''Barbosania'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

of crestless

targaryendraconia Targaryendraconia is an extinct clade of lanceodontian pterosaurs that lived from the Early to Late Cretaceous period in Europe, North America, South America, and Australia. Classification Below is a cladogram following a topology by Pêga ...

pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...

from the

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...

Romualdo Formation The Romualdo Formation is a geologic Konservat-Lagerstätte in northeastern Brazil's Araripe Basin where the states of Pernambuco, Piauí and Ceará come together. The geological formation, previously designated as the Romualdo Member of the Sa ...

of the

Santana Group The Santana Group is a geologic group, formerly included as the middle part of the Araripe Group, in the Araripe Basin of northeastern Brazil. The group comprises the Crato, Ipubi and Romualdo Formations and is dated to the Aptian to Albian stage ...

of northeastern

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

, dating to the

Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...

Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 M ...

Discovery and naming

''Barbosania'' was named and described by Ross A. Elgin and

Eberhard Frey Eberhard is an old Germanic name meaning the strength or courage of a wild boar. People First name *Eberhard of Friuli (815–866), Duke and key figure in the Carolingian Empire *Eberhard of Béthune (died 1212), Flemish grammarian *Eberhard I, Du ...

2011 File:2011 Events Collage.png, From top left, clockwise: a protester partaking in Occupy Wall Street heralds the beginning of the Occupy movement; protests against Libyan dictator Muammar Gaddafi, who was killed that October; a young man celebrate ...

and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''Barbosania gracilirostris''. The generic name honours Professor

Miguel Barbosa Miguel Ângelo Freitas Barbosa, known as Miguel Barbosa (born 16 October 1996), is a Portuguese football defender who plays for FK Sloboda Užice in the Serbian First League. Club career Born in Funchal, the capital of Madeira islands, a Portu ...

of the Portuguese Museu de História Natural de Sintra in whose collection the

type specimen In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to a ...

is present. The

specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...

is derived from

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

''rostrum'', "snout", and ''gracilis'', "slender", in reference to the slender form of the anterior skull. The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

, MNHS/00/85, was originally acquired for Barbosa's personal collection, the basis of the new

Sintra Sintra (, ) is a town and municipality in the Greater Lisbon region of Portugal, located on the Portuguese Riviera. The population of the municipality in 2011 was 377,835, in an area of . Sintra is one of the most urbanized and densely populated ...

museum, from Brazilian fossil dealers. Its provenance is probably the Serra da Mãozinha (in the original article was Sierra de Maõsina, but this name is wrong), implying an

Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Ma to 100.5 Ma. Geology Pro ...

late Albian age, about a hundred million years old. It consists of an almost complete

skeleton A skeleton is the structural frame that supports the body of an animal. There are several types of skeletons, including the exoskeleton, which is the stable outer shell of an organism, the endoskeleton, which forms the support structure inside ...

including the

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...

, that is partially articulated and uncompressed. The lower legs and most of the neck are missing. The specimen was prepared in the German

Staatliches Museum für Naturkunde Karlsruhe The State Museum of Natural History Karlsruhe (german: link=no, Staatliches Museum für Naturkunde Karlsruhe), abbreviated SMNK, is one of the two state of Baden-Württemberg's natural history museums. Together with the State Museum of Natural H ...

. It is probably of a subadult individual, though the closure of the respective sutures provides somewhat contradictory information, the fusion of some elements indicating it was full-grown. Several specimens previously referred to ''

Brasileodactylus ''Brasileodactylus'' a genus of pterosaur from the Aptian-age (Early Cretaceous period) lower Santana formation of Chapada do Araripe, Ceará, Brazil. The genus was named by paleontologist Alexander Wilhelm Armin Kellner in 1984. The genus name ...

'' may instead be individuals of ''Barbosania''.

Description

''Barbosania'' was a medium-sized

pterodactyloid Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional ...

with a skull length of 392 millimetres. Its body was 209.5 millimetres long. The skull is elongated with a slight upwards bend in the snout. Its lack of a rostral and dentary median sagittal crest allows a distinction from other related pterosaurs. Also a parietal crest on top of the skull is absent. The authors considered it unlikely that this morphology was age-related and rejected the explanation of such traits by

sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...

unless there was specific evidence to prove this. The describers established a single

autapomorphy In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to t ...

, unique trait: the possession of thirteen dorsal vertebrae instead of the normal twelve. The first four tooth pairs are extremely long, forming a rosette to catch slippery prey like fish or squids. This rosette is however not laterally expanded, which was reflected in the specific name. There are at least twenty-four teeth in the upper jaw and twenty in the lower jaw for a total of eighty-eight.

Classification

''Barbosania'' was assigned to the

Ornithocheiroidea Ornithocheiroidea (or ornithocheiroids) is a group of pterosaurs within the extinct suborder Pterodactyloidea. They were typically large pterosaurs that lived from the Early to Late Cretaceous periods (Valanginian to Maastrichtian stages), with f ...

''sensu'' Unwin and more precisely to the

Ornithocheiridae Ornithocheiridae (or ornithocheirids, meaning "bird hands") is a group of pterosaurs within the suborder Pterodactyloidea. These pterosaurs were among the last to possess teeth. Members that belong to this group lived from the Early to Late Cr ...

. In 2019, Pêgas ''et al.'' assigned ''Barbosania'' to the clade

Targaryendraconia Targaryendraconia is an extinct clade of lanceodontian pterosaurs that lived from the Early to Late Cretaceous period in Europe, North America, South America, and Australia. Classification Below is a cladogram following a topology by Pêga ...

, more specifically to the family Targaryendraconidae, as shown below:Rodrigo V. Pêgas, Borja Holgado & Maria Eduarda C. Leal (2019) On ''Targaryendraco wiedenrothi'' gen. nov. (Pterodactyloidea, Pteranodontoidea, Lanceodontia) and recognition of a new cosmopolitan lineage of Cretaceous toothed pterodactyloids, Historical Biology,

References

{{Portal bar, Paleontology, Cretaceous, Brazil Pteranodontoids Early Cretaceous pterosaurs of South America Albian life Aptian life Cretaceous Brazil Fossils of Brazil Romualdo Formation Fossil taxa described in 2011

Discovery and naming

Description

Classification

See also

References