''Blechnum'', known as hard fern, is a
genus of
ferns in the family
Blechnaceae, subfamily Blechnoideae, according to the
Pteridophyte Phylogeny Group
The Pteridophyte Phylogeny Group, or PPG, is an informal international group of systematic botanists who collaborate to establish a consensus on the classification of pteridophytes (lycophytes and ferns) that reflects knowledge about plant relation ...
classification of 2016 (PPG I).
[ Two very different circumscriptions of the genus are used by different authors. In the PPG I system, based on Gasper et al. (2016),][ ''Blechnum'' is one of 18 genera in the subfamily Blechnoideae, and has about 30 species.][ Other sources use a very broadly defined ''Blechnum'' s.l., including accepting only two other genera in the subfamily. The genus then has about 250 species.][ In the PPG I circumscription, the genus is mostly neotropical, with a few southern African species.
]
Description
Plants in the genus ''Blechnum'' (as circumscribed in the PPG I classification) are mainly terrestrial or grow on rocks; few are epiphyte
An epiphyte is an organism that grows on the surface of a plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphytes grow are called phoroph ...
s. Their rhizome
In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
s may be upright or creeping and have scales with entire margins or at most a few very small teeth. They generally form stolon
In biology, stolons (from Latin '' stolō'', genitive ''stolōnis'' – "branch"), also known as runners, are horizontal connections between organisms. They may be part of the organism, or of its skeleton; typically, animal stolons are external s ...
s, which is a characteristic of the genus. The sterile and fertile fronds are usually of the same form or at most slightly different. The blades of the fronds are of one colour and are usually pinnatisect
The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...
or unipinnate, rarely entire. The leaf veins are usually free, dividing one to three times, each ending near the frond margin in an enlarged tip. The sori have indusia (scale-like coverings).[
]
Species
Using the PPG I circumscription, , the ''Checklist of Ferns and Lycophytes of the World'' accepted the following species and hybrids.[
*'']Blechnum anthracinum
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used b ...
'' R.C.Moran
*'' Blechnum appendiculatum'' Willd.
*''Blechnum arcuatum
''Blechnum arcuatum'', known locally as ''quil-quil'', is a fern native with a natural range in Chile ranging from Ñuble River
The Ñuble River or Rio Ñuble is a river in Ñuble Region, located in the southern portion of central Chile. Its m ...
'' J.Rémy ex Fée
*''Blechnum areolatum
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' V.A.O.Dittrich & Salino
*'' Blechnum asplenioides'' Sw.
*''Blechnum auriculatum
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different Circumscription (taxonomy), circumscriptions ...
'' Cav.
*'' Blechnum australe'' L.
*''Blechnum austrobrasilianum
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' de la Sota
* ''Blechnum'' × ''caudatum'' Cav.
* ''Blechnum'' × ''confluens'' Schltdl. & Cham.
* ''Blechnum'' × ''falciculatum'' C.Presl
*''Blechnum gracile
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' Kaulf.
*''Blechnum gracilipes
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' (Rosenst.) M.Kessler & A.R.Sm.
*''Blechnum guayanense
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' A.Rojas
*''Blechnum hastatum
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' Kaulf.
*''Blechnum heringeri
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' Brade
*''Blechnum laevigatum
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' Cav.
*''Blechnum lanceola
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' Sw.
* ''Blechnum'' × ''leopoldense'' (Dutra) V.A.O.Dittrich & Salino
*''Blechnum longipilosum
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' V.A.O.Dittrich & Salino
*'' Blechnum ludificans'' Herter
*''Blechnum malacothrix
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used b ...
'' Maxon & Morton
*''Blechnum meridense
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used b ...
'' Klotzsch
*''Blechnum occidentale
''Blechnum occidentale'' is a fern in the family Blechnaceae. Its native range is from Mexico through Central America and the Caribbean to tropical South America.
''Blechnum scaberulum'' and ''Blechnum sodiroi'' have been regarded as separate s ...
'' L.
* ''Blechnum'' × ''pampasicum'' de la Sota & M.L.Durán
*''Blechnum polypodioides
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used b ...
'' Raddi
*''Blechnum punctulatum
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' Sw.
*''Blechnum tomentosum
''Blechnum'', known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Two very different circumscriptions of the genus are used by d ...
'' M.Kessler & A.R.Sm.
References
Further reading
* Huxley, A., ed. (1992). ''New RHS Dictionary of Gardening''. Macmillan
* Moore, D. M. (1983). ''Flora of Tierra del Fuego''
External links
Trees and shrubs in the Andes of Ecuador: ''Blechnum''
{{Taxonbar, from=Q2049575
Blechnaceae
Fern genera
Taxa named by Carl Linnaeus