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The black falcon (''Falco subniger'') is a medium-large falcon that is endemic to
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
. It can be found in all mainland states and territories and yet is regarded as Australia's most under-studied falcon.Debus, S.J.S. & Olsen, J. (2011). Some aspects of the biology of the Black Falcon ''Falco subniger''. Corella 35: 29-36


Description

The information in this section is based on recent descriptions by several authors (notably, Debus & Davies 2012, Debus & Olsen 2011, Morcombe 2002 and Birds in Backyards n.d.).Debus, S.J. S & Davies, J. (2012). Birds of prey of Australia: a field guide (2nd ed). CSIRO Publishing, Collingwood, VicMorcombe, M. (2002). Field guide to Australian birds. Steve Parish Publishing, Archerfield, QldBirds in Backyards (n.d.). Black Falcon. Retrieved 19 October 2013 http://www.birdsinbackyards.net/species/Falco-subniger Size (adult, beak to tail): 45 – 56 cm (average 50 cm), tail makes up about half the length. Note: Females are larger than males - this is a form of sexual dimorphism. Average weight: 833 g (female) 582 g (male). Wing span: 95 –115 cm. Colouration is uniform dark brown to sooty black; juveniles are generally darker than adults; underwing feathers are two-tone (flight feathers slightly paler); adults may have an evident dark stripe below the eye. Occasionally the birds may have a white chin, spots on underwing coverts or bars on undertail coverts. Cere, eye ring and feet are pale grey (or pale blue-grey); eye is dark brown and beak tip is black. Talons are black. Chick is white down. The falcon's body is streamlined with a relatively long tail and slim build. Wings are long and pointed tapering toward wing tip.


Notes on identification

When attempting to identify a flying raptor Debus & Davies recommend concentrating on the bird's silhouette (including shape of wings and proportions), flight style and vocalisations, rather than on details of colouration. Australian falcons can be differentiated from
hawk Hawks are bird of prey, birds of prey of the family Accipitridae. They are widely distributed and are found on all continents except Antarctica. * The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks and others. Th ...
s by their toothed upper mandible (with a corresponding notch in their lower mandible) and by their long pointed wings.
Brown falcon The brown falcon (''Falco berigora'') is a relatively large falcon native to Australia and New Guinea. A number of plumage morphs occur, with the primary distinction being between the pale morph and the dark morph. Both morphs usually have dark ...
s (''Falco berigora'') are a common and widespread species in Australia, and they are the most likely to be confused with the black falcon (particularly brown falcon juveniles and black morphs). In fact, Debus & Olsen suggest many observations and behaviour of brown falcons have mistakenly been attributed to the black falcon, due to incorrect identification. The two species differ in the relative length of exposed tarsi (the lower, visible half of the bird's leg) compared with thigh feathers, flight style, wing attitude, and bold underwing barring and tail barring (which appear on the brown falcon only).Debus, S.J.S & Zuccon, A.E. (2013). Observations on hunting and breeding behaviour of the Black Falcon (''Falco subniger''). Sunbird 43(1): 12-26Morris, F.T. (1976). Birds of prey of Australia: A field guide. Lansdowne Editions, Melbourne, Vic


Taxonomy

The black falcon belongs to the family
Falconidae The falcons and caracaras are around 60 species of diurnal birds of prey that make up the family Falconidae (representing all extant species in the order Falconiformes). The family is divided into three subfamilies, Herpetotherinae, which inclu ...
, as do the other three falcon species which are found in Australia, the brown falcon,
grey falcon The grey falcon (''Falco hypoleucos'') is a medium-sized falcon native to Australia, possibly the rarest. It is uncommon throughout its range and is currently classified as Vulnerable. Taxonomy The description of the species was published by ...
(''Falco hypoleucos'') and
peregrine falcon The peregrine falcon (''Falco peregrinus''), also known as the peregrine, and historically as the duck hawk in North America, is a Cosmopolitan distribution, cosmopolitan bird of prey (Bird of prey, raptor) in the family (biology), family Falco ...
(''F. peregrinus''). Genetic analysis has revealed that the black falcon may be an early offshoot of the Old World
hierofalcon The hierofalcons are four closely related species of falcon which make up the subgenus ''Hierofalco'': * Lanner falcon, ''Falco biarmicus'' * Laggar falcon, ''Falco jugger'' * Saker falcon, ''Falco cherrug'' * Gyrfalcon, ''Falco rusticolus'' Th ...
s - such as the saker falcon (''F. cherrug'') and laggar falcon (''F. jugger'').


