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The Hemiscylliidae are a
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of
shark Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachimo ...
s in the order
Orectolobiformes Carpet sharks are sharks classified in the order Orectolobiformes . Sometimes the common name "carpet shark" (named so because many species resemble ornately patterned carpets) is used interchangeably with "wobbegong", which is the common name ...
, commonly known as longtail carpet sharks and sometimes as bamboo sharks. They are found in shallow waters of the tropical
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
. They are relatively small sharks, with the largest species reaching no more than in adult body length. They have elongated, cylindrical bodies, with short
barbel Barbel may refer to: *Barbel (anatomy), a whisker-like organ near the mouth found in some fish (notably catfish, loaches and cyprinids) and turtles *Barbel (fish), a common name for certain species of fish **''Barbus barbus'', a species of cyprinid ...
s and large
spiracle Spiracle or spiraculum may refer to: * Spiracle (arthropods), opening in the exoskeletons of some arthropods * Spiracle (vertebrates), openings on the surface of some vertebrates * Spiraculum, a genus of land snails in family Cyclophoridae Cycl ...
s. As their common name suggests, they have unusually long tails, which exceed the length of the rest of their bodies. They are sluggish fish, feeding on bottom-dwelling invertebrates and smaller fish.


Genera and species


''Chiloscyllium''

This genus is distinguished by a relatively long snout with subterminal
nostril A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called turbi ...
s. The eyes and supraorbital ridges are hardly elevated. The mouth is closer to the eyes than to the tip of the snout, with lower labial folds usually connected across the chin by a flap of skin. The
pectoral Pectoral may refer to: * The chest region and anything relating to it. * Pectoral cross, a cross worn on the chest * a decorative, usually jeweled version of a gorget * Pectoral (Ancient Egypt), a type of jewelry worn in ancient Egypt * Pectorali ...
and
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods. Structure and function Structure In actinopterygians, the pelvic fin consists of two en ...
s are thin and not very muscular. No black hood on the head or large black spot on the side is present (though juveniles often are strongly marked with dark spots/bars). * ''
Chiloscyllium arabicum The Arabian carpetshark (''Chiloscyllium arabicum'') is a species of carpet shark in the family Hemiscylliidae, inhabiting coral reefs and other shallow coastal habitats from the Persian Gulf to India. Reaching long, this shark is characterized b ...
'' Gubanov, 1980 (Arabian carpetshark) * ''
Chiloscyllium burmensis The Burmese bamboo shark, ''Chiloscyllium burmensis'', is an extremely rare bamboo shark in the family Hemiscylliidae. The first specimen was caught 1963 off the coast from Rangoon in Burma in a depth of 29 – 33 m. This holotype is an adul ...
'' Dingerkus & DeFino, 1983 (Burmese bamboo shark) * ''
Chiloscyllium griseum The grey bamboo shark, ''Chiloscyllium griseum'', is a species of carpet shark in the family Hemiscylliidae, found in the Indo-West Pacific Oceans from the Arabian Sea to Pakistan, India, Malaysia, Thailand, Indonesia, China, Japan, the Philippin ...
'' J. P. Müller &
Henle Henle can refer to: * Friedrich Gustav Jakob Henle, a German physician, pathologist and anatomist (1809–1885) ** Loop of Henle in the kidney, named after Henle *Fritz Henle, a photographer, known as "Mr. Rollei" for his use of the 2.25" square for ...
, 1838
(grey bamboo shark) * ''
Chiloscyllium hasselti Hasselt's bamboo shark (''Chiloscyllium hasseltii'') is a bamboo shark in the family Hemiscylliidae found around Thailand, Malaysia and Indonesia, between latitudes 23° N and 10° S, and longitude 91° E and 133° E; residing inshore. Its le ...
''
Bleeker Bleeker is a Dutch occupational surname. Bleeker is an old spelling of ''(linnen)bleker'' ("linen bleacher").Chiloscyllium indicum The slender bamboo shark, ''Chiloscyllium indicum'', is a bamboo shark in the family Hemiscylliidae found in the Indo-West Pacific Oceans between latitudes 40° N and 10° S, and longitude 65° E and 160° E. It is harmless to humans. Descr ...
'' ( J. F. Gmelin, 1789) (slender bamboo shark) * '' Chiloscyllium plagiosum'' (Anonymous, referred to
Bennett Bennett may refer to: People *Bennett (name), including a list of people with the surname and given name Places Canada * Bennett, Alberta *Bennett, British Columbia * Bennett Lake, in the British Columbia and Yukon Territory **Bennett Range **Benn ...
, 1830)
(white-spotted bamboo shark) * '' Chiloscyllium punctatum'' J. P. Müller &
Henle Henle can refer to: * Friedrich Gustav Jakob Henle, a German physician, pathologist and anatomist (1809–1885) ** Loop of Henle in the kidney, named after Henle *Fritz Henle, a photographer, known as "Mr. Rollei" for his use of the 2.25" square for ...
, 1838
(brown-banded bamboo shark)


''Hemiscyllium''

