''Buellia spuria'', the disc lichen, is a white to light ashy gray

crustose Crustose is a habit of some types of algae and lichens in which the organism grows tightly appressed to a substrate, forming a biological layer. ''Crustose'' adheres very closely to the substrates at all points. ''Crustose'' is found on rocks and ...

areolate Lichens are composite organisms made up of multiple species: a fungal partner, one or more photosynthetic partners, and sometimes a basidiomycete yeast. They are regularly grouped by their external appearance – a characteristic known as their ...

lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.epilithic Lithophytes are plants that grow in or on rocks. They can be classified as either epilithic (or epipetric) or endolithic; epilithic lithophytes grow on the surfaces of rocks, while endolithic lithophytes grow in the crevices of rocks (and are also ...

) in

montane Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...

habitats.Nash, T.H., Ryan, B.D., Gries, C., Bugartz, F., (eds.) 2001. Lichen Flora of the Greater Sonoran Desert Region. Vol 3

/ref> It has a black edge from the conspicuous, more or less continuous

prothallus A prothallus, or prothallium, (from Latin ''pro'' = forwards and Greek ''θαλλος'' (''thallos'') = twig) is usually the gametophyte stage in the life of a fern or other pteridophyte. Occasionally the term is also used to describe the young ...

, which can also be seen in the cracks between the areolas forming a

hypothallus In true slime molds (myxogastria), lichens, and in species of the family Clavicipitaceae, the hypothallus is the layer on which the fruit body sits, lying in contact with the substrate. The word is derived from the Ancient Greek root ''hypó'' ("u ...

, and in sharp contrast with the whitish or ashy colored areolas. It prefers

mafic A mafic mineral or rock is a silicate mineral or igneous rock rich in magnesium and iron. Most mafic minerals are dark in color, and common rock-forming mafic minerals include olivine, pyroxene, amphibole, and biotite. Common mafic rocks include ...

(

siliceous Silicon dioxide, also known as silica, is an oxide of silicon with the chemical formula , most commonly found in nature as quartz and in various living organisms. In many parts of the world, silica is the major constituent of sand. Silica is one ...

) rock substrates. In

Joshua Tree National Park Joshua Tree National Park is an American national park in southeastern California, east of San Bernardino and Los Angeles and north of Palm Springs, California, Palm Springs. It is named after the Joshua trees (''Yucca brevifolia'') native to t ...

is can be seen on vertical

granite Granite () is a coarse-grained (phaneritic) intrusive igneous rock composed mostly of quartz, alkali feldspar, and plagioclase. It forms from magma with a high content of silica and alkali metal oxides that slowly cools and solidifies undergro ...

and

gneiss Gneiss ( ) is a common and widely distributed type of metamorphic rock. It is formed by high-temperature and high-pressure metamorphic processes acting on formations composed of igneous or sedimentary rocks. Gneiss forms at higher temperatures an ...

faces in washes.Photo Gallery, Joshua Tree Lichens, Joshua Tree National Park website, National Park Service

/ref> It is common worldwide in the Northern Hemisphere. It is very common in the

Sonoran Desert The Sonoran Desert ( es, Desierto de Sonora) is a desert in North America and ecoregion that covers the northwestern Mexican states of Sonora, Baja California, and Baja California Sur, as well as part of the southwestern United States (in Arizona ...

from southern

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...

Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...

Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...

, and

Sonora Sonora (), officially Estado Libre y Soberano de Sonora ( en, Free and Sovereign State of Sonora), is one of the 31 states which, along with Mexico City, comprise the Administrative divisions of Mexico, Federal Entities of Mexico. The state is d ...

, Chihuahua, and

Sinaloa Sinaloa (), officially the Estado Libre y Soberano de Sinaloa ( en, Free and Sovereign State of Sinaloa), is one of the 31 states which, along with Mexico City, comprise the Administrative divisions of Mexico, Federal Entities of Mexico. It is d ...

Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...

. It is similar in appearance to '' Buellia stellulata'', but has a different secondary chemistry, and ''B. spuria'' is common throughout the Sonoran Desert region, while ''B. stellulata'' is restricted to coastal regions.

References

spuria Lichen species Lichens described in 1828 Lichens of Asia Lichens of Europe Lichens of North America Taxa named by Ludwig Schaerer {{Caliciales-stub

See also

References