Population and threat status

The black falcon is listed by the IUCN as least concern. It is generally considered scarce, uncommon and nomadic but has at least one stronghold in inland Queensland. It is listed as Vulnerable in NSW and Victoria.NSW Government Office of Environment and Heritage (2013). Black Falcon - profile. Retrieved 20 October 2013 http://www.environment.nsw.gov.au/threatenedSpeciesApp/profile.aspx?id=20269 In 2009
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
estimated the black falcon's population as between 670 and 6700 mature individuals, with a stable population trend.BirdLife International (2013). Species factsheet: ''Falco subniger''. Retrieved 17 October 2013 http://www.birdlife.org/datazone/speciesfactsheet.php?id=3616&m=1


Distribution and habitat

The black falcon is widely distributed across mainland Australia, except densely forested areas. The species is only occasionally seen in the southern interior of WA and is sparse in coastal areas of south eastern Australia. BirdLife International also lists occasional, non-breeding vagrants to New Zealand. Total distribution area has been estimated at 5,910,000 km2. The black falcon's habitat is usually in the arid and semi arid zones. It is usually found near watercourses or utilizing patches of isolated trees. It hunts over open wooded grasslands, saltbush plains, bluebush plains and other low vegetation. In arid areas it will hunt over wetlands or near artificial or temporary water bodies – areas which tend to attract the most abundant birdlife.


Behaviour and ecology

The black falcon is known to both harass and be harassed by other bird species, including raptors and corvids. It may be found resting on power poles during the day but will not sit on wires.


Flight and hunting style

The falcon's flight has been described as variable. Generally it flies with relaxed ‘crow-like’ wingbeats with occasional soaring. It also flies with a stiffer, shorter wingbeat. It glides and soars with slightly drooped or sometimes horizontal wings, carpals forward (wings are stretched out straight when gliding). Tail is normally folded (except when soaring, when it is fanned with notched corners). It often soars for long periods without flapping and catches prey in the air or on the ground. In pursuit of prey it uses powerful, rapid wingbeats. While resting the bird's wing tips are shorter than the tail and legs appear short. The species usually hunts alone but has also been observed hunting cooperatively in pairs and occasionally in larger numbers, when prey is abundant (specifically, when stubble fires yield an abundant prey source. Twelve birds were seen at one such fire near Gundagai, NSW. They have often been observed using contour hunting (over treetops), surprising and taking birds on the wing. They will sometimes tail chase other birds over long distances, and less often engage in a Peregrine-like vertical stoop when hunting. The species preys upon and is sometimes the prey of other raptors.


Diet

The black falcon's diet primarily consists of bird species, from finch to cockatoo size but they have also been observed feeding on small mammals (i.e.
rabbit Rabbits, also known as bunnies or bunny rabbits, are small mammals in the family Leporidae (which also contains the hares) of the order Lagomorpha (which also contains the pikas). ''Oryctolagus cuniculus'' includes the European rabbit speci ...
s,
mice A mouse ( : mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus' ...
and
rat Rats are various medium-sized, long-tailed rodents. Species of rats are found throughout the order Rodentia, but stereotypical rats are found in the genus ''Rattus''. Other rat genera include ''Neotoma'' ( pack rats), ''Bandicota'' (bandicoot ...
s), insects and carrion. Avian prey species include: * various pigeons (including crested pigeon ''Ocyphaps lophotes'' and the feral rock dove ''Columba livia'') *
Australian pipit The Australian pipit (''Anthus australis'') is a fairly small passerine bird of open country in Australia and New Guinea. It belongs to the pipit genus ''Anthus'' in the family Motacillidae. It was formerly lumped together with the Richard's, A ...
(''Anthus australis'') * Australian magpie (''Gymnorhina tibicen'') * Australian ringneck (''Barnardius zonarius'') *
Black-faced Cuckooshrike The black-faced cuckooshrike (''Coracina novaehollandiae'') is a common omnivorous passerine bird native to Australia and southern New Guinea. It has a protected status in Australia, under the National Parks and Wildlife Act, 1974. They are wid ...
(''Coracina novaehollandiae'') *
Blue bonnet The blue bonnet was a type of soft woollen hat that for several hundred years was the customary working wear of Scottish labourers and farmers. Although a particularly broad and flat form was associated with the Scottish Lowlands, where it was so ...
(''Northiella haematogaster'') * Budgerigar (''Melopsittacus undulatus'') *
Common starling The common starling or European starling (''Sturnus vulgaris''), also known simply as the starling in Great Britain and Ireland, is a medium-sized passerine bird in the starling family, Sturnidae. It is about long and has glossy black plumage ...
(''Sturnus vulgaris'') * Eastern rosella (''Platycercus eximius'') * Fairy martin (''Petrochelidon ariel'') * Galah (''Eolophus roseicapillus'') * Grey Teal (''Anas gracilis'') * Honeyeaters (Meliphagidae) * Horsfield's bush lark (''Mirafra javanica'') *
Little buttonquail The little buttonquail (''Turnix velox'') is a species of buttonquail, part of a small family of birds which resemble, but are unrelated to, the true quails. This species is resident in Australia, where it is one of the more common buttonquails. ...
(''Turnix velox'') * Little corella (''Cacatua sanguinea'') *
Musk lorikeet The musk lorikeet (''Glossopsitta concinna'') is a lorikeet, now the only species in the genus ''Glossopsitta''. It inhabits south-central/eastern Australia. The little lorikeet and the purple-crowned lorikeet were previously included in the ge ...
(''Glossopsitta concinna'') * Stubble quail (''Coturnix pectoralis'') * Zebra finch (''Taeniopygia guttata'') Debus & Zuccon also observed the black falcon hunting turquoise parrot (''Neophema pulchella'') and
apostlebird The apostlebird (''Struthidea cinerea''), also known as the grey jumper, lousy jack, happy jack, Donny bird or caw bird is a quick-moving, gray or black bird about 33 cm (13 in) long. It is a native to Australia where it roams woodlands ...
(''Struthidea cinerea''), but on these occasions the attacks were unsuccessful.


