The big-scale sand smelt (''Atherina boyeri'') is a species of

fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...

in the family

Atherinidae The Old World silversides are a family, Atherinidae, of fish in the order Atheriniformes. Atherinidae are abundant and considered bony fish (teleost) that are widespread globally, living in rivers, estuaries, and coastal waters. They occur worl ...

. It is a

euryhaline Euryhaline organisms are able to adapt to a wide range of salinities. An example of a euryhaline fish is the molly (''Poecilia sphenops'') which can live in fresh water, brackish water, or salt water. The green crab (''Carcinus maenas'') is an e ...

amphidromous fish Fish migration is animal migration, mass relocation by fish from one area or body of water to another. Many types of fish migrate on a regular basis, on time scales ranging from daily to annually or longer, and over distances ranging from a few ...

, up to 20 cm in length.

Description

It is a small pelagic fish species which occurs near the surface in the littoral estuarine zone: in lagoons, salt marshes (77 psu), shallow brackish areas (2 psu) and inland waters which are rather unsuitable for other fish species, due to their high ionic strength and salinity. Body is rather long, slender, moderately flattened. Eyes are large. Head and body are scaly. Mouth is protractible, upwardly directed, with small teeth. Lower jaw has an upper expansion within mouth (high dentary bone). There are two separate dorsal fins, with all rays of first and 1-2 anterior rays of second dorsal fin being unsegmented. The anal fin is similar to the second dorsal fin, while the caudal fin is forked. The first dorsal fin has 6-10 flexible spines. It is an omnivorous species feeding on zoo-plankton and small bottom-living animals (

crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

gammarids Gammaridae is a family of amphipods. In North America they are included among the folk taxonomic category of " scuds", and otherwise gammarids is usually used as a common name. They have a wide distribution, centered on Eurasia, and are eury ...

polychaete Polychaeta () is a paraphyletic class (biology), class of generally marine invertebrate, marine annelid worms, common name, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that ...

worms and

molluscs Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant taxon, extant species of molluscs are recognized. The number of fossil sp ...

Range

It is found in the eastern

Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...

from

Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...

and

Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...

Nouadhibou Nouadhibou (; ar, نواذيبو, Nwādībū, Berber: Nwadibu, formerly in French: ) is the second largest city in Mauritania and serves as a major commercial centre. The city itself has about 118,000 inhabitants expanding to over 140,000 in the ...

(

Mauritania Mauritania (; ar, موريتانيا, ', french: Mauritanie; Berber: ''Agawej'' or ''Cengit''; Pulaar: ''Moritani''; Wolof: ''Gànnaar''; Soninke:), officially the Islamic Republic of Mauritania ( ar, الجمهورية الإسلامية ...

) and

Madeira ) , anthem = ( en, "Anthem of the Autonomous Region of Madeira") , song_type = Regional anthem , image_map=EU-Portugal_with_Madeira_circled.svg , map_alt=Location of Madeira , map_caption=Location of Madeira , subdivision_type=Sovereign st ...

. Also it occurs in the

Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the e ...

, including the inshore

lagoon A lagoon is a shallow body of water separated from a larger body of water by a narrow landform, such as reefs, barrier islands, barrier peninsulas, or isthmuses. Lagoons are commonly divided into ''coastal lagoons'' (or ''barrier lagoons'') a ...

s, such as

Trasimeno Lake Trasimeno ( , also ; it, Lago Trasimeno ; la, Trasumennus; ett, Tarśmina), also referred to as Trasimene ( ) or Thrasimene in English, is a lake in the province of Perugia, in the Umbria region of Italy on the border with Tuscany. Th ...

and Lesina in

Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...

Hyères Hyères (), Provençal Occitan: ''Ieras'' in classical norm, or ''Iero'' in Mistralian norm) is a commune in the Var department in the Provence-Alpes-Côte d'Azur region in southeastern France. The old town lies from the sea clustered around t ...

in the southern

France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...

such as

Marseille Marseille ( , , ; also spelled in English as Marseilles; oc, Marselha ) is the prefecture of the French department of Bouches-du-Rhône and capital of the Provence-Alpes-Côte d'Azur region. Situated in the camargue region of southern Franc ...

and Lake Qarun in

Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Mediter ...

; an isolated population is found near the coasts of

England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...

and the

Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...

. In the

Black Sea The Black Sea is a marginal mediterranean sea of the Atlantic Ocean lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bounded by Bulgaria, Georgia, Roma ...

, it is widespread along all coasts, in lagoons and

estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environment ...

, in the downstreams of rivers

Danube The Danube ( ; ) is a river that was once a long-standing frontier of the Roman Empire and today connects 10 European countries, running through their territories or being a border. Originating in Germany, the Danube flows southeast for , pa ...

Dniester The Dniester, ; rus, Дне́стр, links=1, Dnéstr, ˈdⁿʲestr; ro, Nistru; grc, Τύρᾱς, Tyrās, ; la, Tyrās, la, Danaster, label=none, ) ( ,) is a transboundary river in Eastern Europe. It runs first through Ukraine and th ...

Southern Bug , ''Pivdennyi Buh'' , name_etymology = , image = Sunset S Bug Vinnitsa 2007 G1.jpg , image_size = 270 , image_caption = Southern Bug River in the vicinity of Vinnytsia, Ukraine , map = PietinisBug ...

Inhulets The Inhulets () is a river, a right tributary of the Dnieper, that flows through Ukraine. It has a length of 557 km and a drainage basin of 14,460 km². The Inhulets has its source in the Dnieper Upland in a ravine (balka) to the we ...

