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''Beesia'' is a genus of
flowering plants Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
in the
buttercup family Ranunculaceae (buttercup or crowfoot family; Latin "little frog", from "frog") is a family of over 2,000 known species of flowering plants in 43 genera, distributed worldwide. The largest genera are ''Ranunculus'' (600 species), ''Delphinium' ...
. It was named in 1915 after the plant nursery firm
Bees of Chester Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyletic lineage within the superfamil ...
, who financed the plant hunting trips of
George Forrest George Forrest may refer to: *G. Topham Forrest (George Topham Forrest, 1872–1945), principal architect for the London County Council *George Forrest (author) (1915–1999), American author and musician *George Forrest (botanist) (1873–1932), S ...
and
Frank Kingdon-Ward Francis Kingdon-Ward, born Francis Kingdon Ward OBE, (6 November 1885 in Manchester – 8 April 1958) was an English botanist, explorer, plant collector and author. He published most of his books as Frank Kingdon-Ward and this hyphenated form ...
in China.


Species

There are two
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of ''Beesia'': *''Beesia calthifolia'', which is
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to the following WGSRPD floristic regions: North-Central
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
, South-Central China, Southeast China, East Himalaya, and
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
. *''Beesia deltophylla'', which is native to southeastern
Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa people, ...
.


Description


Morphology

Beesia is an
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
perennial and grows as a dense basal rosette of heart-shaped
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
with delicate serrated edges. Leaves are soft and waxy; new growth of ''B. deltophylla'' flushes dark green to black, while ''B. calthifolia'' is a lighter shade of green. Small white
flowers A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
bloom mid-summer off an upright spike. Flowers are star-like: although ''Beesia'' flowers do not have petals, they do have 5 elliptic, petal-like
sepals A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
. Flowers also have many stamens.


Phytochemistry

A class of
organic molecules In chemistry, organic compounds are generally any chemical compounds that contain carbon-hydrogen or carbon-carbon bonds. Due to carbon's ability to catenate (form chains with other carbon atoms), millions of organic compounds are known. The s ...
termed ''beesiosides'' have been isolated from ''Beesia'' plants. Beesiosides are
cycloartane Cycloartane is a triterpene, also known as 4,4,14-trimethyl-9,19-cyclo-5alpha,9beta-cholestane. Its derivative cycloartenol is the starting point for the synthesis of almost all plant steroids. See also * Lanostane * Cycloastragenol Cycloastr ...
glycosides In chemistry, a glycoside is a molecule in which a sugar is bound to another functional group via a glycosidic bond. Glycosides play numerous important roles in living organisms. Many plants store chemicals in the form of inactive glycosides. ...
(a type of triterpene that forms a
sugar Sugar is the generic name for sweet-tasting, soluble carbohydrates, many of which are used in food. Simple sugars, also called monosaccharides, include glucose, fructose, and galactose. Compound sugars, also called disaccharides or double ...
and a non-sugar upon
hydrolysis Hydrolysis (; ) is any chemical reaction in which a molecule of water breaks one or more chemical bonds. The term is used broadly for substitution reaction, substitution, elimination reaction, elimination, and solvation reactions in which water ...
).


Phylogeny

''
Anemonopsis ''Anemonopsis'', the false anemone, is a monotypic genus in the family Ranunculaceae, containing only the species ''Anemonopsis macrophylla'', endemic to Japan's main island of Honshu. The generic name ''Anemonopsis'' refers to it being ''Ane ...
'', a monotypic genus in the Ranunculaceae native to
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
, is a
sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...
to ''Beesia.'' ''
Eranthis ''Eranthis'' is a genus of eight species of flowering plants in the buttercup family (biology), family Ranunculaceae, native plant, native to southern Europe and east across Asia to Japan. The common name winter aconite comes from the early flowe ...
'' and '' Actaea'' are also closely related to ''Beesia''.


Uses


Ornamental

''Beesia'' is easily divided or grown from seed, and can be grown in USDA hardiness zones 6a to 8b. It grows best in partial shade or filtered light in rich, moist
soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former te ...
.


Medicinal

''Beesia''
rhizomes In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
are used in
Chinese herbal medicine Chinese herbology () is the theory of traditional Chinese herbal therapy, which accounts for the majority of treatments in traditional Chinese medicine (TCM). A ''Nature'' editorial described TCM as "fraught with pseudoscience", and said that t ...
as a treatment for
rheumatoid arthritis Rheumatoid arthritis (RA) is a long-term autoimmune disorder that primarily affects joints. It typically results in warm, swollen, and painful joints. Pain and stiffness often worsen following rest. Most commonly, the wrist and hands are involv ...
and
influenza Influenza, commonly known as "the flu", is an infectious disease caused by influenza viruses. Symptoms range from mild to severe and often include fever, runny nose, sore throat, muscle pain, headache, coughing, and fatigue. These symptoms ...
.


References


External links


GRIN Genus Profile
Ranunculaceae Ranunculaceae genera Taxa named by William Wright Smith Taxa named by Isaac Bayley Balfour {{ranunculales-stub