Boraginaceae, the
borage or
forget-me-not
''Myosotis'' ( ) is a genus of flowering plants in the family Boraginaceae. The name comes from the Ancient Greek "mouse's ear", which the foliage is thought to resemble. In the northern hemisphere they are colloquially known as forget-me-nots ...
family, includes about 2,000 species of
shrub
A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s,
tree
In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s and
herb
In general use, herbs are a widely distributed and widespread group of plants, excluding vegetables and other plants consumed for macronutrients, with savory or aromatic properties that are used for flavoring and garnishing food, for medicinal ...
s in 146, to 156
genera
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
with a worldwide distribution.
The
APG IV system
The APG IV system of flowering plant classification is the fourth version of a modern, mostly molecular-based, system of plant taxonomy for flowering plants (angiosperms) being developed by the Angiosperm Phylogeny Group (APG). It was publishe ...
from 2016 classifies the Boraginaceae as single family of the order
Boraginales
Boraginales is an order of flowering plants in the asterid clade. It includes the Boraginaceae and a number of other families, with a total of about 125 genera and 2,700 species. Its herbs, shrubs, trees and lianas (vines) have a worldwide distr ...
within the
asterids
In the APG IV system (2016) for the classification of flowering plants, the name asterids denotes a clade (a monophyletic group). Asterids is the largest group of flowering plants, with more than 80,000 species, about a third of the total floweri ...
.
[ ] Under the older
Cronquist system
The Cronquist system is a taxonomic classification system of flowering plants. It was developed by Arthur Cronquist in a series of monographs and texts, including ''The Evolution and Classification of Flowering Plants'' (1968; 2nd edition, 1988) a ...
it was included in Lamiales, but it is now clear that it is no more similar to the other families in this order than they are to families in several other asterid orders. A revision of the Boraginales, also from 2016, split the Boraginaceae in eleven distinct families:
Boraginaceae ''
sensu
''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular co ...
stricto'', Codonaceae,
Coldeniaceae
''Coldenia'', named after C. Colden, is a monotypic genus of flowering plants traditionally included in the borage family, Boraginaceae ''sensu lato''. It was assigned to the subfamily Ehretioideae, but molecular data revealed it to be more clos ...
,
Cordiaceae
Cordioideae is a subfamily of the flowering plant family Boraginaceae.
Genera
*''Coldenia'' L.
*''Cordia
''Cordia'' is a genus of flowering plants in the borage family, Boraginaceae. It contains about 300 species of shrubs and trees, that a ...
,
Ehretiaceae
Ehretioideae is a subfamily of the flowering plant family Boraginaceae.
Genera
*''Bourreria'' P.Browne
*'' Cortesia'' Cav.
*''Ehretia'' P.Browne
*''Halgania'' Gaudich.
*''Ixorhea'' Fenzl
*''Lepidocordia'' Ducke
*'' Menais'' Loefl.
*'' Patag ...
,
Heliotropiaceae Heliotropiaceae is a cosmopolitan family of flowering plants with approximately 450 species worldwide, though it is concentrated especially in the tropics and subtropics.
There are the following four genera in the family:
* ''Heliotropium (''incl. ...
,
Hoplestigmataceae
''Hoplestigma'' is a genus of flowering plants in the family Hoplestigmataceae.Vernon H. Heywood, Richard K. Brummitt, Ole Seberg, and Alastair Culham. 2007. ''Flowering Plant Families of the World''. Firefly Books: Ontario, Canada. . One study o ...
,
Hydrophyllaceae,
Lennoaceae
Lennooideae is a subfamily of parasitic flowering plants of southwestern North America and northwestern South America.
The relationships of this subfamily to other plants remain uncertain. It was traditionally treated at family rank as Lennoacea ...
,
Namaceae, and Wellstediaceae.
These plants have alternately arranged leaves, or a combination of alternate and opposite leaves. The leaf blades usually have a narrow shape; many are linear or lance-shaped. They are smooth-edged or toothed, and some have
petioles. Most species have bisexual flowers, but some taxa are
dioecious
Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...
