Belligerent Sculpin
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The belligerent sculpin (''Megalocottus platycephalus''), or flathead sculpin, is a species of marine ray-finned fish belonging to the family Cottidae, the typical sculpins. This species occurs in the northern Pacific Ocean.


Taxonomy

The belligerent sculpin was first formally described in 1814 as ''Cottus platycephalus'' by the German zoologist Peter Simon Pallas with its
type locality Type locality may refer to: * Type locality (biology) * Type locality (geology) See also * Local (disambiguation) * Locality (disambiguation) {{disambiguation ...
given as Kamchatka and the
Sea of Okhotsk The Sea of Okhotsk ( rus, Охо́тское мо́ре, Ohótskoye móre ; ja, オホーツク海, Ohōtsuku-kai) is a marginal sea of the western Pacific Ocean. It is located between Russia's Kamchatka Peninsula on the east, the Kuril Islands ...
. In 1861 the American
biologist A biologist is a scientist who conducts research in biology. Biologists are interested in studying life on Earth, whether it is an individual cell, a multicellular organism, or a community of interacting populations. They usually specialize in ...
Theodore Gill classified this species in the monospecific genus ''
Megalocottus ''Megalocottus'' is a small genus of marine ray-finned fishes belonging to the family Cottidae, the typical sculpins. These fishes are found in the western Pacific Ocean. Taxonomy ''Megalocottus'' was first proposed as a monospecific genu ...
''. Some authorities consider the belligerent sculpin to be the still the only species in its genus, either recognising the
Southern flathead sculpin The southern flathead sculpin (''Megalocottus taeniopterus'') is a species of marine ray-finned fish belonging to the family Cottidae, the typical sculpins. This species occurs in the northern Pacific Ocean. Taxonomy The southern flathead sc ...
(''M. taeniopterus'') as a
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
or
junior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linna ...
of ''M. platycephalus''. Fishbase still recognises two species within the genus ''Megalocottus''. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
''playcephalus'' means "flat head", a reference to the wide, flat head with a projecting lower jaw.


Description

The belligerent sculpin has a broad, strongly flattened head, much wider than it is deep with a projecting lower jaw. The top and the sides of the head are very warty and there are large spines or bumps behind the eye and on the nape, these may have short cirri on them. The first and second spines on the
preoperculum This glossary of ichthyology is a list of definitions of terms and concepts used in ichthyology, the study of fishes. A B C ...
are curved, the third and fourth are poorly developed. There are irregular rows of spiny plates over and under the
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...
. The colour is grey-brown marked with yellow or white spotting and blotching. The fins have spots and bars. This species reaches a maximum published total length of .


Distribution and habitat

The belligerent sculpin is found in the northern Pacific Ocean. Its range extends from the Sea of Japan, through the Sea of Okhotsk and the northern Kuril Islands and the southeastern Kamchatka Peninsula east to the
Alaska Peninsula The Alaska Peninsula (also called Aleut Peninsula or Aleutian Peninsula, ale, Alasxix̂; Sugpiaq: ''Aluuwiq'', ''Al'uwiq'') is a peninsula extending about to the southwest from the mainland of Alaska and ending in the Aleutian Islands. The ...
and north along the
Bering Sea The Bering Sea (, ; rus, Бе́рингово мо́ре, r=Béringovo móre) is a marginal sea of the Northern Pacific Ocean. It forms, along with the Bering Strait, the divide between the two largest landmasses on Earth: Eurasia and The Ameri ...
coasts of Alaska and Russia to the
Chukchi Sea Chukchi Sea ( rus, Чуко́тское мо́ре, r=Chukotskoye more, p=tɕʊˈkotskəjə ˈmorʲɪ), sometimes referred to as the Chuuk Sea, Chukotsk Sea or the Sea of Chukotsk, is a marginal sea of the Arctic Ocean. It is bounded on the west b ...
. It is a
demersal fish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They occ ...
, typically found in the subtidal zone down to depths of , although has been claimed. It is found on substrates of sand, mud and gravel, especially where there are beds of '' Fucus'' and ''
Zostera ''Zostera'' is a small genus of widely distributed seagrasses, commonly called marine eelgrass, or simply seagrass or eelgrass, and also known as seaweed by some fishermen and recreational boaters including yachtsmen. The genus ''Zostera'' con ...
''. It is generally found in brackish water close to river mouths, near to the shore and in shallow water.


Biology

The belligerent sculpin's biology is not well known. They are oviparous. In the Spring they move closer inshore and into estuaries, even into the lower reaches of rivers. The juveniles feed on algae, and crustaceans such as
copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...
s, cumaceans,
amphipod Amphipoda is an order of malacostracan crustaceans with no carapace and generally with laterally compressed bodies. Amphipods range in size from and are mostly detritivores or scavengers. There are more than 9,900 amphipod species so far descr ...
s, and mysids while the adults have a similar diet but they also eat small fishes.


References

{{Taxonbar, from=Q2044901 Belligerent sculpin Fish described in 1814 Taxa named by Peter Simon Pallas