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Calyptratae is a subsection of
Schizophora The Schizophora are a section of true flies containing 78 families, which are collectively referred to as muscoids, although technically the term "muscoid" should be limited to flies in the superfamily Muscoidea; this is an example of informal, hi ...
in the
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
order
Diptera Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced ...
, commonly referred to as the calyptrate muscoids (or simply calyptrates). It consists of those flies which possess a
calypter A calypter is either of two posterior lobes of the posterior margin of the forewing of flies between the extreme posterior wing base and the alula, which covers the halteres. The lower calypter is the proximal calypter (synonyms: squama (of som ...
that covers the
haltere ''Halteres'' (; singular ''halter'' or ''haltere'') (from grc, ἁλτῆρες, weights held in the hands to give an impetus in leaping) are a pair of small club-shaped organs on the body of two orders of flying insects that provide infor ...
s, among which are some of the most familiar of all flies, such as the
house fly The housefly (''Musca domestica'') is a fly of the suborder Cyclorrhapha. It is believed to have evolved in the Cenozoic Era, possibly in the Middle East, and has spread all over the world as a commensal of humans. It is the most common fly ...
. About 18,000 described species are in this group, or about 12% of all the flies yet described.


Subsection

*Superfamily
Muscoidea Muscoidea is a superfamily of flies in the subsection Calyptratae. Muscoidea, with approximately 7000 described species, is nearly 5% of the known species level diversity of the Diptera, the true flies. Most muscoid flies are saprophagous, coproph ...
*:
Anthomyiidae The Anthomyiidae are a large and diverse family of Muscoidea flies. Most look rather like small houseflies, but are commonly drab grey. The genus ''Anthomyia'', in contrast, is generally conspicuously patterned in black-and-white or black-and- ...
- cabbage flies *:
Fanniidae The Fanniidae are a small (285 species in five genera) group of true flies largely confined to the Holarctic and temperate Neotropical realms; there are 11 Afrotropical species, 29 Oriental, and 14 Australasian. Adults are medium-sized to small ...
*:
Muscidae Muscidae are a family of flies found in the superfamily Muscoidea. Muscidae, some of which are commonly known as house flies or stable flies due to their synanthropy, are worldwide in distribution and contain almost 4,000 described species i ...
- house flies *:
Scathophagidae The Scathophagidae are a small family of Muscoidea which are often known as dung flies, although this name is not appropriate except for a few species of the genus ''Scathophaga'' which do indeed pass their larval stages in animal dung. The nam ...
- dung flies *Superfamily
Oestroidea Oestroidea is a superfamily of Calyptratae including the blow flies, bot flies, flesh flies, and their relatives. It occurs worldwide and has about 15,000 described species. The superfamily includes the families: *Calliphoridae *Mesembrinellidae ...
*:
Calliphoridae The Calliphoridae (commonly known as blow flies, blow-flies, carrion flies, bluebottles, greenbottles, or cluster flies) are a family of insects in the order Diptera, with almost 1,900 known species. The maggot larvae, often used as fishing ba ...
*:
Mystacinobiidae The New Zealand bat fly (''Mystacinobia zelandica'') is a small, wingless insect which lives in a commensal relationship with the New Zealand lesser short-tailed bat. It is a true fly, in the order Diptera, placed in its own genus, ''Mystacinobia ...
*:
Oestridae Botflies, also known as warble flies, heel flies, and gadflies, are a family of flies known as the Oestridae. Their larvae are internal parasites of mammals, some species growing in the host's flesh and others within the gut. ''Dermatobia homin ...
*:
Rhinophoridae Rhinophoridae is a family of flies (Diptera), commonly known as Woodlouse Flies, found in all zoogeographic regions except Oceania, but mainly in the Palaearctic and Afrotropical regions. They are small, slender, black, bristly flies phylog ...
*:
Sarcophagidae Sarcophagidae () are a family of flies commonly known as flesh flies. They differ from most flies in that they are ovoviviparous, opportunistically depositing hatched or hatching maggots instead of eggs on carrion, dung, decaying material, or op ...
*:
Tachinidae The Tachinidae are a large and variable family of true flies within the insect order Diptera, with more than 8,200 known species and many more to be discovered. Over 1,300 species have been described in North America alone. Insects in this family ...
*:
Ulurumyiidae ''Ulurumyia macalpinei'' is a species of fly in the superfamily Oestroidea endemic to Australia. It was first discovered in the 1970s, but was not described until 2017, and in the intervening decades was informally known to entomologists as McAlp ...
*Superfamily
Hippoboscoidea Hippoboscoidea is a superfamily of the Calyptratae. The flies in this superfamily are blood-feeding obligate parasites of their hosts. Four families are often placed here: * Glossinidae - Tsetse flies *Hippoboscidae - Ked flies *Nycteribiidae - B ...
*: Glossinidae *:
Hippoboscidae __NOTOC__ Hippoboscidae, the louse flies or keds, are obligate parasites of mammals and birds. In this family, the winged species can fly at least reasonably well, though others with vestigial or no wings are flightless and highly apomorphic. As ...
*:
Nycteribiidae Nycteribiidae is a family of the true fly superfamily Hippoboscoidea are known as "bat flies", together with their close relatives the Streblidae. As the latter do not seem to be a monophyletic group, it is conceivable that bat flies cannot be u ...
*:
Streblidae The Streblidae are a family of flies in the superfamily Hippoboscoidea, and together with their relatives the Nycteribiidae, are known as bat flies. They are winged or wingless ectoparasites of bats, and often have long legs. They appear to be ho ...
The
Mormotomyiidae The family Mormotomyiidae (Diptera: Ephydroidea) contains only one known species, ''Mormotomyia hirsuta'', commonly known as the frightful hairy flyKirk-Spriggs, A.H., Kotrba, M. & Copeland, R.S. 2011. Further details of the morphology of the en ...
belong to the
Ephydroidea The Ephydroidea are a superfamily of muscomorph flies, with over 6,000 species. Description A characteristic of adult Ephydroidea (shared with their relatives such as Calyptratae) is that the pedicel of the antenna has a dorsoventral seam or ...
and not to Hippoboscoidea as previously construed. The Streblidae are probably not
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
.Petersen, Frederik Torp; Meier, Rudolf; Kutty, Sujatha Narayanan & Wiegmann, Brian M. (2007): The phylogeny and evolution of host choice in the Hippoboscoidea (Diptera) as reconstructed using four molecular markers. '' Mol. Phylogenet. Evol.'' 45(1): 111–122. (HTML abstract)


References

{{Calyptratae-stub