. Most
pollination
Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds ...
is by
hymenoptera
Hymenoptera is a large order (biology), order of insects, comprising the sawfly, sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are Par ...
ns, such as
bee
Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyly, monophyletic lineage within the ...
s. Most species have
inflorescence
An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s that have a coiling shape, at least when new, called scorpioid cymes. The flower has a usually five-lobed
calyx. The corolla varies in shape from rotate to bell-shaped to tubular, but it generally has five lobes. It can be green, white, yellow, orange, pink, purple, or blue. There are five
stamen
The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10
Morphology and terminology
A stamen typically consists of a stalk called the filame ...
s and one
style
Style is a manner of doing or presenting things and may refer to:
* Architectural style, the features that make a building or structure historically identifiable
* Design, the process of creating something
* Fashion, a prevailing mode of clothing ...
with one or two stigmas. The fruit is a
drupe
In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...
, sometimes fleshy.
Most members of this family have hairy leaves. The coarse character of the hairs is due to
cystolith
Cystolith (Gr. "cavity" and "stone") is a botanical term for outgrowths of the epidermal cell wall, usually of calcium carbonate, formed in a cellulose matrix in special cells called lithocysts, generally in the leaf of plants.
Cystoliths are pr ...
s of
silicon dioxide
Silicon dioxide, also known as silica, is an oxide of silicon with the chemical formula , most commonly found in nature as quartz and in various living organisms. In many parts of the world, silica is the major constituent of sand. Silica is one ...
and
calcium carbonate. These hairs can induce an adverse skin reaction, including itching and rash in some individuals, particularly among people who handle the plants regularly, such as gardeners. In some species,
anthocyanin
Anthocyanins (), also called anthocyans, are water-soluble vacuolar pigments that, depending on their pH, may appear red, purple, blue, or black. In 1835, the German pharmacist Ludwig Clamor Marquart gave the name Anthokyan to a chemical compo ...
s cause the flowers to change color from red to blue with age. This may be a signal to
pollinator
A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains.
Insects are the maj ...
s that a flower is old and depleted of
pollen
Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
and
nectar
Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists ...
.
[Hess, D. 2005. ''Systematische Botanik''. ]
Well-known members of the family include:
*
alkanet
Alkanet is the common name of several related plants in the borage family (Boraginaceae):
* Alkanet, ''Alkanna tinctoria'', the source of a red dye; this is the plant most commonly called simply "alkanet"
* Various other plants of the genus '' Alk ...
(''Alkanna tinctoria'')
*
borage
Borage ( or ; ''Borago officinalis''), also known as starflower, is an annual herb in the flowering plant family Boraginaceae. It is native to the Mediterranean region, and has naturalized in many other locales.
It grows satisfactorily in gard ...
(''Borago officinalis'')
*
comfrey
''Symphytum'' is a genus of flowering plants in the borage family, Boraginaceae, known by the common name comfrey (pronounced ). There are 59 recognized species.WFO (2022): Symphytum L. Published on the Internet; http://www.worldfloraonline.org/ ...
(''Symphytum'' spp.)
*
fiddleneck
''Amsinckia'' is a genus of flowering plants commonly known as fiddlenecks. The common name is derived from the flower stems, which curl over at the top in a manner reminiscent of the head of a fiddle. Fiddlenecks are in the family Boraginac ...
(''Amsinckia'' spp.)
*
forget-me-not
''Myosotis'' ( ) is a genus of flowering plants in the family Boraginaceae. The name comes from the Ancient Greek "mouse's ear", which the foliage is thought to resemble. In the northern hemisphere they are colloquially known as forget-me-nots ...
(''Myosotis'' spp.)
*
geigertree (''Cordia sebestena'')
*
green alkanet
''Pentaglottis'' is a monotypic genus of flowering plants in the family Boraginaceae. It is represented by a single species, ''Pentaglottis sempervirens'', commonly known as the green alkanet, evergreen bugloss or alkanet, and is a bristly, pe ...
(''Pentaglottis sempervirens'')
*
heliotrope (''Heliotropium'' spp.)
*
hound's tongue
''Cynoglossum'' is a genus of small-flowered plants in the family Boraginaceae (borage family).
''Cynoglossum officinale'', the common hound's-tongue, is a native of Asia, Africa, and Europe. It has been introduced into North America, and it is ...
(''Cynoglossum'' spp.)
*
lungwort
''Pulmonaria'' (lungwort) is a genus of flowering plants in the family Boraginaceae, native to Europe and western Asia, with one species (''P. mollissima'') east to central Asia. According to various estimates there may be between 10 and 18 spe ...
