the
Great Ordovician Biodiversification Event The Great Ordovician Biodiversification Event (GOBE), was an evolutionary radiation of animal life throughout the Ordovician period, 40 million years after the Cambrian explosion, whereby the distinctive Cambrian fauna fizzled out to be replaced wi ...
. Some Cambrian organisms ventured onto land, producing the trace fossils ''
Protichnites ''Protichnites'' is an ichnogenus of trace fossil consisting of the imprints made by the walking activity of certain arthropods. It consists of two rows of tracks and a medial furrow between the two rows. This furrow, which may be broken, set at ...
'' and ''
Climactichnites ''Climactichnites'' is an enigmatic, Cambrian fossil formed on or within sandy tidal flats around . It has been interpreted in many different ways in the past, but is now thought to be a trace fossil of a slug-like organism that moved by crawling ...
''. Fossil evidence suggests that
euthycarcinoid Euthycarcinoidea are an enigmatic group of extinct possibly amphibious arthropods that ranged from Cambrian to Triassic times. Fossils are known from Europe, North America, Argentina, Australia and Antarctica. Description The euthycarcinoid body ...
s, an extinct group of arthropods, produced at least some of the ''Protichnites''. Fossils of the track-maker of ''Climactichnites'' have not been found; however, fossil trackways and resting traces suggest a large,
slug Slug, or land slug, is a common name for any apparently shell-less terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced shell, or only a smal ...
-like
mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is esti ...
. In contrast to later periods, the Cambrian fauna was somewhat restricted; free-floating organisms were rare, with the majority living on or close to the sea floor; and mineralizing animals were rarer than in future periods, in part due to the unfavourable ocean chemistry. Many modes of preservation are unique to the Cambrian, and some preserve soft body parts, resulting in an abundance of . These include Sirius Passet, the Sinsk Algal Lens, the Maotianshan Shales, the Emu Bay Shale, and the Burgess Shale,.


Symbol

The United States
Federal Geographic Data Committee The Federal Geographic Data Committee (FGDC) is a United States government committee which promotes the coordinated development, use, sharing, and dissemination of geospatial data on a national basis. Its 32 members are representatives from the E ...
uses a "barred capital C" character to represent the Cambrian Period. The
Unicode Unicode, formally The Unicode Standard,The formal version reference is is an information technology Technical standard, standard for the consistent character encoding, encoding, representation, and handling of Character (computing), text expre ...
character is .Unicode Character 'LATIN CAPITAL LETTER C WITH BAR' (U+A792)
fileformat.info. Retrieved 15 June 2015


Gallery

File:CambrianStromatolites.jpg, Stromatolites of the Pika Formation (Middle Cambrian) near Helen Lake, Banff National Park, Canada File:Elrathia kingii growth series.jpg,
Trilobite Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the At ...
s were very common during this time File:20191203 Anomalocaris canadensis.png, ''
Anomalocaris ''Anomalocaris'' ("unlike other shrimp", or "abnormal shrimp") is an extinct genus of Radiodonta, radiodont, an Order (biology), order of early-diverging stem-group arthropods. The first fossils of ''Anomalocaris'' were discovered in the ''Ogygop ...
'' was an early marine predator, among the various
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s of the time File:20191108 Opabinia regalis.png, ''
Opabinia ''Opabinia regalis'' is an extinct, stem group arthropod found in the Middle Cambrian Burgess Shale Lagerstätte (505 million years ago) of British Columbia. ''Opabinia'' was a soft-bodied animal, measuring up to 7 cm in body length, and it ...
'' was a creature with an unusual body plan; it was probably related to arthropods File:Pikaia NT small.jpg, '' Pikaia'' was a stem-chordate from the Middle Cambrian File:Protichnites, Blackberry Hill, Wisconsin, Cambrian 2.jpg, ''
Protichnites ''Protichnites'' is an ichnogenus of trace fossil consisting of the imprints made by the walking activity of certain arthropods. It consists of two rows of tracks and a medial furrow between the two rows. This furrow, which may be broken, set at ...
'' were the trackways of arthropods that walked Cambrian beaches File:20210830 Hallucigenia sparsa diagrammatic reconstruction.png, ''
Hallucigenia sparsa ''Hallucigenia'' is a genus of Cambrian animal resembling worms, known from articulated fossils in Burgess Shale-type deposits in Canada and China, and from isolated spines around the world. The generic name reflects the type species' unusual ...
'' was a member of group
lobopodian The lobopodians, members of the informal group Lobopodia (from the Greek language, Greek, meaning "blunt feet"), or the formally erected phylum Lobopoda Cavalier-Smith (1998), are panarthropods with stubby legs called lobopods, a term which may ...
, that is considered to related to modern
velvet worms Onychophora (from grc, ονυχής, , "claws"; and , , "to carry"), commonly known as velvet worms (due to their velvety texture and somewhat wormlike appearance) or more ambiguously as peripatus (after the first described genus, '' Peripatus ...
. File:20200329 Cambroraster falcatus.png, ''
Cambroraster ''Cambroraster'' is an extinct monotypic genus of hurdiid radiodont, dating to the middle Cambrian, and represented by the single formally described species ''Cambroraster falcatus''. Hundreds of specimens were found in the Burgess Shale, and des ...
falcatus'' was a large arthropod for the era


See also

* Cambrian–Ordovician extinction event – circa 488 mya * Dresbachian extinction event—circa 499 mya * End Botomian extinction event—circa 513 mya * List of fossil sites ''(with link directory)'' *
Type locality (geology) Type locality, also called type area, is the locality where a particular rock type, stratigraphic unit or mineral species is first identified. If the stratigraphic unit in a locality is layered, it is called a stratotype, whereas the standard o ...
, the locality where a particular rock type, stratigraphic unit, fossil or mineral species is first identified


References


Further reading

* * * * * * * * * * *


External links

*
Biostratigraphy
– includes information on Cambrian trilobite
biostratigraphy Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock Stratum, strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictiona ...

Dr. Sam Gon's trilobite pages
(contains numerous Cambrian trilobites)


Paleomap Project

Report on the web on Amthor and others from ''Geology'' vol. 31




{{Authority control Geological periods