, and

Dnieper } The Dnieper () or Dnipro (); , ; . is one of the major transboundary rivers of Europe, rising in the Valdai Hills near Smolensk, Russia, before flowing through Belarus and Ukraine to the Black Sea. It is the longest river of Ukraine and B ...

, with a permanent population is in the

Kakhovka Reservoir The Kakhovka Reservoir (, ''Kakhovs′ke vodoskhovyshche'') is a water reservoir on the Dnieper River in Ukraine. It was created in 1956, when the Kakhovka Hydroelectric Power Plant was built. It is one of several reservoirs in the Dnieper reservo ...

.Movchan Yu.V. (1988) True loaches,

catfishes Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive, ...

, canal catfishes, freshwater eels, congers, needlefishes,

cods Cod is the common name for the demersal fish genus '' Gadus'', belonging to the family Gadidae. Cod is also used as part of the common name for a number of other fish species, and one species that belongs to genus ''Gadus'' is commonly not call ...

stickleback The sticklebacks are a family of ray-finned fishes, the Gasterosteidae which have a Holarctic distribution in fresh, brackish and marine waters. They were thought to be related to the pipefish and seahorses but are now thought to be more closel ...

s, syngnathids, mosquitofishes, zeids,

barracuda A barracuda, or cuda for short, is a large, predatory, ray-finned fish known for its fearsome appearance and ferocious behaviour. The barracuda is a saltwater fish of the genus ''Sphyraena'', the only genus in the family Sphyraenidae, which was ...

grey mullets Grey mullet can mean any of several fish in the family ''Mugilidae'' (the mullets) and having a greyish hue: * Flathead grey mullet, ''Mugil cephalus'' * Thicklip grey mullet, ''Chelon labrosus'' * Boxlip grey mullet, ''Oedalechilus labeo The bo ...

old world silverside The Old World silversides are a family, Atherinidae, of fish in the order Atheriniformes. Atherinidae are abundant and considered bony fish (teleost) that are widespread globally, living in rivers, estuaries, and coastal waters. They occur worldw ...

s, cusk eels n:''Fauna of Ukraine'', Vol. 8, No 3,

Kiev Kyiv, also spelled Kiev, is the capital and most populous city of Ukraine. It is in north-central Ukraine along the Dnieper, Dnieper River. As of 1 January 2021, its population was 2,962,180, making Kyiv the List of European cities by populat ...

: Naukova Dumka, 399 pp. (in Russian). The isolated population in the

Caspian Sea The Caspian Sea is the world's largest inland body of water, often described as the world's largest lake or a full-fledged sea. An endorheic basin, it lies between Europe and Asia; east of the Caucasus, west of the broad steppe of Central Asia ...

is characterised as

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

''A. b. caspia'' (Eichwald, 1838).

Fishing

The major small-scale fishing gears exploiting this species are coastal beach seines, small mesh size (10 mm)

gill nets Gillnetting is a fishing method that uses gillnets: vertical panels of netting that hang from a line with regularly spaced floaters that hold the line on the surface of the water. The floats are sometimes called "corks" and the line with corks is ...

and lift-nets. It is often used as bait fish on small and medium longlines, handlines, fishing using rods and

reels A reel is an object around which a length of another material (usually long and flexible) is wound for storage (usually hose are wound around a reel). Generally a reel has a cylindrical core (known as a '' spool'') with flanges around the ends ...

, as

trolling In slang, a troll is a person who posts or makes inflammatory, insincere, digressive, extraneous, or off-topic messages online (such as in social media, a newsgroup, a forum, a chat room, a online video game), or in real life, with the i ...

bait, even as bait in

fish trap A fish trap is a trap used for fishing. Fish traps include fishing weirs, lobster traps, and some fishing nets such as fyke nets. Traps are culturally almost universal and seem to have been independently invented many times. There are two main ...

Gastronomy

This small fish is appreciated in the

Spanish Spanish might refer to: * Items from or related to Spain: **Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many Latin American countries **Spanish cuisine Other places * Spanish, Ontario, Cana ...

, French, Turkish, and

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

cuisines. The fish are lightly powdered with

wheat flour Wheat flour is a powder made from the grinding of wheat used for human consumption. Wheat varieties are called "soft" or "weak" if gluten content is low, and are called "hard" or "strong" if they have high gluten content. Hard flour, or ''bread ...

before being fried in hot

olive oil Olive oil is a liquid fat obtained from olives (the fruit of ''Olea europaea''; family Oleaceae), a traditional tree crop of the Mediterranean Basin, produced by pressing whole olives and extracting the oil. It is commonly used in cooking: f ...

Etymology

The specific name of this species honours the Medieval poet and scientist Guillaume Boyer who was a native of

Nice Nice ( , ; Niçard: , classical norm, or , nonstandard, ; it, Nizza ; lij, Nissa; grc, Νίκαια; la, Nicaea) is the prefecture of the Alpes-Maritimes department in France. The Nice agglomeration extends far beyond the administrative c ...

, the type locality for this species.

References

{{Taxonbar, from=Q301035

big-scale sand smelt The big-scale sand smelt (''Atherina boyeri'') is a species of fish in the family Atherinidae. It is a euryhaline amphidromous fish, up to 20 cm in length. Description It is a small pelagic fish species which occurs near the surface in th ...

Fish of the Mediterranean Sea Fish of the North Sea Fish of the Black Sea Fish of the Caspian Sea Fish of Europe

Taxonomy articles created by Polbot Taxa named by Antoine Risso