(''Pulmonaria'' spp.)
*
oysterplant (''Mertensia maritima'')
*
purple viper's bugloss/Salvation Jane (''Echium plantagineum'')
*
Siberian bugloss (''Brunnera macrophylla'')
*
viper's bugloss
''Echium vulgare'', known as viper's bugloss and blueweed,Dickinson, T.; Metsger, D.; Bull, J.; & Dickinson, R. (2004) ROM Field Guide to Wildflowers of Ontario. Toronto:Royal Ontario Museum, p. 203. is a species of flowering plant in the borag ...
(''Echium vulgare'')
Genera
According to
Kew
Kew () is a district in the London Borough of Richmond upon Thames. Its population at the 2011 census was 11,436. Kew is the location of the Royal Botanic Gardens ("Kew Gardens"), now a World Heritage Site, which includes Kew Palace. Kew is a ...
;
[
* '']Actinocarya
''Actinocarya'' is a small genus of annual herbs in the family Boraginaceae. Species in the genus are found in Pakistan, China, and India
India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-la ...
''
* '' Adelinia''
* ''Adelocaryum
''Adelocaryum'' is a genus of flowering plants belonging to the family Boraginaceae.
Its native range is southern Arabian Peninsula and the Indian subcontinent
The Indian subcontinent is a list of the physiographic regions of the world, p ...
''
* '' Aegonychon''
* ''Afrotysonia
''Afrotysonia'' is a genus of flowering plants belonging to the family Boraginaceae
Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide distribution. ...
''
* '' Ailuroglossum''
* ''Alkanna
''Alkanna'' is a genus of herbaceous plants including about 60 species of the family Boraginaceae.
Selected species
* '' A. amana'' Rech.f.
* '' A. angustifolia'' Sümbül
* '' A. areolata'' Boiss.
* '' A. attilae'' P.H.D ...
''
* '' Amblynotus''
* ''Amphibologyne
''Amphibologyne'' is a genus of flowering plants belonging to the family Boraginaceae
Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide distri ...
''
* ''Amsinckia
''Amsinckia'' is a genus of flowering plants commonly known as fiddlenecks. The common name is derived from the flower stems, which curl over at the top in a manner reminiscent of the head of a fiddle. Fiddlenecks are in the family Boraginac ...
''
* '' Amsinckiopsis''
* ''Anchusa
The genus ''Anchusa'' belongs to the borage family (Boraginaceae). It includes about 35 species found growing in Europe, North Africa, South Africa and Western Asia. They are introduced in the United States.
They consist of annual plants, bie ...
''
* '' Ancistrocarya''
* '' Andersonglossum''
* '' Anoplocaryum''
* ''Antiotrema
''Antiotrema'' is a genus of flowering plants belonging to the family Boraginaceae
Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide distributi ...
''
* '' Antiphytum''
* ''Arnebia
''Arnebia'' is a genus of flowering plants in the family Boraginaceae. There are about 38 species, most are located in the Mediterranean region and eastwards to the Himalayas and one species extending down to tropical Africa.
The generic name " ...
''
* '' Asperugo''
* '' Auxemma''
* ''Borago
''Borago'', or borage, is a genus of five species of herbs native to the Mediterranean, with one species, ''Borago officinalis'', cultivated and naturalized throughout the world.
Uses
''Borago officinalis'' is used medicinally, in companion pl ...
''
* '' Bothriospermum''
* ''Bourreria
''Bourreria'' is a genus of flowering plants in the borage family, Boraginaceae. Members of the genus are commonly known as strongbark or strongback. The generic name was chosen by Patrick Browne to honour German pharmacist Johann Ambrosius Beur ...
''
* '' Brachybotrys''
* '' Brandella''
* '' Brunnera''
* ''Buglossoides
''Buglossoides'' is a genus consisting of 15 species of annual or perennial herbs, native to Europe and Asia. They grow naturally in habitats ranging from sunny scrub to rocky slopes and woodland areas. These plants are covered in fine bristles ...
''
* '' Caccinia''
* ''Cerinthe
''Cerinthe'' is a genus of flowering plants in the family Boraginaceae, known as honeyworts. The genus is characterised by a calyx made up of separate, rather than fused, sepals, a tubular corolla, and the schizocarpic fruit that divides into ...