This genus is confined to tropical waters of
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
,
Papua New Guinea Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilong Papua Niugini; ho, Independen Stet bilong Papua Niu Gini), is a country i ...
, and
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
, but an individual from this genus, possibly representing an undescribed species, has been photographed at the
Seychelles Seychelles (, ; ), officially the Republic of Seychelles (french: link=no, République des Seychelles; Creole: ''La Repiblik Sesel''), is an archipelagic state consisting of 115 islands in the Indian Ocean. Its capital and largest city, V ...
. They have short snouts with the
nostril A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called turbi ...
s placed almost at the tip, and well-elevated eyes and supraorbital ridges. The mouth is closer to the tip of the snout than the eyes, and lacks the connecting dermal fold across the chin. The pectoral and pelvic fins are thick and heavily muscular. Either a black hood on the head or a large black spot on the sides of the body is present. Nine recognized species are in this genus: * '' Hemiscyllium freycineti'' ( Quoy &
Gaimard Joseph Paul Gaimard (31 January 1793 – 10 December 1858) was a French naval surgeon and naturalist. Biography Gaimard was born at Saint-Zacharie on January 31, 1793. He studied medicine at the naval medical school in Toulon, subsequent ...
, 1824)
(Indonesian speckled carpetshark) * '' Hemiscyllium galei'' G. R. Allen & Erdmann, 2008 (Cenderwasih epaulette shark) * '' Hemiscyllium hallstromi'' Whitley, 1967 (Papuan epaulette shark) * '' Hemiscyllium halmahera'' G. R. Allen, Erdmann &
Dudgeon Dudgeon is a surname. Notable people with the surname include: * Cecil Dudgeon, politician * Gus Dudgeon, record producer * James Dudgeon, footballer * Joe Dudgeon, English footballer * John Dudgeon, physician * Neil Dudgeon, actor * Patrick Dud ...
, 2013
(Halmahera epaulette shark) * '' Hemiscyllium henryi'' G. R. Allen & Erdmann, 2008 (Henry's epaulette shark) * '' Hemiscyllium michaeli'' G. R. Allen &
Dudgeon Dudgeon is a surname. Notable people with the surname include: * Cecil Dudgeon, politician * Gus Dudgeon, record producer * James Dudgeon, footballer * Joe Dudgeon, English footballer * John Dudgeon, physician * Neil Dudgeon, actor * Patrick Dud ...
, 2010
(Milne Bay epaulette shark) * '' Hemiscyllium ocellatum'' ( Bonnaterre, 1788) (epaulette shark) * ''
Hemiscyllium strahani The hooded carpetshark (''Hemiscyllium strahani'') is a bamboo shark in the family Hemiscylliidae found around Papua New Guinea, between latitudes 5° S and 10° S, and longitude 144° E and 153° E. Its length is up to 75 cm. Like other ...
'' Whitley, 1967 (hooded carpetshark) * ''
Hemiscyllium trispeculare The speckled carpetshark, ''Hemiscyllium trispeculare'', is a bamboo shark in the family Hemiscylliidae found around north and west Australia between latitudes 8° S and 22° S, and longitude 114° E and 152° E. Its length is up to 79 c ...
'' J. Richardson, 1843 (speckled carpetshark) * '' Hemiscyllium sp.'' Not yet described (Seychelles carpetshark)


Fossil taxa

* †'' Acanthoscyllium sahelalmae'' (Pictet & Humbert, 1866) * †'' Adnetoscyllium angloparisensis'' (Guinot et al, 2013) * †'' Almascyllium cheikeliasi'' (Signeaux, 1949) * †'' Chiloscyllium broenirnani'' Casier, 1958 * †'' Hemiscyllium bruxelliensis'' Herman, 1977 * †'' Mesiteia daimeriesi'' (Herman, 1973) * †'' Pseudospinax heterodon'' Underwood & Mitchell, 1999


Captivity

Hemiscylliid sharks are sometimes kept in home aquaria. Species from this family are ideal aquarium sharks because their natural habitats are tidepools, coral beds, and around boulders. This predisposition towards relatively confined spaces helps them adapt better to home aquaria compared to other species. Their generally small size for sharks, and their preference for water temperatures comparable to those enjoyed by other common aquarium fish, have also endeared them to marine aquarists. Multiple species of hemiscylliids have been successfully induced to breed in captivity. Full-sized adult epaulette sharks are most successfully housed in tanks at or exceeding , while adult bamboo sharks require more space and are known to do well in aquaria. Hemiscyliids in captivity are provided artificial caves in which to hide. However, unstable tank decor has been known to cause fatal injuries when the structure is disturbed by the sharks' digging behavior.


Parthenogenesis

The British press on February 10, 2016 reported that a bamboo shark at Great Yarmouth’s Sea Life Centre was pregnant with two fertilized eggs. It is known that the shark has not come into contact with any other bamboo sharks since 2013. Although parthenogenesis is observed in a small number of species, this is such a rare occurrence in this species that it became a news story.Martin, Sean (2016-02-10
A miracle? Female bamboo shark set for VIRGIN BIRTH at British zoo
Express.co.uk. Retrieved on 2017-03-08.


References

{{Taxonbar, from=Q805999 Orectolobiformes Taxa named by Theodore Gill Shark families Extant Late Jurassic first appearances