''Breviceratops'' (meaning "short horned face") is a genus of

protoceratopsid Protoceratopsidae is a family of basal (primitive) ceratopsians from the Late Cretaceous period. Although ceratopsians have been found all over the world, protoceratopsids are only definitively known from Cretaceous strata in Asia, with most spec ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

that lived during the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...

in what is now the

Barun Goyot Formation The Barun Goyot Formation (also known as Baruungoyot Formation or West Goyot Formation) is a geological formation dating to the Late Cretaceous Period. It is located within and is widely represented in the Gobi Desert Basin, in the Ömnögovi Pro ...

Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...

Discovery and naming

Cretaceous-aged dinosaur fossil localities of Mongolia

The first fossils were discovered during the 1960s by the Polish expedition to the

Nemegt Basin The Nemegt Basin is a geographical area in the northwestern Gobi Desert, in Ömnögovi Province, southern Mongolia. It is known locally as the "Valley of the Dragons", since it is a source of many fossil finds, including dinosaurs, dinosaur eggs, ...

Gobi Desert The Gobi Desert (Chinese: 戈壁 (沙漠), Mongolian: Говь (ᠭᠣᠪᠢ)) () is a large desert or brushland region in East Asia, and is the sixth largest desert in the world. Geography The Gobi measures from southwest to northeast an ...

. The specimens were originally described by

Teresa Maryańska Teresa Maryańska (1937 – 3 October 2019) was a Polish paleontologist who specialized in Mongolian dinosaurs, particularly pachycephalosaurians and ankylosaurians. Peter Dodson (1998 p. 9) states that in 1974 Maryanska together with Halszka ...

and

Halszka Osmólska Halszka Osmólska (September 15, 1930 – March 31, 2008) was a Polish paleontologist who had specialized in Mongolian dinosaurs. Biography She was born in 1930 in Poznań. In 1949, she began to study biology at Faculty of Biology and Earth Scie ...

in 1975 and named as a second species of ''

Protoceratops ''Protoceratops'' (; ) is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 75 to 71 million years ago. The genus ''Protoceratops'' includes two species: ''P. andrewsi'' and the larger ''P. hellenik ...

'', ''Protoceratops kozlowskii'', the specific name honouring Polish

paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...

Roman Kozłowski Roman Stanisław Jakub Kozłowski (1 February 1889 – 2 May 1977) was a Polish palaeontologist, best known for his work on graptolites. Kozłowski was born in Włocławek, north-west of Warsaw, on 1 February 1889. He studied at universities in Sw ...

. The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

, ZPAL MgD-I/117, was found in the Khulsan locality of the

dating from the Late

Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...

. It consists of a partial juvenile postcranial skeleton with skull. Other specimens were referred from the Hermiin Tsav and Khulsan localities: MgD-I/116, a skull and lower jaws of a small juvenile; MgD-I/118, fragmentary postcrania and lower jaws of a juvenile; MgD-I/119, a dentary and three neural arches; MgD-I/20, two dentaries and loose teeth; MgD-I/21, a maxilla fragment with four teeth of a juvenile; and MgD-I/22, teeth. They were then renamed as a separate genus, ''Breviceratops'', by Sergei Mikhailovich Kurzanov in 1990, the generic name combining the

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

''brevis'' (meaning short), with a reference to the

Ceratopsia Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. ...

. Kurzanov also referred an additional number of fossils from Hermiin Tsav, which is the type locality of ''

Bagaceratops ''Bagaceratops'' (meaning "small-horned face") is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 72 to 71 million years ago. ''Bagaceratops'' remains have been reported from the Barun Goyot Forma ...

'', a coeval protoceratopsid. The Hermiin Tsav material has been noted as closely resembling ''Bagaceratops'', which has led to the proposal that ''Breviceratops'' is a juvenile stage and synonym of this protoceratopsid. However, the holotype from Khulsan has primitive premaxillary teeth, a feature inconsistent with the derived morphology of ''Bagaceratops''. Although most of these specimens have been re-identified in fact as ''Bagaceratops'' specimens, ''Breviceratops'' is now restricted to the holotype and juvenile ZPAL MgD-I/116.

Description

''Breviceratops'' had a

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...

combining primitive and derived traits, such as premaxillary

teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...

(also shared with ''Protoceratops andrewsi'') and an

antorbital fenestra An antorbital fenestra (plural: fenestrae) is an opening in the skull that is in front of the eye sockets. This skull character is largely associated with archosauriforms, first appearing during the Triassic Period. Among extant archosaurs, birds ...

(also shared with ''Bagaceratops''), which is an opening of the located in front of the

eye socket In anatomy, the orbit is the cavity or socket of the skull in which the eye and its appendages are situated. "Orbit" can refer to the bony socket, or it can also be used to imply the contents. In the adult human, the volume of the orbit is , of ...

. This

fenestra A fenestra (fenestration; plural fenestrae or fenestrations) is any small opening or pore, commonly used as a term in the biological sciences. It is the Latin word for "window", and is used in various fields to describe a pore in an anatomical st ...

was distinct from ''Bagaceratops'' in that it was narrow to straight-shaped. The skull of ''Breviceratops'' ended in a moderatively pronounced

neck frill A neck frill is the relatively extensive margin seen on the back of the heads of reptiles with either a bony support such as those present on the skulls of dinosaurs of the suborder Marginocephalia or a cartilaginous one as in the frill-necked ...

formed by the and , a common trait among protoceratopsids and overall ceratopsians.

Classification

''Breviceratops'' belonged to the

, a group of herbivorous dinosaurs with

parrot Parrots, also known as psittacines (), are birds of the roughly 398 species in 92 genera comprising the order Psittaciformes (), found mostly in tropical and subtropical regions. The order is subdivided into three superfamilies: the Psittacoid ...

-like beaks which thrived in

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

and

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...

during the Cretaceous period. It has been assigned to the

Protoceratopsidae Protoceratopsidae is a family of basal (primitive) ceratopsians from the Late Cretaceous period. Although ceratopsians have been found all over the world, protoceratopsids are only definitively known from Cretaceous strata in Asia, with most spec ...

in 2019 by Łukasz Czepiński, were ''Bagaceratops'' and ''Protoceratops'' appear to be close relatives. Below are the proposed relationships among Protoceratopsidae by Czepiński:

References

External links

* * {{Taxonbar, from=Q138061 Ceratopsians Ornithischian genera Campanian genera Late Cretaceous dinosaurs of Asia Fossil taxa described in 1990 Taxa named by Sergei Kurzanov Controversial dinosaur taxa

Discovery and naming

Description

Classification

See also

References

External links