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Cambrian Substrate Revolution
The "Cambrian substrate revolution" or "Agronomic revolution", evidenced in trace fossils, is the diversification of animal burrowing during the early Cambrian period. Before this "widening of the behavioural repertoire", bottom-dwelling animals mainly grazed on the microbial mats that lined the surface, crawling above (like how freshwater snails still do) or burrowing just below them. These microbial mats created a barrier between the water and the sediment underneath, which was less water-logged than modern sea-floors, and almost completely anoxic (lacking in oxygen). As a result, the substrate was inhabited by sulfate-reducing bacteria, whose emissions of hydrogen sulfide (H2S) made the substrate toxic to most other organisms. Around the start of the Cambrian, organisms began to burrow vertically, forming a great diversity of different fossilisable burrow forms as they penetrated the sediment for protection or to feed. These burrowing animals broke down the microbial mats, ...
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Trace Fossil
A trace fossil, also known as an ichnofossil (; from el, ἴχνος ''ikhnos'' "trace, track"), is a fossil record of biological activity but not the preserved remains of the plant or animal itself. Trace fossils contrast with body fossils, which are the fossilized remains of parts of organisms' bodies, usually altered by later chemical activity or mineralization. The study of such trace fossils is ichnology and is the work of ichnologists. Trace fossils may consist of impressions made on or in the substrate by an organism. For example, burrows, borings (bioerosion), urolites (erosion caused by evacuation of liquid wastes), footprints and feeding marks and root cavities may all be trace fossils. The term in its broadest sense also includes the remains of other organic material produced by an organism; for example coprolites (fossilized droppings) or chemical markers (sedimentological structures produced by biological means; for example, the formation of stromatolites). H ...
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Skolithos
''Skolithos'' (formerly spelled ''Scolithus'' or ''Skolithus'') is a common trace fossil ichnogenus that is, or was originally, an approximately vertical cylindrical burrow. It is produced by a variety of organisms in shallow marine environments globally and appear as lineated features in sedimentary rocks. Depositional environments ''Skolithos'' burrows range in age from early Cambrian to the present and are found throughout the world. They occur in sediments and sedimentary rocks, primarily sands and sandstones. They are typically marine in origin, and are commonly associated with high-energy environments close to the shoreline. They have also been reported from freshwater lacustrine settings, but have rarely been described from carbonate rocks. Vertical ''Skolithos'' burrows can also occur in alluvial sediments such as braided river deposits, where the periodic fluctuation of water is an important factor in the formation of this structure.  This periodic water fluctuation ...
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Helicoplacus
''Helicoplacus'' (often misspelled ''Helioplacus'') is the earliest well-studied fossil echinoderm. Fossil plates are known from several regions. Complete specimens were found in Lower Cambrian strata of the White Mountains of California. The animal was a cigar-shaped creature up to long that stood upright on one end. Unlike more typical echinoderms such as sea stars, ''Helicoplacus'' does not have fivefold symmetry. Instead, there is a spiral food groove on the outside along which food was moved to a mouth that is thought to be on the side. The respiratory system appears to be primitive. Although the animal does not look like a typical echinoderm, the plates are composed of the characteristic calcareous plates known as '' stereom'' that are common to all echinoderms. The ambulacrum is similar to that of the Edrioasteroidea; as a result, ''Helicoplacus'' may belong to Pelmatozoa. Other contemporaneous echinoderms are known to have existed from their dissociated plates, but other ...
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Echinoderm
An echinoderm () is any member of the phylum Echinodermata (). The adults are recognisable by their (usually five-point) radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the sea lilies or "stone lilies". Adult echinoderms are found on the sea bed at every ocean depth, from the intertidal zone to the abyssal zone. The phylum contains about 7,000 living species, making it the second-largest grouping of deuterostomes, after the chordates. Echinoderms are the largest entirely marine phylum. The first definitive echinoderms appeared near the start of the Cambrian. The echinoderms are important both ecologically and geologically. Ecologically, there are few other groupings so abundant in the biotic desert of the deep sea, as well as shallower oceans. Most echinoderms are able to reproduce asexually and regenerate tissue, organs, and limbs; in some cases, they can undergo complete regeneration from a single limb. ...
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Sessility (zoology)
Sessility is the biological property of an organism describing its lack of a means of self-locomotion. Sessile organisms for which natural ''motility'' is absent are normally immobile. This is distinct from the botanical concept of sessility, which refers to an organism or biological structure attached directly by its base without a stalk. Sessile organisms can move via external forces (such as water currents), but are usually permanently attached to something. Organisms such as corals lay down their own substrate from which they grow. Other sessile organisms grow from a solid such as a rock, dead tree trunk, or a man-made object such as a buoy or ship's hull. Mobility Sessile animals typically have a motile phase in their development. Sponges have a motile larval stage and become sessile at maturity. Conversely, many jellyfish develop as sessile polyps early in their life cycle. In the case of the cochineal, it is in the nymph stage (also called the crawler stage) that the ...