''
* '' Chionocharis''
* ''Choriantha
''Choriantha'' is a genus of flowering plants belonging to the family Boraginaceae
Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide distributi ...
''
* ''Codon
The genetic code is the set of rules used by living cells to translate information encoded within genetic material ( DNA or RNA sequences of nucleotide triplets, or codons) into proteins. Translation is accomplished by the ribosome, which links ...
''
* ''Coldenia
''Coldenia'', named after C. Colden, is a monotypic genus of flowering plants traditionally included in the borage family, Boraginaceae ''sensu lato''. It was assigned to the subfamily Ehretioideae, but molecular data revealed it to be more clos ...
''
* ''Cordia
''Cordia'' is a genus of flowering plants in the borage family, Boraginaceae. It contains about 300 species of shrubs and trees, that are found worldwide, mostly in warmer regions. Many of the species are commonly called manjack, while ''bocote' ...
''
* '' Craniospermum''
* '' Crucicaryum''
* ''Cryptantha
''Cryptantha'' is a genus of flowering plants in the borage family, Boraginaceae. They are known commonly as cat's eyes and popcorn flowers (the latter name is also used to refer to the closely related genus ''Plagiobothrys'',Hasenstab-Lehman, ...
''
* '' Cynoglossopsis''
* ''Cynoglossum
''Cynoglossum'' is a genus of small-flowered plants in the family Boraginaceae (borage family).
''Cynoglossum officinale'', the common hound's-tongue, is a native of Asia, Africa, and Europe. It has been introduced into North America, and it is ...
''
* '' Cynoglottis''
* '' Cystostemon''
* '' Dasynotus''
* '' Decalepidanthus''
* '' Draperia''
* '' Echiochilon''
* ''Echium
''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae.
Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands where ...
''
* ''Ehretia
''Ehretia'' is a genus of flowering plants in the borage family, Boraginaceae. It contains about 50 species.
The generic name honors German botanical illustrator Georg Dionysius Ehret (1708–1770).
Species
Accepted species and other notable t ...
''
* '' Ellisia''
* ''Embadium
''Embadium'' is a genus of flowering plants belonging to the family Boraginaceae.
Its native range is Central and Southern Australia.
Species:
*''Embadium jobnstonii''
*''Embadium stagnense''
*''Embadium uncinatum''
References
{{Taxo ...
''
* '' Emmenanthe''
* ''Eremocarya
''Eremocarya'' is a genus of flowering plants in the family Boraginaceae. There are about 63 species and its native range extends through western United States to northwestern Mexico. It is part of subtribe of ''Amsinckiinae''.
It was once thoug ...
''
* ''Eriodictyon
''Eriodictyon'' is a genus of plants known by the common name yerba santa within the Hydrophylloideae subfamily of the borage family, Boraginaceae. They are distributed throughout the southwestern United States and Mexico.
Description
Most spe ...
''
* ''Eritrichium
''Eritrichium'' (alpine forget-me-not) is a genus of flowering plants in the family Boraginaceae. It contains 78 species. Notable members include '' Eritrichium howardii'' and ''Eritrichium nanum''.
Its native range stretches from temperate Eura ...
''
* '' Eucrypta''
* ''Euploca
''Euploca'' is an almost cosmopolitan genus of plants with around 100 species. It was first described by Thomas Nuttall in 1837. While part of the broadly defined Boraginaceae in the APG IV system from 2016, a revision of the order Boraginales fr ...
''
* '' Gastrocotyle''
* '' Glandora''
* ''Greeneocharis
''Greeneocharis'' is a genus of flowering plants in the family Boraginaceae. There are about 63 species and its native range extends from western U.S.A. down to northwestern Mexico and to western Argentina. It is part of subtribe of ''Amsinckiin ...
''
* '' Gyrocaryum''
* ''Hackelia
''Hackelia'' (stickseeds) is a genus of plants in the borage family, Boraginaceae. It includes 54 species found in North America, western South America, temperate Eurasia, and Australia. 12 species are native to California
California is a ...
''
* '' Halacsya''
* ''Halgania
''Halgania'' is a genus of small shrubs in the family Boraginaceae. The genus comprises about 20 species that are endemic to Australia. ''Halgania'' is named for Emmanuel Halgan, a vice-admiral in the French Navy.