Breeding

Laying time: between May and November (usually July to September). Clutch size: 1 - 5 (usually 3 or 4). Egg size: 42x32 mm. Incubation: approximately 34 days (by female, possibly by male for short periods). Nestling period: 5.5 – 7 weeks. Life span: at least 12 years in the wild, and 20 years in captivity. The black falcon nests in living or dead trees, using the stick nests of corvids or other raptors. It may take over nests occupied by other birds. It appears there may be interspecific competition for nest sites between the black falcon and other raptors and corvids.Debus, S.J.S. & Tsang, L.R. (2011). Notes on Black Falcons ''Falco subniger'' breeding near Tamworth, New South Wales. Australian Field Ornithology 28: 13-26 Nest sites may be a limiting factor for black falcons, specifically where there has large-scale vegetation clearing, such as occurred in the sheep-wheat belt of NSW. During breeding season males perform courtship displays such as horizontal figure eights around the nest. The male bring food to the female during incubation and brooding. During the later nestling period both sexes may forage to feed the young. Post-fledging behaviour and development of young falcons appears to be very similar to that of the peregrine falcon, and includes spending time with siblings and parents, and practicing hunting, territorial and courtship behaviours.


Vocalisations

The most common call is similar to the 'cackle' of the peregrine falcon but is slower and deeper – ‘gaak-gaar-gaak’. The guttural call is shorter and more rapid when being attacked or when intruders are nearby – ‘gak-gak-gak-gak’. Some authors suggest the bird screams when attacking prey but this attribution could be a result of misidentifications of brown falcons, with other authors contesting that the birds are generally silent during attack. The male also makes a sharp, rather high-pitched ‘eeik..eeik..eeik’ or ‘ee-chip…ee-chip’ during courtship displays, and the female has a whine or wail when begging for food or copulating.


Home range, dispersal

The species’ home range is undetermined, but likely to be greater than 100 km2. The species appears to be wide-ranging in the non-breeding season, but is also known to stay in regular non-breeding territories for lengthy periods.


Threats and management

Declining numbers of black falcons is a growing concern, with two states (Victoria and New South Wales) already having listed the species as vulnerable. Several authors have pointed out that population and other ecological data on the species is deficient, and addressing this issue should be a major priority for managers in the future. The primary threats to the species are all
anthropogenic Anthropogenic ("human" + "generating") is an adjective that may refer to: * Anthropogeny, the study of the origins of humanity Counterintuitively, anthropogenic may also refer to things that have been generated by humans, as follows: * Human im ...
. These include clearing of habitat, degradation of habitat by over grazing, collision with vehicles and shooting. Clearing of big, old trees and breeding sites in riparian areas is of particular concern. The species also faces competition from corvids and other raptors over nest sites and possibly prey. Disturbance of nest sites and harassment by other birds (including cockatoos) also appears to impact the falcon's breeding success. Recommendations for ongoing management include monitoring of populations and further research into its biology and ecology. The Office of Environment and Heritage (NSW Government) has identified eight priority actions that aim to help recover the species. These include: protecting and monitoring known nest sites; protecting old stick nests that could make suitable breeding sites for the species; protecting and recruiting large old trees; engaging landholders in the management of habitat; expanding habitat (particularly along riparian areas); research into dietary requirements and awareness raising.NSW Government Office of the Environment and Heritage (n.d.). Priority actions by type of threatened species: Black Falcon. Retrieved 24 October 2013 from http://www.environment.nsw.gov.au/threatenedSpeciesApp/PasSearchSpecies.aspx?speciesName=Black+Falcon&generalType=Birds Victoria does not appear to have a recovery plan in place for the black falcon.


References


External links


Atlas of Living Australia - ''Falco subniger''
{{Taxonbar, from=Q286194 black falcon Endemic birds of Australia Diurnal raptors of Australia black falcon black falcon