''Leucocarbo'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

of birds in the family

Phalacrocoracidae Phalacrocoracidae is a family of approximately 40 species of aquatic birds commonly known as cormorants and shags. Several different classifications of the family have been proposed, but in 2021 the IOC adopted a consensus taxonomy of seven gen ...

with the members commonly known as blue-eyed shags. This is a group of closely related cormorant taxa. Many have a blue, purple or red ring around the eye (not a blue iris); other shared features are white underparts (at least in some individuals) and pink feet. They are found around the colder parts of the Southern Hemisphere, especially near southern

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...

Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...

, and

New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...

. Many are

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

to remote islands. Determining which types are species and which are subspecies of what larger species is problematic; various recent authorities have recognized from 8 to 14 species and have placed them in a variety of genera. The common names are even more confusing, "like myriad footprints criss-crossing in the snow and about as easy to disentangle." Only one common name is given for most species here.

Taxonomy

The genus ''Leucocarbo'' was introduced in 1856 by the French naturalist

Charles Lucien Bonaparte Charles Lucien Jules Laurent Bonaparte, 2nd Prince of Canino and Musignano (24 May 1803 – 29 July 1857), was a French naturalist and ornithologist. Lucien and his wife had twelve children, including Cardinal Lucien Bonaparte. Life and career ...

. He did not specify a

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

but this was designated as the

Guanay cormorant The Guanay cormorant or Guanay shag (''Leucocarbo bougainvilliorum'') is a member of the cormorant family found on the Pacific coast of Peru and northern Chile. (The Argentinian population on the Patagonian Atlantic coast appears to be extirpat ...

by William Ogilvie-Grant in 1898. The name ''Leucocarbo'' combines the

Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...

''leukos'' meaning "white" with the genus name ''Carbo'' introduced by

Bernard Germain de Lacépède Bernard-Germain-Étienne de La Ville-sur-Illon, comte de Lacépède or La Cépède (; 26 December 17566 October 1825) was a French naturalist and an active freemason. He is known for his contribution to the Comte de Buffon's great work, the ...

in 1799. A molecular phylogenetic study published in 2014 found that ''Leucocarbo'' is

sister A sister is a woman or a girl who shares one or more parents with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a family, familial relationship, it is sometimes used endearingly to r ...

to the American cormorants in the genus ''

Nannopterum ''Nannopterum'' is a genus of cormorant comprising three species. They are found throughout the Americas, hence the common name American cormorants. These species were formerly classified in the genus '' Phalacrocorax''. A molecular phylogenet ...

''; the genera split between 6.7 - 8.0 million years ago. The genus contains 16 species: *

Rock shag The rock shag (''Leucocarbo magellanicus''), also known as the Magellanic cormorant, is a marine cormorant found around the southernmost coasts of South America. Its breeding range is from around Valdivia, Chile, south to Cape Horn and Tierra de ...

, ''Leucocarbo magellanicus'' *

, ''Leucocarbo bougainvillii'' *

Bounty shag The Bounty shag (''Leucocarbo ranfurlyi''), also known as the Bounty Island shag, is a species of cormorant of the family Phalacrocoracidae. They are found only on the tiny and remote Subantarctic Bounty Islands, 670 km southeast of New Ze ...

, ''Leucocarbo ranfurlyi'' *

New Zealand king shag The New Zealand king shag (''Leucocarbo carunculatus''), also known as the rough-faced shag, king shag or kawau, is a rare bird endemic to New Zealand. Some taxonomic authorities, including the International Ornithologists' Union, place this spe ...

or rough-faced shag, ''Leucocarbo carunculatus '' *

Chatham shag The Chatham shag (''Leucocarbo onslowi''), also known as the Chatham Island shag, is a species of bird in the cormorant and shag family, Phalacrocoracidae. It is endemic to the Chatham Islands of New Zealand. For a long time the species was plac ...

, ''Leucocarbo onslowi'' *

Otago shag The Otago shag, (''Leucocarbo chalconotus''), together with the Foveaux shag formerly known as the Stewart Island shag and in its dark phase as the bronze shag, is a species of shag now found only in coastal Otago, New Zealand. Description ...

, ''Leucocarbo chalconotus'' *

Foveaux shag The Foveaux shag (''Leucocarbo stewarti''), together with the Otago shag formerly known as the Stewart Island shag and in its dark phase as the bronze shag, is a species of shag endemic to Stewart Island/Rakiura and Foveaux Strait, from which i ...

, ''Leucocarbo stewarti'' *

Auckland shag The Auckland shag (''Leucocarbo colensoi'') or Auckland Islands shag is a species of cormorant from New Zealand. The species is endemic to the Auckland Islands archipelago. It is a sedentary bird that primarily eats various crustaceans and fish. ...

, ''Leucocarbo colensoi'' *

Campbell shag The Campbell shag (''Leucocarbo campbelli''), also known as the Campbell Island shag, is a species of bird in the family Phalacrocoracidae. It is endemic to Campbell Island. Its natural habitats are open seas and rocky shores. It is a medium- ...

, ''Leucocarbo campbelli'' *

Imperial shag The imperial shag or imperial cormorant (''Leucocarbo atriceps'') is a black and white cormorant native to southern South America, primarily in rocky coastal regions, but locally also at large inland lakes. Some taxonomic authorities, including ...

or blue-eyed shag, ''Leucocarbo atriceps'' *

South Georgia shag The South Georgia shag (''Leucocarbo georgianus''), also known as the South Georgia cormorant, is a marine cormorant native to South Georgia and a few other subantarctic islands in the South Atlantic Ocean. Taxonomy The South Georgia shag is one ...

, ''Leucocarbo georgianus'' *

Crozet shag The Crozet shag (''Leucocarbo melanogenis''), also known as the South Georgia cormorant, is a marine cormorant native to the Crozet, Prince Edward and Marion islands in the South Atlantic Ocean. Characteristics The Crozet shag is a member o ...

, ''Leucocarbo melanogenis'' *

Antarctic shag The Antarctic shag (''Leucocarbo bransfieldensis''), sometimes referred to as the imperial cormorant, king cormorant, imperial shag, blue-eyed shag or Antarctic cormorant, is the only species of the cormorant family found in the Antarctic. It is s ...

, ''Leucocarbo bransfieldensis'' *

Kerguelen shag The Kerguelen shag (''Leucocarbo verrucosus'') is a species of cormorant endemic to the Kerguelen Islands in the southern Indian Ocean, one of the most isolated places on Earth. Many authorities consider it a subspecies of the imperial shag. R ...

, ''Leucocarbo verrucosus'' *

Heard Island shag The Heard Island shag (''Leucocarbo nivalis''), or Heard Island cormorant, is a marine cormorant native to the Australian territory comprising the Heard and McDonald Islands in the Southern Ocean, about 4100 km south-west of Perth, Wester ...

, ''Leucocarbo nivalis'' *

Macquarie shag The Macquarie shag (''Leucocarbo purpurascens''), Macquarie Island shag or Macquarie Island cormorant, is a marine cormorant native to Macquarie Island in the Southern Ocean, about halfway between Australia and Antarctica. Taxonomy The Macquarie ...

, ''Leucocarbo purpurascens''

References

{{DEFAULTSORT:Blue-Eyed Shag Leucocarbo Taxa named by Charles Lucien Bonaparte