Brontops
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''Megacerops'' ("large-horned face", from '' méga-'' "large" + '' kéras'' "horn" + '' ōps'' "face") is an extinct
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of the prehistoric
odd-toed ungulate Odd-toed ungulates, mammals which constitute the taxonomic order Perissodactyla (, ), are animals—ungulates—who have reduced the weight-bearing toes to three (rhinoceroses and tapirs, with tapirs still using four toes on the front legs) o ...
(hoofed mammal)
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Brontotheriidae Brontotheriidae is a family of extinct mammals belonging to the order Perissodactyla, the order that includes horses, rhinoceroses, and tapirs. Superficially, they looked rather like rhinos, although they were actually more closely related to ...
, an extinct group of
rhinoceros A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species ...
-like browsers related to
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...
s. It was endemic to North America during the
Late Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "d ...
epoch (38–33.9 mya), existing for approximately .


Description

All of the species had a pair of blunt
horn Horn most often refers to: *Horn (acoustic), a conical or bell shaped aperture used to guide sound ** Horn (instrument), collective name for tube-shaped wind musical instruments *Horn (anatomy), a pointed, bony projection on the head of various ...
s on their snout (the size varying between species), with the horns of males being much larger than those of the females. This could indicate that they were social animals which butted heads for breeding privileges. Despite resembling the rhinoceros, it was larger than any living rhinoceros: the living animal easily approached the size of the
African forest elephant The African forest elephant (''Loxodonta cyclotis'') is one of the two living African elephant species. It is native to humid forests in West Africa and the Congo Basin. It is the smallest of the three living elephant species, reaching a shoulde ...
, the third-largest land animal today. It stood about tall at the shoulders with an overall length(including tail) of . Its skull reached in greatest length, with some specimens possessing substantial canines, up to 70 mm long. ''Megacerops'' resembled a large rhinoceros, possessing blunt Y-shaped horn-like protrusions on its nose up to 43 cm in length. Its mass is estimated to be in the range of The dorsal
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristi ...
e above the shoulders had extra long spines to support the huge neck muscles needed to carry the heavy
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...
. The shape of its teeth suggests that it preferred food such as soft stems and leaves, rather than tough vegetation. It may have had fleshy lips and a long tongue for carefully selecting food.


Paleobiology

The skeleton of an adult male was found with partially healed rib fractures, which supports the theory that males used their 'horns' to fight each other. No creature living in ''Megacerops'' time and area except another ''Megacerops'' could have inflicted such an injury. The breathing movements prevented the fractures from completely healing. The adults may have also used their horns to defend themselves and their calves from predators, such as
hyaenodonts Hyaenodontidae ("hyena teeth") is a family of extinct predatory mammals from extinct superfamily Hyaenodontoidea within extinct order Hyaenodonta. Hyaenodontids arose during the early Eocene and persisted well into the early Miocene. Fossils of t ...
,
entelodonts Entelodontidae, the entelodonts, are an extinct family of pig-like artiodactyls (even-toed ungulates) which inhabited the Northern Hemisphere (Asia, Europe, and North America) from the late Eocene to the Middle Miocene epochs, about 38-19 mil ...
, '' Bathornis'' or
nimravids Nimravidae is an extinct family of carnivorans, sometimes known as false saber-toothed cats, whose fossils are found in North America and Eurasia. Not considered to belong to the true cats (family Felidae), the nimravids are generally considered ...
.


Discovery

Fossils were uncovered in the northern plains states. Life-sized models of ''Megacerops'' families (a male, female, and juvenile) are displayed at the James E. Martin Paleontological Research Laboratory, South Dakota School of Mines & Technology, and a different set at the Canadian Museum of Nature. Many remains have been found in
South Dakota South Dakota (; Sioux: , ) is a U.S. state in the North Central region of the United States. It is also part of the Great Plains. South Dakota is named after the Lakota and Dakota Sioux Native American tribes, who comprise a large porti ...
and
Nebraska Nebraska () is a state in the Midwestern region of the United States. It is bordered by South Dakota to the north; Iowa to the east and Missouri to the southeast, both across the Missouri River; Kansas to the south; Colorado to the sout ...
. In the past, specimens exposed by severe
rain Rain is water droplets that have condensed from atmospheric water vapor and then fall under gravity. Rain is a major component of the water cycle and is responsible for depositing most of the fresh water on the Earth. It provides water ...
storms were found by Native Americans of the Sioux tribes. The Sioux called them "thunder beasts", a name preserved in the ancient Greek translation (bronto-, ''thunder;'' therion, ''beast''). Many of the skeletons found by the Sioux belonged to herds which were killed by
volcanic eruptions Several types of volcanic eruptions—during which lava, tephra (ash, lapilli, volcanic bombs and volcanic blocks), and assorted gases are expelled from a volcanic vent or fissure—have been distinguished by volcanologists. These are often ...
of the
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico ...
, which were volcanically active at the time. File:Brontotherium hatcheri.jpg, Skeleton of ''Megacerops'' File:Megacerops sp.jpg, Skeleton of ''Megacerops'' in Field Museum of Natural History File:Megacerops models.jpg, Reconstruction of ''Megacerops'' in Canadian Museum of Nature File:Annual report of the Director to the Board of Trustees for the year .. (1931) (17811304663).jpg, Life-size sculpture of Megacerops family


Taxonomy

''Megacerops'' was named by Leidy (1870). Its type species is ''Megacerops coloradensis''. It was synonymized subjectively with ''Menodus'' by Clark and Beerbower (1967). It was assigned to Brontotheriidae by Leidy (1870), Carroll (1988), Mader (1989), and Mader (1998). According to Mihlbachler and others, ''Megacerops'' includes the species of the genera ''Menodus'', ''Brontotherium'', ''Brontops'', ''Menops'', ''Ateleodon'', and ''Oreinotherium''. File:Brontotherium skull IMG 4441.jpg, Skull in Zurich


See also

*


References

{{Taxonbar, from=Q135590 Brontotheres White River Fauna Eocene mammals of North America Eocene odd-toed ungulates Priabonian genus extinctions Fossil taxa described in 1870 Taxa named by Joseph Leidy