''Calycopis pisis'', the pisis groundstreak, is a butterfly found in several countries in Latin America.

Taxonomic history and synonyms

This species was described in 1887 by

Frederick DuCane Godman Frederick DuCane Godman DCL FRS FLS FGS FRGS FES FZS MRI FRHS (15 January 1834 – 19 February 1919) was an English lepidopterist, entomologist and ornithologist. He was one of the twenty founding members of the British Ornithologists' Un ...

and

Osbert Salvin Osbert Salvin FRS (25 February 1835 – 1 June 1898) was an English naturalist, ornithologist, and herpetologist best known for co-authoring ''Biologia Centrali-Americana'' (1879–1915) with Frederick DuCane Godman. This was a 52 volume encyc ...

in ''

Biologia Centrali-Americana The ''Biologia Centrali-Americana'' is an encyclopedia of the natural history of Mexico and Central America, privately issued in 215 parts from 1879 to 1915 by the editors Frederick DuCane Godman and Osbert Salvin, of the British Museum (Natural ...

''. They placed it in the genus ''

Thecla Thecla ( grc, Θέκλα, ) was a saint of the early Christian Church, and a reported follower of Paul the Apostle. The earliest record of her life comes from the ancient apocryphal ''Acts of Paul and Thecla''. Church tradition The ''Acts of ...

''. In 1969, William D. Field transferred the species to his newly-circumscribed genus ''Calystryma''. Then, in 1991, Kurt Johnson transferred it his newly-circumscribed genus ''Klaufera'' to be its

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

. In the same paper, Johnson described a species he named ''Serratoterga larsoni'' to honor the American cartoonist

Gary Larson Gary Larson (born August 14, 1950) is an American cartoonist, environmentalist, and former musician. He is the creator of ''The Far Side'', a single-panel cartoon series that was syndicated internationally to more than 1,900 newspapers for fif ...

; this was the type species of ''Serratoterga''. In 2004, R. K. Robins

synonym A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...

ized the two species and transferred it to the genus ''

Calycopis ''Calycopis'' is a genus of butterflies in the family Lycaenidae. Massively split up by Kurt Johnson in 1991, most modern authors consider the changes proposed at that time to be unjustified. Most of the species of this genus are found in the N ...

'', making ''C. larsoni'' an

invalid Invalid may refer to: * Patient, a sick person * one who is confined to home or bed because of illness, disability or injury (sometimes considered a politically incorrect term) * .invalid, a top-level Internet domain not intended for real use As t ...

junior synonym.

Distribution

It has been found in several parts of Latin America, including Guatemala, Nicaragua, Panama, Honduras, Ecuador, Belize, Mexico (in the states of

Veracruz Veracruz (), formally Veracruz de Ignacio de la Llave (), officially the Free and Sovereign State of Veracruz de Ignacio de la Llave ( es, Estado Libre y Soberano de Veracruz de Ignacio de la Llave), is one of the 31 states which, along with Me ...

and

Oaxaca Oaxaca ( , also , , from nci, Huāxyacac ), officially the Free and Sovereign State of Oaxaca ( es, Estado Libre y Soberano de Oaxaca), is one of the 32 states that compose the political divisions of Mexico, Federative Entities of Mexico. It is ...

) and in

northern Brazil The North Region of Brazil ( pt, Região Norte do Brasil; ) is the largest region of Brazil, corresponding to 45.27% of the national territory. It is the second least inhabited of the country, and contributes with a minor percentage in the national ...

. H. L. Lewis listed a specimen of ''C. pisis'' as being from the island of

Trinidad Trinidad is the larger and more populous of the two major islands of Trinidad and Tobago. The island lies off the northeastern coast of Venezuela and sits on the continental shelf of South America. It is often referred to as the southernmos ...

, but other lepidopterists have said they believe this to be an error. A type locality was not initially specified, but the specimen designated as the

lectotype In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the ...

was collected in Bugaba,

Chiriquí Province Chiriquí () is a province of Panama located on the western coast; it is the second most developed province in the country, after the Panamá Province. Its capital is the city of David. It has a total area of 6,490.9 km², with a population ...

, Panama. The type locality for the junior synonym ''C. larsoni'' is

Santo Domingo, Ecuador Santo Domingo de los Colorados, often simply referred to as Santo Domingo (Quechua: Tsachila), is an Ecuadorian city and seat of the canton that bears its name and the Santo Domingo de los Tsáchilas Province. It is the fourth most populous city ...

Description

The

forewing Insect wings are adult outgrowths of the insect exoskeleton that enable insects to fly. They are found on the second and third thoracic segments (the mesothorax and metathorax), and the two pairs are often referred to as the forewings and hindwin ...

s measure for the males and for the female. The wings' underside for both sexes are the same: they are a pale brownish gray with a slight yellow iridescence and with a pattern of red, eye-like markings on the

hindwing Insect wings are adult outgrowths of the insect exoskeleton that enable insects to fly. They are found on the second and third thoracic segments (the mesothorax and metathorax), and the two pairs are often referred to as the forewings and hindwi ...

around the M₃ and Cu₁ cells (see the

Comstock–Needham system The Comstock–Needham system is a naming system for insect wing veins, devised by John Comstock and George Needham in 1898. It was an important step in showing the homology of all insect wings. This system was based on Needham's ''pretracheati ...

insect wing Insect wings are adult outgrowths of the insect exoskeleton that enable insects to fly. They are found on the second and third thoracic segments (the mesothorax and metathorax), and the two pairs are often referred to as the forewings and hindwin ...

nomenclature). The upper surface of the male wings are a purple metallic color. The upper surface of the female forewing is dark brown with some blue tint near the

basal area Basal area is the cross-sectional area of trees at breast height (1.3m or 4.5 ft above ground). It is a common way to describe stand density. In forest management, basal area usually refers to merchantable timber and is given on a per hectar ...

. The upper surface of the female hindwings are also dark brown with iridescent blue, with dark brown spots in the M₃ and Cu₁ cells.

Notes

References

External links

* {{Taxonbar, from=Q25804809 Butterflies of Central America Lycaenidae of South America Butterflies described in 1887 Taxa named by Frederick DuCane Godman Taxa named by Osbert Salvin Calycopis