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Holdfast (biology)
A holdfast is a root-like structure that anchors aquatic sessile organisms, such as seaweed, other sessile algae, stalked crinoids, benthic cnidarians, and sponges, to the substrate. Holdfasts vary in shape and form depending on both the species and the substrate type. The holdfasts of organisms that live in muddy substrates often have complex tangles of root-like growths. These projections are called haptera and similar structures of the same name are found on lichens. The holdfasts of organisms that live in sandy substrates are bulb-like and very flexible, such as those of sea pens, thus permitting the organism to pull the entire body into the substrate when the holdfast is contracted. The holdfasts of organisms that live on smooth surfaces (such as the surface of a boulder) have flattened bases which adhere to the surface. The organism derives no nutrition from this intimate contact with the substrate, as the process of liberating nutrients from the substrate requires ...
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Bioturbation
Bioturbation is defined as the reworking of soils and sediments by animals or plants. It includes burrowing, ingestion, and defecation of sediment grains. Bioturbating activities have a profound effect on the environment and are thought to be a primary driver of biodiversity. The formal study of bioturbation began in the 1800s by Charles Darwin experimenting in his garden. The disruption of aquatic sediments and terrestrial soils through bioturbating activities provides significant ecosystem services. These include the alteration of nutrients in aquatic sediment and overlying water, shelter to other species in the form of burrows in terrestrial and water ecosystems, and soil production on land.Shaler, N. S., 1891, The origin and nature of soils, in Powell, J. W., ed., USGS 12th Annual report 1890-1891: Washington, D.C., Government Printing Office, p. 213-45. Bioturbators are deemed ecosystem engineers because they alter resource availability to other species through the physical ch ...
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Fauna
Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is '' funga''. Flora, fauna, funga and other forms of life are collectively referred to as '' biota''. Zoologists and paleontologists use ''fauna'' to refer to a typical collection of animals found in a specific time or place, e.g. the "Sonoran Desert fauna" or the "Burgess Shale fauna". Paleontologists sometimes refer to a sequence of faunal stages, which is a series of rocks all containing similar fossils. The study of animals of a particular region is called faunistics. Etymology ''Fauna'' comes from the name Fauna, a Roman goddess of earth and fertility, the Roman god Faunus, and the related forest spirits called Fauns. All three words are cognates of the name of the Greek god Pan, and ''panis'' is the Greek equivalent of fauna. ''Fauna'' is also the word for a book that catalogues the animals in such a manner. The term was first used b ...
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Anoxic Waters
Anoxic waters are areas of sea water, fresh water, or groundwater that are depleted of dissolved oxygen. The US Geological Survey defines anoxic groundwater as those with dissolved oxygen concentration of less than 0.5 milligrams per litre. Anoxic waters can be contrasted with hypoxic waters, which are low (but not lacking) in dissolved oxygen. This condition is generally found in areas that have restricted water exchange. In most cases, oxygen is prevented from reaching the deeper levels by a physical barrier, as well as by a pronounced density stratification, in which, for instance, heavier hypersaline waters rest at the bottom of a basin. Anoxic conditions will occur if the rate of oxidation of organic matter by bacteria is greater than the supply of dissolved oxygen. Anoxic waters are a natural phenomenon, and have occurred throughout geological history. The Permian–Triassic extinction event, a mass extinction of species from the world's oceans, may have resulted from w ...
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Predation
Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the host) and parasitoidism (which always does, eventually). It is distinct from scavenging on dead prey, though many predators also scavenge; it overlaps with herbivory, as seed predators and destructive frugivores are predators. Predators may actively search for or pursue prey or wait for it, often concealed. When prey is detected, the predator assesses whether to attack it. This may involve ambush or pursuit predation, sometimes after stalking the prey. If the attack is successful, the predator kills the prey, removes any inedible parts like the shell or spines, and eats it. Predators are adapted and often highly specialized for hunting, with acute senses such as vision, hearing, or smell. Many predatory animals, both vertebrate and inv ...
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Exoskeletons
An exoskeleton (from Greek ''éxō'' "outer" and ''skeletós'' "skeleton") is an external skeleton that supports and protects an animal's body, in contrast to an internal skeleton (endoskeleton) in for example, a human. In usage, some of the larger kinds of exoskeletons are known as " shells". Examples of exoskeletons within animals include the arthropod exoskeleton shared by chelicerates, myriapods, crustaceans, and insects, as well as the shell of certain sponges and the mollusc shell shared by snails, clams, tusk shells, chitons and nautilus. Some animals, such as the turtle, have both an endoskeleton and an exoskeleton. Role Exoskeletons contain rigid and resistant components that fulfill a set of functional roles in many animals including protection, excretion, sensing, support, feeding and acting as a barrier against desiccation in terrestrial organisms. Exoskeletons have a role in defense from pests and predators, support and in providing an attachment framework ...
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