''Barsboldia'' (meaning "of Barsbold", a well-known Mongolian

paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...

) is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

of large hadrosaurid

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

from the

early Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval fro ...

Nemegt Formation The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians, ...

of Ömnogöv',

Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...

. It is known from a partial

vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristi ...

l column, partial pelvis, and some

ribs The rib cage, as an enclosure that comprises the ribs, vertebral column and sternum in the thorax of most vertebrates, protects vital organs such as the heart, lungs and great vessels. The sternum, together known as the thoracic cage, is a semi- ...

Discovery

In 1970, a Polish-Mongolian expedition near the Nemegt found the skeleton of an ornithopod and first assigned it to '' Saurolophus angustirostris''. However, two Polish paleontologists,

Teresa Maryańska Teresa Maryańska (1937 – 3 October 2019) was a Polish paleontologist who specialized in Mongolian dinosaurs, particularly pachycephalosaurians and ankylosaurians. Peter Dodson (1998 p. 9) states that in 1974 Maryanska together with Hals ...

and

Halszka Osmólska Halszka Osmólska (September 15, 1930 – March 31, 2008) was a Polish paleontologist who had specialized in Mongolian dinosaurs. Biography She was born in 1930 in Poznań. In 1949, she began to study biology at Faculty of Biology and Earth Scie ...

, came to the conclusion that it was a lambeosaurine that had to represent a separate species. They named and described ''Barsboldia sicinskii'' based on the holotype specimen ZPAL MgD-1/110. The genus name honors the Mongolian paleontologist Rinchen Barsbold, while the species name honors Wojciech Siciński, the technician at the Warsaw Paleobiological Institute who prepared the skeleton. The holotype specimen, found in a layer of the

dating to the early Maastrichtian, consisted of a partial skeleton consisting of nine back

e, nine hip vertebrae, fifteen tail vertebrae, a left ilium, parts of the left and right pubis, several ribs, and a few fragments of the hind limbs, with the backbone largely articulated. The anterior and posteriormost portions of the skeleton were lost due to erosion.

Description

Barsboldia was a large hadrosaur, previously measured in length and in body mass. In 2011, the tibial length was measured at , rivalling that of '' Shantungosaurus'' measured at and that of '' Magnapaulia'' measured at ; this indicates that ''Barsboldia'' could have possibly reached within the range of in total body length. The most distinctive features of ''Barsboldia'' are found in the

neural spines The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...

. These are very tall, particularly over the hips, and were described as second only to those of '' Hypacrosaurus altispinus'' and the tips of those found in the first few vertebrae of the tail are club-shaped, possibly a sign of old age.

Phylogeny

Maryańska and Osmólska described their new genus as a lambeosaurine (or hollow-crested duckbill), the first from the Nemegt Formation, although it lacked a skull. However, the sacrum has a keel along the bottom, a possible lambeosaurine feature, and the bones closely resemble those of ''Hypacrosaurus''. With only one partial skeleton known, and no skull, the genus has been considered

dubious Doubt is a mental state in which the mind remains suspended between two or more contradictory propositions, unable to be certain of any of them. Doubt on an emotional level is indecision between belief and disbelief. It may involve uncertainty ...

or a possible lambeosaurine of uncertain placement. A newer study published in 2011 suggests that ''Barsboldia'' is actually a valid saurolophine. The following

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...

was recovered in the 2011 phylogenetic analysis of

Hadrosauroidea Hadrosauroidea is a clade or superfamily of ornithischian dinosaurs that includes the "duck-billed" dinosaurs, or hadrosaurids, and all dinosaurs more closely related to them than to ''Iguanodon''. Their remains have been recovered in Asia, Europ ...

by Prieto-Márquez (the relationships within Lambeosaurinae and between basal hadrosauroids aren't shown). Barsboldia vertebrae

Paleobiology

As a hadrosaurid, ''Barsboldia'' would have been a

bipedal Bipedalism is a form of terrestrial locomotion where an organism moves by means of its two rear limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' 'double' ...

quadruped Quadrupedalism is a form of locomotion where four limbs are used to bear weight and move around. An animal or machine that usually maintains a four-legged posture and moves using all four limbs is said to be a quadruped (from Latin ''quattuor' ...

al herbivore, eating

plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclu ...

s with a sophisticated skull that permitted a grinding motion analogous to

chewing Chewing or mastication is the process by which food is crushed and ground by teeth. It is the first step of digestion, and it increases the surface area of foods to allow a more efficient break down by enzymes. During the mastication process, th ...

, and was furnished with hundreds of continually-replaced

teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, t ...

. If it was a lambeosaurine, it would have had a hollow crest formed out of expanded skull bones containing the nasal passages, with a function relating to identification by sight and sound.

References

{{Taxonbar, from1=Q18606529, from2=Q264109 Saurolophines Late Cretaceous dinosaurs of Asia Maastrichtian life Nemegt fauna Fossil taxa described in 1981 Taxa named by Teresa Maryańska Taxa named by Halszka Osmólska Ornithischian genera

Discovery

Description

Phylogeny

Paleobiology

See also

References