Features
''Halgania'' species ...
''
* '' Harpagonella''
* '' Heliocarya''
* ''Heliotropium
''Heliotropium'' is a genus of flowering plants in the heliotrope family, Heliotropiaceae. There are around 325 species in this almost cosmopolitan genus, which are commonly known as heliotropes (sg. ). It is highly toxic for dogs and cats.
Et ...
''
* ''Hesperochiron
''Hesperochiron'' is a small genus of plants in the waterleaf family containing two species native to western North America. These are thick-rooted perennial herbs growing in squat patches at ground level and producing bluish-white flowers with ...
''
* '' Heterocaryum''
* '' Huynhia''
* '' Hydrophyllum''
* '' Iberodes''
* '' Ivanjohnstonia''
* ''Ixorhea
''Ixorhea'' is a genus of flowering plants belonging to the family Boraginaceae
Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide distribution.
...
''
* ''Johnstonella
''Johnstonella'' is a genus of flowering plants belonging to the family Boraginaceae.
Its native range is south western and southern USA to Mexico, Peru to southern South America. It is found in the American states of Arizona, California, Nevada ...
''
* ''Lappula
''Lappula'' is a genus of flowering plants in the Boraginaceae, borage family known generally as stickseeds. They are Indigenous (ecology), native to the northern hemisphere. These are Annual plant, annual Herbaceous plant, herbs producing funnel ...
''
* ''Lasiocaryum
''Lasiocaryum'' is a genus of flowering plants belonging to the family Boraginaceae.
Its native range is Himalaya
The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the T ...
''
* ''Lennoa
''Lennoa'' is a monotypic genus of flowering plants belonging to the family Boraginaceae. It only contains one known species, Lennoa madreporoides Lex. It is within the subfamily of Lennoaceae.
Its native range is Mexico to Venezuela. It is foun ...
''
* ''Lepechiniella
''Lepechiniella'' is a genus of flowering plants belonging to the family Boraginaceae.
Its native range is Iran and Western Himalaya, to Xinjiang (in China). It is also found in the countries of Afghanistan, Kazakhstan, Kyrgyzstan, Pakistan, Taji ...
''
* ''Lepidocordia
''Lepidocordia'' is a genus of flowering plants belonging to the family Boraginaceae
Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide distribut ...
''
* '' Lindelofia''
* ''Lithodora
''Lithodora'' is a genus of flowering plants in the family Boraginaceae, native to southwestern Europe, southern Greece, Turkey and Algeria. They are low-growing, evergreen shrubs and subshrubs, producing 5-lobed blue or white flowers. The Greek ...
''
* ''Lithospermum
''Lithospermum'' is a genus of plants belonging to the family Boraginaceae. The genus is distributed nearly worldwide, but most are native to the Americas and the center of diversity is in the southwestern United States and Mexico. Species are ...
''
* '' Lobostemon''
* '' Maharanga''
* '' Mairetis''
* '' Mattiastrum''
* '' Megacaryon''
* '' Melanortocarya''
* '' Memoremea''
* ''Mertensia
''Mertensia'' is a genus of flowering plants in the family Boraginaceae. They are perennial herbaceous plants with blue or sometimes white flowers that open from pink-tinged buds. Such a change in flower color is common in Boraginaceae and is ca ...
''
* '' Microcaryum''
* '' Microparacaryum''
* '' Microula''
* '' Mimophytum''
* '' Moltkia''
* ''Moltkiopsis
''Moltkiopsis'' is a monotypic genus of flowering plants belonging to the family Boraginaceae. The only species is ''Moltkiopsis ciliata''.
Its native range is northern Africa to Sahel and the Arabian Peninsula. It is found in the regions of Alg ...
''
* ''Moritzia
''Moritzia'' is a genus of flowering plants belonging to the family Boraginaceae.
Its native range is Costa Rica to Southern Tropical America. It is found in Brazil, Colombia, Costa Rica, Ecuador, Panamá, Peru and Venezuela.
The genus name of ...
''
* ''Myosotidium
''Myosotidium'' is a genus of plants belonging to the family Boraginaceae. This genus is represented by the single species ''Myosotidium hortensia'', the Chatham Islands lily, giant forget-me-not or Chatham Islands forget-me-not, which is endemi ...
