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''Bothriechis'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
pit viper The Crotalinae, commonly known as pit vipers,Mehrtens JM (1987). ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . crotaline snakes (from grc, κρόταλον ''krotalon'' castanet), or pit adders, are a subfa ...
s, commonly called palm vipersMehrtens JM. 1987. ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . or palm-pitvipersCampbell JA, Lamar WW. 2004. ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publishing Associates. 870 pp. 1500 plates. . found predominantly in
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
and
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
, although the most common
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
, '' B. schlegelii'', ranges as far south as
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
and
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
. All members are relatively slender and arboreal. The name ''Bothriechis'' is derived from the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
words ''bothros'' and ''echis'' that mean "pit" and "viper" respectively. Ten species and no subspecies are currently generally recognized.


Description

Species that belong to this genus typically reach lengths of 60–80 cm, while '' B. aurifer'', '' B. bicolor'' and '' B. lateralis'' are known to grow to 1 m or more. General characteristics include a sharply defined
canthus rostralis In snakes and amphibians, the canthus, canthal ridge or ''canthus rostralis'',Spawls S, Branch B. 1995. ''The Dangerous Snakes of Africa''. Ralph Curtis Books. Dubai: Oriental Press. 192 pp. . is the angle between the flat crown of the head and the ...
, an unelevated snout, a
rostral scale The rostral scale, or rostral, in snakes and other scaled reptiles is the median plate on the tip of the snout that borders the mouth opening. Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Ithaca and London: ...
that is not as high as it is broad, and a prehensile tail that accounts for at least 15% of the body length. The color pattern usually consists of a green ground color that may or may not include pale or dark markings. '' B. schlegelii'' is an exception to this rule.


Geographic range

Occurs in southern
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
(southeastern
Oaxaca Oaxaca ( , also , , from nci, Huāxyacac ), officially the Free and Sovereign State of Oaxaca ( es, Estado Libre y Soberano de Oaxaca), is one of the 32 states that compose the political divisions of Mexico, Federative Entities of Mexico. It is ...
and the northern highlands of
Chiapas Chiapas (; Tzotzil language, Tzotzil and Tzeltal language, Tzeltal: ''Chyapas'' ), officially the Free and Sovereign State of Chiapas ( es, Estado Libre y Soberano de Chiapas), is one of the states that make up the Political divisions of Mexico, ...
), through Central America to northern
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
(
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
, western
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
,
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
and northern
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
).


Behavior

As a general rule, species found above 1,500 meters altitude tend to be diurnal, while those found below 1,000 meters tend to be active at night. Those found between 1,000 and 1,500 meters may be active at any time of the day.


Venom

''Bothriechis''
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
is primarily a
haemotoxin Hemotoxins, haemotoxins or hematotoxins are toxins that destroy red blood cells, disrupt blood clotting, and/or cause organ degeneration and generalized tissue damage. The term ''hemotoxin'' is to some degree a misnomer since toxins that damage ...
which causes severe pain, swelling, bleb formation,
bruising A bruise, also known as a contusion, is a type of hematoma of tissue, the most common cause being capillaries damaged by trauma, causing localized bleeding that extravasates into the surrounding interstitial tissues. Most bruises occur clos ...
, and quite often
necrosis Necrosis () is a form of cell injury which results in the premature death of cells in living tissue by autolysis. Necrosis is caused by factors external to the cell or tissue, such as infection, or trauma which result in the unregulated dige ...
. If untreated it can lead to loss of a limb, or even death. Each year several farmers and plantation workers are bitten by eyelash vipers, sometimes resulting in fatalities.
Wyeth Wyeth, LLC was an American pharmaceutical company. The company was founded in Philadelphia, Pennsylvania, in 1860 as ''John Wyeth and Brother''. It was later known, in the early 1930s, as American Home Products, before being renamed to Wyeth in ...
in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
an
Instituto Clodomiro Picado
in
Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the no ...
both manufacture different polyvalent
antivenin Antivenom, also known as antivenin, venom antiserum, and antivenom immunoglobulin, is a specific treatment for envenomation. It is composed of antibodies and used to treat certain venomous bites and stings. Antivenoms are recommended only if th ...
s which can be used to treat eyelash viper envenomations.


Species

There are 11 recognized species. T
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...


Taxonomy

A new species from
Guatemala Guatemala ( ; ), officially the Republic of Guatemala ( es, República de Guatemala, links=no), is a country in Central America. It is bordered to the north and west by Mexico; to the northeast by Belize and the Caribbean; to the east by H ...
and
Honduras Honduras, officially the Republic of Honduras, is a country in Central America. The republic of Honduras is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Oce ...
, '' B. thalassinus'', was described by
Campbell Campbell may refer to: People Surname * Campbell (surname), includes a list of people with surname Campbell Given name * Campbell Brown (footballer), an Australian rules footballer * Campbell Brown (journalist) (born 1968), American television ne ...
& E.N. Smith (2000). Campbell & Lamar (2004) recognize this species, as well as a ninth addition to the genus: '' B. supraciliaris'', which was first described by
Taylor Taylor, Taylors or Taylor's may refer to: People * Taylor (surname) **List of people with surname Taylor * Taylor (given name), including Tayla and Taylah * Taylor sept, a branch of Scottish clan Cameron * Justice Taylor (disambiguation) Plac ...
(1954) as a subspecies of ''B. schlegelii'', and is found in southwestern
Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the no ...
. ''B. guifarroi'' was discovered in 2010 and described by
Townsend Townsend (pronounced tounʹ-zənd) or Townshend may refer to: Places United States *Camp Townsend, National Guard training base in Peekskill, New York *Townsend, Delaware *Townsend, Georgia *Townsend, Massachusetts, a New England town ** Townsend ...
et al. in 2013, in a study that included the following suggested
phylogenetic tree A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ...
of the genus:


References


Further reading

* Campbell JA, Smith EN. 2000. "A new species of arboreal pitviper from the Atlantic versant of northern Central America". ''Revista de Biología Tropical'' 48: 1001-1013. * Cope ED. 1860 (dated 1859). "Catalogue of the venomous serpents in the museum of the Academy of Natural Sciences of Philadelphia, with notes on the families, genera and species". ''Proc. Acad. Nat. Sci. Philadelphia'' 11: 332-347 38 * Peters W. 1859. "''Über die von Hrn. Dr. Hoffmann in Costa Rica gesammelten und an das Königl. Zoologische Museum gesandten Schlangen''". ''Monatsberichte der Preussischen Akademie Wissenschaften zu Berlin'' 1859: 275-278 78 * Posada Arango A. 1889. ''Anales de la Academia de Medicina de Medellin'' 2: 45-49 7 *Posada Arango A. 1889. ''Bulletin de la Société Zoologique de France, Paris'' 14: 343-345 43 * Salvin O. 1860. "On the reptiles of Guatemala". ''Proceedings of the Zoological Society of London'' 1860: 451-461 59 * Taylor EH. 1954. "Further studies on the serpents of Costa Rica". ''University of Kansas Science Bulletin'' 36 (11): 673-801.


External links

{{Taxonbar, from=Q944716 Crotalinae Snake genera Taxa named by Wilhelm Peters