Blattulidae
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Blattulidae is an extinct family of
cockroach Cockroaches (or roaches) are a paraphyletic group of insects belonging to Blattodea, containing all members of the group except termites. About 30 cockroach species out of 4,600 are associated with human habitats. Some species are well-known as ...
es known from the Triassic to the Late Cretaceous. Their distinguishing characteristics include "
forewing Insect wings are adult outgrowths of the insect exoskeleton that enable insects to fly. They are found on the second and third thoracic segments (the mesothorax and metathorax), and the two pairs are often referred to as the forewings and hindwin ...
has long Sc, regular venation with distinct intercalaries and hindwing has simple CuP, branched A1."


Systematics

* †'' Anablatta'' Martins-Neto and Gallego 2007 Potrerillos Formation, Argentina
Ladinian The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between Ma and ~237 Ma (million years ago). The Ladinian was preceded by the Anisian and succeeded by the Carnian (part of the Upper or Late Triassic). ...
* †'' Argentinoblattula'' Martins-Neto and Gallego 2005 Los Rastros Formation, Argentina, Carnian * †'' Batola'' Vršanský 2009 Charentese amber, France, Albian * †'' Blattula'' Handlirsch 1906 40 species: Çakrazboz Formation, Turkey,
Rhaetian The Rhaetian is the latest age of the Triassic Period (in geochronology) or the uppermost stage of the Triassic System (in chronostratigraphy). It was preceded by the Norian and succeeded by the Hettangian (the lowermost stage or earliest age ...
Lilstock Formation The Lilstock Formation is a geologic formation in England. It preserves bivalve, insect and other invertebrate fossils, as well as fossil fish of '' Agkistracanthus mitgelensis'' and the basal theropod dinosaur '' Newtonsaurus cambrensis'' dating ...
, United Kingdom, Rhaetian Cattamarra Coal Measures, Australia,
Toarcian The Toarcian is, in the ICS' geologic timescale, an age and stage in the Early or Lower Jurassic. It spans the time between 182.7 Ma (million years ago) and 174.1 Ma. It follows the Pliensbachian and is followed by the Aalenian. The Toarcian ...
Guanyintan Formation, China,
Hettangian The Hettangian is the earliest age and lowest stage of the Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (My ...
Zaoshang Formation, China, Sinemurian, Shiti Formation, Hanshan Formation China, Bajocian
Jiulongshan Formation The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China. The formation consists of coarse conglomerates ...
, China, Callovian, Tuodian Formation, China, Oxfordian,
Posidonia Shale The Posidonia Shale (german: Posidonienschiefer, also called Schistes Bitumineux in Luxembourg) geologically known as the Sachrang Formation, is an Early Jurassic (Toarcian) geological formation of southwestern and northeast Germany, northern Swit ...
, " Green Series", Germany, Toarcian Karabastau Formation, Kazakhstan, Callovian Sulyukta Formation, Kyrgyzstan,
Toarcian The Toarcian is, in the ICS' geologic timescale, an age and stage in the Early or Lower Jurassic. It spans the time between 182.7 Ma (million years ago) and 174.1 Ma. It follows the Pliensbachian and is followed by the Aalenian. The Toarcian ...
Ulaan-Ereg Formation, Sharteg, Mongolia,
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
Cheremkhovskaya Formation Cheremkhovsky District (russian: Черемхо́вский райо́н) is an administrative district, one of the thirty-three in Irkutsk Oblast, Russia.Charter of Irkutsk Oblast, Article 13 Municipally, it is incorporated as Cheremkhovsk ...
, Russia, Toarcian Itat Formation, Russia, Bajocian Uda Formation, Russia, Oxfordian
Purbeck Group The Purbeck Group is an Upper Jurassic to Lower Cretaceous lithostratigraphic group (a sequence of rock strata) in south-east England. The name is derived from the district known as the Isle of Purbeck in Dorset where the strata are exposed in the ...
, United Kingdom,
Berriasian In the geological timescale, the Berriasian is an age/stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 145.0 ± 4.0 Ma and 139.8 ± 3.0 Ma (million years ago) ...
,
Doronino Formation Doronino (russian: Доронино) is a rural locality (a village) in Yugskoye Rural Settlement, Cherepovetsky District, Vologda Oblast, Russia. The population was 33 as of 2002. There are 4 streets. Geography Doronino is located southeast of ...
, Russia,
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
,
Lushangfen Formation The Lushangfen Formation is an Early Cretaceous aged geologic formation in China. Dinosaur remains are among the fossils that have been recovered from the formation, although none have yet been referred to a specific genus.Weishampel, et al. (2004 ...
, Yixian Formation, China,
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
,
Jinju Formation The Jinju Formation () is an Early Cretaceous geologic Formation (geology), formation in South Korea.Blattulites'' Vishnyakova 1982 Cheremkhovskaya Formation, Russia, Toarcian * †'' Elisama'' Giebel 1856
Haifanggou Formation The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China. The formation consists of coarse conglomerates ...
, China, Callovian/Oxfordian, Sharteg, Mongolia, Tithonian, Purbeck Group, United Kingdom, Berriasian, Crato Formation, Brazil,
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
Dzun-Bain Formation, Mongolia, Aptian * †'' Globula'' Vršanský 2009 Charentese amber, France, Albian * †'' Habroblattula'' Wang et al. 2007 Yixian Formation,
Laiyang Formation Laiyang city () is a county-level city within Yantai bordering Qingdao, located in the middle of the Shandong Peninsula, in Shandong province, China. The majority (70%) of its population are farmers and it is famous for producing the Laiyang pe ...
, China, Aptian * †'' Huablattula'' Qiu et al. 2019 Burmese amber, Myanmar,
Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...
* †'' Kridla'' Vršanský 2005 Kyndal Formation, Russia, Albian * †'' Macaroblattula'' Wang et al. 2007 Yixian Formation, China, Aptian * †'' Nula'' Vršanský 2008 Salignac amber, France, Cenomanian * †'' Ocelloblattula'' Anisyutkin and Gorochov 2008 Crato Formation, Brazil, Aptian
Lebanese amber Lebanese amber is fossilized resin found in Lebanon and southwest Syria. It dates back approximately 130-125 million years to the Barremian of the Early Cretaceous. It formed on what was then the northern coast of Gondwana, believed to be a tropica ...
, Barremian * †'' Svabula'' Vršanský 2005 Sharin-Gol Formation, Mongolia, Barremian * †'' Vrtula'' Vršanský 2008
Shinekhudag Formation The Shinekhudag Formation (also known as Shinekhudug Formation, Shinekhudag or Shinekhudukskaya Svita) is a geological formation in Dundgov, Mongolia whose strata date back to the Early Cretaceous. Dinosaur remains have been recovered from it.Wei ...
, Mongolia, Aptian * †'' Xonpepetla'' Cifuentes-Ruiz and Vršanský 2006
Cerro del Pueblo Formation The Cerro del Pueblo Formation is a geological Formation (geology), formation in Coahuila, Mexico whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel et al., ...
, Mexico, Campanian


References

{{Taxonbar, from=Q21347636 Cockroaches Prehistoric insect families