''
* ''Myosotis
''Myosotis'' ( ) is a genus of flowering plants in the family Boraginaceae. The name comes from the Ancient Greek "mouse's ear", which the foliage is thought to resemble. In the northern hemisphere they are colloquially known as forget-me-nots ...
''
* ''Myriopus
''Myriopus'' is a genus of flowering plants in the borage and forget-me-not family Boraginaceae (or Heliotropiaceae Heliotropiaceae is a cosmopolitan family of flowering plants with approximately 450 species worldwide, though it is concentrated ...
''
* '' Nama''
* '' Neatostema''
* ''Nemophila
''Nemophila'' is a genus found in the flowering plant family Boraginaceae.
Most of the species in ''Nemophila'' contain the phrase "baby blue eyes" in their common names. ''N. menziesii'' has the common name of "baby blue eyes". ''N. parviflora' ...
''
* '' Nesocaryum''
* ''Nihon
Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
''
* '' Nogalia''
* ''Nonea
''Nonea'' is a genus of flowering plants in the family Boraginaceae. Sometimes known as monkswort, these are herbaceous perennial plant, perennials or Annual plant, annual plants, native to Europe, Asia and Africa.
The leaves are grayish-green, ...
''
* '' Ogastemma''
* ''Omphalodes
''Omphalodes'' (navelwort) is a genus of flowering plants in the family Boraginaceae, widely distributed in the temperate Northern Hemisphere. In spring they produce blue or white flowers similar to forget-me-nots.
Both the Greek ''Omphalodes'' ...
''
* '' Omphalolappula''
* ''Omphalotrigonotis
''Omphalotrigonotis'' is a genus of flowering plants belonging to the family Boraginaceae
Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide dist ...
''
* '' Oncaglossum''
* ''Onosma
''Onosma'' is a genus of flowering plants in the family Boraginaceae. They are native to the Mediterranean and western Asia. They grow in dry, sunny habitats with rocky, sandy substrates.Koyuncu, O., et al. (2013)A new ''Onosma'' (Boraginaceae) s ...
''
* ''Oreocarya
''Oreocarya'' (Perennial Cat's-Eye) is a genus of flowering plants in the family Boraginaceae. There are about 63 species and its native range extends from western and central Canada, through western United States to north Mexico. It is part of s ...
''
* '' Paracaryum''
* '' Paramoltkia''
* '' Pardoglossum''
* ''Pectocarya
''Pectocarya'' is a plant genus of about 15 species in the family Boraginaceae. Plants in this genus are known generally as combseeds. They are small annual plants which bear tiny white flowers no more than 3 millimeters in diameter. Their fruits ...
''
* ''Pentaglottis
''Pentaglottis'' is a monotypic genus of flowering plants in the family Boraginaceae. It is represented by a single species, ''Pentaglottis sempervirens'', commonly known as the green alkanet, evergreen bugloss or alkanet, and is a bristly, pe ...
''
* ''Phacelia
''Phacelia'' (phacelia, scorpionweed, heliotrope) is a genus of about 200 species of annual or perennial herbaceous plants in the borage family, native to North and South America. California is particularly rich in species with over 90 recorded ...
''
* ''Pholisma
The genus ''Pholisma'' (Nutt. ex Hook.) consists of three to five species of desert-dwelling, primarily subterranean plants.
Most notable of the genus is '' Pholisma sonorae'', native to the Southwestern United States and Mexico. The species is ...
''
* ''Pholistoma
''Pholistoma'' is a small genus of flowering plants in the borage family known generally as fiestaflowers. There are three species, all native to a section of western North America between Oregon and Baja California
Baja California (; 'Lower ...
''
* ''Plagiobothrys
''Plagiobothrys'' is a genus of flowering plants known commonly as popcorn flowers. These are small herbaceous plants which bear tiny white or yellow flowers. Their fruits are nutlets. Although these plants are found predominantly in North Ameri ...
''
* '' Podonosma''
* '' Pontechium''
* '' Pseudoheterocaryum''
* '' Pseudolappula''
* ''Pulmonaria
''Pulmonaria'' (lungwort) is a genus of flowering plants in the family Boraginaceae, native to Europe and western Asia, with one species (''P. mollissima'') east to central Asia. According to various estimates there may be between 10 and 18 spe ...
