Bathyspondylus
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''Bathyspondylus'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
plesiosaur The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared ...
from the
Kimmeridge Clay Formation The Kimmeridge Clay is a sedimentary deposit of fossiliferous marine clay which is of Late Jurassic to lowermost Cretaceous age and occurs in southern and eastern England and in the North Sea. This rock formation is the major source rock for Nort ...
of
Swindon Swindon () is a town and unitary authority with Borough status in the United Kingdom, borough status in Wiltshire, England. As of the 2021 Census, the population of Swindon was 201,669, making it the largest town in the county. The Swindon un ...
,
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
. Because it is known only from its
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
vertebrae (and so few of those have been recovered),
paleontologists Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
are not entirely sure of the
taxonomy Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
of ''Bathyspondylus''; the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
it belongs to is not currently known.Cladistic Analysis of the Plesiosauria (Reptilia: Sauropterygia)Adam Stuart Smith, Supervised by M. J. Benton. University of Bristol, Department of Earth Sciences. September 2003. pg. 18 The type, and only known,
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
is ''B. swindoniensis'', which was described from the same material as its genus.


Etymology

The
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
name ''Bathyspondylus'' is a compound of two
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
roots: βαθυς (''bathys'') 'deep' and σπονδυλος (''spondylos'') '
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
'. It can thus be translated as "deep-vertebrae". The species name ''B. swindoniensis'' refers to the town of
Swindon Swindon () is a town and unitary authority with Borough status in the United Kingdom, borough status in Wiltshire, England. As of the 2021 Census, the population of Swindon was 201,669, making it the largest town in the county. The Swindon un ...
in
Wiltshire Wiltshire (; abbreviated Wilts) is a historic and ceremonial county in South West England with an area of . It is landlocked and borders the counties of Dorset to the southwest, Somerset to the west, Hampshire to the southeast, Gloucestershire ...
, near which the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen was discovered.


Discovery and naming

The holotype, a set of ?21+
cervical vertebrae In tetrapods, cervical vertebrae (singular: vertebra) are the vertebrae of the neck, immediately below the skull. Truncal vertebrae (divided into thoracic and lumbar vertebrae in mammals) lie caudal (toward the tail) of cervical vertebrae. In ...
, was discovered in
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
(
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...
)-aged
Kimmeridge Clay Formation The Kimmeridge Clay is a sedimentary deposit of fossiliferous marine clay which is of Late Jurassic to lowermost Cretaceous age and occurs in southern and eastern England and in the North Sea. This rock formation is the major source rock for Nort ...
, outcropping at the Great Western Railway Works in
Swindon Swindon () is a town and unitary authority with Borough status in the United Kingdom, borough status in Wiltshire, England. As of the 2021 Census, the population of Swindon was 201,669, making it the largest town in the county. The Swindon un ...
,
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
, in 1774 and the holotype was at one point in time part of the collection of William Cunnington III sometime before his death in 1810. It was probably part of the
Devizes Museum The Wiltshire Museum, formerly known as Wiltshire Heritage Museum and Devizes Museum, is a museum, archive and library and art gallery in Devizes, Wiltshire, England. The museum was established and is run by the Wiltshire Archaeological and Natu ...
collection by 1888, where it is now housed. It was named in 1982 by J. B. Delair.J. B. Delair. (1982). New and little-known Jurassic reptiles from Wiltshire. ''The Wiltshire Archaeological and Natural History Magazine'' 76:155-164


Description

''Bathyspondylus'' had centra set fairly deep in the
vertebrae The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic i ...
relative to its length, as its name (the Greek words for ''deep-vertebrae'') would suggest. The vertebrae themselves are short
antero-posteriorly Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
and can be flat or
concave Concave or concavity may refer to: Science and technology * Concave lens * Concave mirror Mathematics * Concave function, the negative of a convex function * Concave polygon, a polygon which is not convex * Concave set * The concavity In ca ...
on their terminal faces. The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen, from 1774, appears to have features of both
pliosauroids Pliosauroidea is an extinct clade of plesiosaurs, known from the earliest Jurassic to early Late Cretaceous. They are best known for the subclade Thalassophonea, which contained crocodile-like short-necked forms with large heads and massive to ...
and
plesiosauroids Plesiosauroidea (; Greek: 'near, close to' and 'lizard') is an extinct clade of carnivorous marine reptiles. They have the snake-like longest neck to body ratio of any reptile. Plesiosauroids are known from the Jurassic and Cretaceous p ...
incorporated into its bones.


Distribution

''Bathyspondylus'' lived during the
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...
faunal stage In chronostratigraphy, a stage is a succession of rock strata laid down in a single age on the geologic timescale, which usually represents millions of years of deposition. A given stage of rock and the corresponding age of time will by convent ...
of the
Jurassic period The Jurassic ( ) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period of ...
, which occurred roughly 155 to 150 million years ago. The first of its fossils came from deposits near Swindon in Wiltshire,
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
.


See also

*
Timeline of plesiosaur research This timeline of plesiosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, taxonomic revisions, and cultural portrayals of plesiosaurs, an order of marine reptiles that flourished duri ...
*
List of plesiosaur genera This list of plesiosaurs is a comprehensive listing of all genera that have ever been included in the order Plesiosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered inv ...


References


External links

*https://web.archive.org/web/20071210085607/http://www.plesiosaur.com/database/genusIndividual.php?i=65 *https://web.archive.org/web/20091001060613/http://www.dinosauria.com/dml/names/ples.html {{Taxonbar, from=Q4120085 Late Jurassic plesiosaurs of Europe Plesiosauria incertae sedis Fossil taxa described in 1982 Sauropterygian genera