Black-throated Brilliant
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The black-throated brilliant (''Heliodoxa schreibersii'') is a species of
hummingbird Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 361 species and 113 genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are found in the tropics aro ...
in the "brilliants", tribe
Heliantheini Heliantheini is one of the two tribes that make up the subfamily Lesbiinae of the hummingbird family Trochilidae. The other tribe in the subfamily is Lesbiini. The informal name "brilliants" has been proposed for this group as it includes the ge ...
in subfamily
Lesbiinae Lesbiinae is one of the six subfamily, subfamilies that make up the hummingbird family (biology), family Trochilidae. The subfamily is divided into two Tribe (biology), tribes: Heliantheini ("brilliants") containing 14 genera and Lesbiini ("coqu ...
. It is found in
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
,
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
,
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
, and
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
.HBW and BirdLife International (2020) ''Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world'' Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip xls zipped 1 MBretrieved 27 May 2021


Taxonomy and systematics

The taxonomy of the black-throated brilliant has not been settled. The
International Ornithological Committee The International Ornithologists' Union, formerly known as the International Ornithological Committee, is a group of about 200 international ornithologists, and is responsible for the International Ornithological Congress and other international ...
(IOC), the
Clements taxonomy ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 2022 ...
, and the South American Classification Committee of the
American Ornithological Society The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its m ...
(SACC) assign two subspecies, the nominate ''H. s. schreibersii'' and ''H. s. whitelyana''.Clements, J. F., T. S. Schulenberg, M. J. Iliff, S. M. Billerman, T. A. Fredericks, J. A. Gerbracht, D. Lepage, B. L. Sullivan, and C. L. Wood. 2021. The eBird/Clements checklist of Birds of the World: v2021. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ Retrieved August 25, 2021Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 31 January 2022. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved February 1, 2022
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
's
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. T ...
(HBW) treats those taxa as full species, the "black-throated" and "black-breasted" brilliants respectively. In addition, for a time in the early 20th century, the two subspecies were treated as species, the only members of genus ''Ionolaima'', which was subsequently merged into the present ''Heliodoxa''. As of May 2022 the SACC has a proposal pending to resurrect ''Ionolaima'' but it has gained little support.


Description

The nominate subspecies ''H. s. schreibersii'' of black-throated brilliant is long. Males weigh about and females . Both have an almost straight dark bill about long and a small white spot behind the eye. The male has shining green upperparts with a glittering green forehead. Its underparts are black with a small glittering purple patch and narrow glittering green band on the lower throat. The steel blue tail is very long and deeply forked. The female's upperparts are like the male's, and in addition it has a whitish to rufous malar stripe. Its underparts are gray with bronzy green spots. Its central tail feathers are green and the tail is less deeply forked than the male's. Juveniles resemble females with a more intensely colored malar stripe.del Hoyo, J., I. Heynen, N. Collar, P. F. D. Boesman, and G. M. Kirwan (2020). Black-throated Brilliant (''Heliodoxa schreibersii''), version 1.0. In Birds of the World (S. M. Billerman, B. K. Keeney, P. G. Rodewald, and T. S. Schulenberg, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.bltbri1.01 retrieved 7 May 2022 Subspecies ''H. s. whitelyana'' is long, also with a long bill. The male's entire upperparts are shining green including the forehead. Its underparts and tail are the same as the nominate's. Females and juveniles are the same as those of the nominate subspecies.


Distribution and habitat

The nominate subspecies of black-throated brilliant is found from southeastern Colombia south through eastern Ecuador into northeastern Peru as far as
San Martín San Martín or San Martin may refer to: People Saints * Saint Martin (disambiguation)#People, name of various saints in Spanish Political leaders *Vicente San Martin (1839 -1901), Military, National hero of Mexico. *Basilio San Martin (1849 ...
Department and also east into the upper Rio Negro watershed in extreme northwestern Brazil. It inhabits the interior of mature humid forest and scrublands. In elevation it mostly ranges between . However, in eastern Ecuador it is found locally as high as and occasionally up to . ''H. s. whitelyana'' is found in central and southeastern Peru. It inhabits humid
montane forest Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
at elevations between . (The map shows only the range of the nominate subspecies.)


Behavior


Movement

The nominate subspecies of black-throated brilliant is known to be sedentary and ''H. s. whitelyana'' is presumed to be.


Feeding

The black-throated brilliant primarily forages for nectar in the forest's understory, typically between high. Flowering plants of families
Ericaceae The Ericaceae are a family of flowering plants, commonly known as the heath or heather family, found most commonly in acidic and infertile growing conditions. The family is large, with c.4250 known species spread across 124 genera, making it th ...
and
Malvaceae Malvaceae, or the mallows, is a family of flowering plants estimated to contain 244 genera with 4225 known species. Well-known members of economic importance include okra, cotton, cacao and durian. There are also some genera containing familiar ...
are among its targets. It also captures small insects by
hawking Hawking may refer to: People * Stephen Hawking (1942–2018), English theoretical physicist and cosmologist *Hawking (surname), a family name (including a list of other persons with the name) Film * ''Hawking'' (2004 film), about Stephen Haw ...
from a perch.


Breeding

The breeding season of the nominate black-throated brilliant spans at least from February to May but may extend as late as October. The female incubates the clutch of two eggs. Nothing else is known about its breeding
phenology Phenology is the study of periodic events in biological life cycles and how these are influenced by seasonality, seasonal and interannual variations in climate, as well as environmental factor, habitat factors (such as elevation). Examples includ ...
or that of subspecies ''H. s. whitelyana''.


Vocalization

What is thought to be the song of the nominate black-throated brilliant is "a repeated, drawn-out, descending reeling trill of c. 4–5 seconds." It also makes single "chup" notes. The presumed song of ''H. s. whitelyana'' is a similar trill lasting about 3 seconds, but there are too few recordings to be sure if it is truly different from the nominate's.


Status

The
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
follows HBW taxonomy and so has treated the two taxa as separate species. Both are assessed as being of Least Concern. The population size of neither is known and both are believed to be decreasing. No immediate threats have been identified. Both taxa appear to be uncommon and patchily distributed, but there are few reliable data from which to make decisions.


References

{{Taxonbar, from=Q1261008 black-throated brilliant Hummingbird species of South America Birds of the Colombian Amazon Birds of the Ecuadorian Amazon Birds of the Peruvian Amazon black-throated brilliant Taxonomy articles created by Polbot