''
* '' Rindera''
* ''Rochefortia
''Rochefortia'' is a genus of flowering plants in family Boraginaceae.
Its species are native to North America, the Caribbean, and South America.
Species
Species include:
* '' Rochefortia acanthophora'' (syn: ''Ehretia
''Ehretia'' is a gen ...
''
* '' Rochelia''
* '' Romanzoffia''
* ''Rotula
A ''rotulus'' (plural ''rotuli'') or ''rotula'' (pl. ''rotulae'') is often referred to as a "vertical roll," is a long and narrow strip of writing material, historically papyrus or parchment, that is wound around a wooden axle or rod. Rotuli are ...
''
* ''Sauria
Sauria is the clade containing the most recent common ancestor of archosaurs (such as crocodilians, dinosaurs, etc.) and lepidosaurs ( lizards and kin), and all its descendants. Since most molecular phylogenies recover turtles as more closely re ...
''
* ''Selkirkia
''Selkirkia'' is a genus of predatory, tubicolous priapulid worms known from the Middle Cambrian Burgess Shale, Ogygopsis Shale and Puncoviscana Formation. 142 specimens of ''Selkirkia'' are known from the Greater Phyllopod bed, where they compr ...
''
* '' Simpsonanthus''
* '' Sinojohnstonia''
* '' Solenanthus''
* '' Stenosolenium''
* '' Stephanocaryum''
* '' Suchtelenia''
* ''Symphytum
''Symphytum'' is a genus of flowering plants in the borage family, Boraginaceae, known by the common name comfrey (pronounced ). There are 59 recognized species.WFO (2022): Symphytum L. Published on the Internet; http://www.worldfloraonline.org/ ...
''
* '' Thaumatocaryum''
* '' Thyrocarpus''
* '' Tianschaniella''
* ''Tiquilia
''Tiquilia'' is a genus of flowering plants in the borage family, Boraginaceae. The 27 species in this genus are known by the common name crinklemat. They are native to the Western Hemisphere and are mostly found in desert regions.
Selected spe ...
''
* ''Tournefortia
''Tournefortia'', commonly known as soldierbush, is a genus of flowering plants in the borage family, Boraginaceae.
It was first published under the name ''Pittonia'' by Charles Plumier in 1703, in honour of Joseph Pitton de Tournefort. Later, ...
''
* '' Trachelanthus''
* ''Trachystemon
''Trachystemon'' is a genus of flowering plants belonging to the family Boraginaceae.
Its native range is Bulgaria to Turkey. It is a monotypic genus, with the only member being ''Trachystemon orientalis''(L.) D.Don.
Trachystemon orientalis
' ...
''
* ''Trichodesma
''Trichodesma'' is a genus of flowering plants in the borage family, Boraginaceae. There are about 40 to 45 species distributed in tropical
The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tr ...
''
* '' Trigonocaryum''
* '' Trigonotis''
* '' Turricula''
* '' Varronia''
* '' Valentiniella''
* ''Wellstedia
''Wellstedia'' is a genus of flowering plants traditionally included in the family Boraginaceae ''s.l.'', but placed in its own family, Wellstediaceae within the Boraginales order, by the Boraginales Working Group.
Species include:
* '' Wellste ...
''
* '' Wigandia''
References
External links
Distribution Map
Missouri Botanical Garden
BoraginaceaeMabberley's Plant-Book
Boraginaceae
Plant Names
IPNI
Home page of James L. Reveal and C. Rose Broome
Boraginaceae (Search Exact)
Name Search
Tropicos
Missouri Botanical Garden
Topwalks.net: Walking Routes in Spain.
Boraginaceae.
Integrated Taxonomic Information System (ITIS).
Further reading
* Diane, N., et al. 2002
A systematic analysis of ''Heliotropium'', ''Tournefortia'', and allied taxa of the Heliotropiaceae (Boraginales) based on ITS1 sequences and morphological data.
''American Journal of Botany'' 89(2), 287-95.
* Gottschling, M., et al. (2001). Secondary structure of the ITS1 transcript and its application in a reconstruction of the phylogeny of Boraginales. ''Plant Biology'' 3, 629-36.
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Asterid families