In the

APG IV system The APG IV system of flowering plant classification is the fourth version of a modern, mostly molecular-based, system of plant taxonomy for flowering plants (angiosperms) being developed by the Angiosperm Phylogeny Group (APG). It was publishe ...

(2016) for the classification of

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

s, the name asterids denotes a

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...

group). Asterids is the largest group of

s, with more than 80,000 species, about a third of the total flowering plant species. Well-known plants in this clade include the

common daisy ''Bellis perennis'' (), the daisy, is a European species of the family Asteraceae, often considered the archetypal species of the name daisy. To distinguish this species from other plants known as daisies, it is sometimes qualified as common dai ...

forget-me-not ''Myosotis'' ( ) is a genus of flowering plants in the family Boraginaceae. The name comes from the Ancient Greek "mouse's ear", which the foliage is thought to resemble. In the northern hemisphere they are colloquially known as forget-me-nots ...

nightshades The Solanaceae , or nightshades, are a family of flowering plants that ranges from annual and perennial herbs to vines, lianas, epiphytes, shrubs, and trees, and includes a number of agricultural crops, medicinal plants, spices, weeds, and orna ...

(including

potato The potato is a starchy food, a tuber of the plant ''Solanum tuberosum'' and is a root vegetable native to the Americas. The plant is a perennial in the nightshade family Solanaceae. Wild potato species can be found from the southern Unit ...

es,

eggplant Eggplant ( US, Canada), aubergine ( UK, Ireland) or brinjal (Indian subcontinent, Singapore, Malaysia, South Africa) is a plant species in the nightshade family Solanaceae. ''Solanum melongena'' is grown worldwide for its edible fruit. Mos ...

tomatoes The tomato is the edible Berry (botany), berry of the plant ''Solanum lycopersicum'', commonly known as the tomato plant. The species originated in western South America, Mexico, and Central America. The Mexican Nahuatl word gave rise to th ...

chili peppers Chili peppers (also chile, chile pepper, chilli pepper, or chilli), from Nahuatl '' chīlli'' (), are varieties of the berry-fruit of plants from the genus ''Capsicum'', which are members of the nightshade family Solanaceae, cultivated for ...

and

tobacco Tobacco is the common name of several plants in the genus '' Nicotiana'' of the family Solanaceae, and the general term for any product prepared from the cured leaves of these plants. More than 70 species of tobacco are known, but the ...

), the

common sunflower The common sunflower (''Helianthus annuus'') is a large annual forb of the genus ''Helianthus'' grown as a crop for its edible oily seeds. Apart from cooking oil production, it is also used as livestock forage (as a meal or a silage plant), as ...

petunias ''Petunia'' is genus of 20 species of flowering plants of South American origin. The popular flower of the same name derived its epithet from the French, which took the word ''petun'', meaning "tobacco," from a Tupi–Guarani language. A tende ...

, yacon,

morning glory Morning glory (also written as morning-glory) is the common name for over 1,000 species of flowering plants in the family Convolvulaceae, whose current taxonomy and systematics are in flux. Morning glory species belong to many genera, some of ...

sweet potato The sweet potato or sweetpotato (''Ipomoea batatas'') is a dicotyledonous plant that belongs to the Convolvulus, bindweed or morning glory family (biology), family, Convolvulaceae. Its large, starchy, sweet-tasting tuberous roots are used as a r ...

coffee Coffee is a drink prepared from roasted coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulant, stimulating effect on humans, primarily due to its caffeine content. It is the most popular hot drink in the world. S ...

lavender ''Lavandula'' (common name lavender) is a genus of 47 known species of flowering plants in the mint family, Lamiaceae. It is native to the Old World and is found in Cape Verde and the Canary Islands, and from Europe across to northern and easte ...

lilac ''Syringa'' is a genus of 12 currently recognized species of flowering plant, flowering woody plants in the olive family or Oleaceae called lilacs. These lilacs are native to woodland and scrub from southeastern Europe to eastern Asia, and wid ...

olive The olive, botanical name ''Olea europaea'', meaning 'European olive' in Latin, is a species of small tree or shrub in the family Oleaceae, found traditionally in the Mediterranean Basin. When in shrub form, it is known as ''Olea europaea'' ...

jasmine Jasmine ( taxonomic name: ''Jasminum''; , ) is a genus of shrubs and vines in the olive family (Oleaceae). It contains around 200 species native to tropical and warm temperate regions of Eurasia, Africa, and Oceania. Jasmines are widely cultiva ...

honeysuckle Honeysuckles are arching shrubs or twining vines in the genus ''Lonicera'' () of the family Caprifoliaceae, native to northern latitudes in North America and Eurasia. Approximately 180 species of honeysuckle have been identified in both contin ...

ash tree ''Fraxinus'' (), commonly called ash, is a genus of flowering plants in the olive and lilac family, Oleaceae. It contains 45–65 species of usually medium to large trees, mostly deciduous, though a number of subtropical species are evergree ...

teak Teak (''Tectona grandis'') is a tropical hardwood tree species in the family Lamiaceae. It is a large, deciduous tree that occurs in mixed hardwood forests. ''Tectona grandis'' has small, fragrant white flowers arranged in dense clusters (panicl ...

snapdragon ''Antirrhinum'' is a genus of plants commonly known as dragon flowers, snapdragons and dog flower because of the flowers' fancied resemblance to the face of a dragon that opens and closes its mouth when laterally squeezed. They are native to r ...

sesame Sesame ( or ; ''Sesamum indicum'') is a flowering plant in the genus ''Sesamum'', also called benne. Numerous wild relatives occur in Africa and a smaller number in India. It is widely naturalized in tropical regions around the world and is cu ...

psyllium Psyllium , or ispaghula , is the common name used for several members of the plant genus ''Plantago'' whose seeds are used commercially for the production of mucilage. Psyllium is mainly used as a dietary fiber to relieve symptoms of both constip ...

garden sage ''Salvia officinalis'', the common sage or just sage, is a perennial, evergreen subshrub, with woody stems, grayish leaves, and blue to purplish flowers. It is a member of the mint family Lamiaceae and native to the Mediterranean region, thoug ...

, table herbs such as

mint MiNT is Now TOS (MiNT) is a free software alternative operating system kernel for the Atari ST system and its successors. It is a multi-tasking alternative to TOS and MagiC. Together with the free system components fVDI device drivers, XaAES g ...

basil Basil (, ; ''Ocimum basilicum'' , also called great basil, is a culinary herb of the family Lamiaceae (mints). It is a tender plant, and is used in cuisines worldwide. In Western cuisine, the generic term "basil" refers to the variety also kno ...

, and

rosemary ''Salvia rosmarinus'' (), commonly known as rosemary, is a shrub with fragrant, evergreen, needle-like leaves and white, pink, purple, or blue flowers, native plant, native to the Mediterranean Region, Mediterranean region. Until 2017, it was kn ...

, and rainforest trees such as

Brazil nut The Brazil nut (''Bertholletia excelsa'') is a South American tree in the family Lecythidaceae, and it is also the name of the tree's commercially harvested edible seeds. It is one of the largest and longest-lived trees in the Amazon rainforest. ...

. Most of the taxa belonging to this clade had been referred to as

Asteridae ''Asteridae'' is an obsolete botanical name at the rank of subclass. Composition of the subclass has also varied; however, by definition it always includes the family Asteraceae (Compositae). In the modern APG IV system of classification, asterid ...

in the

Cronquist system The Cronquist system is a taxonomic classification system of flowering plants. It was developed by Arthur Cronquist in a series of monographs and texts, including ''The Evolution and Classification of Flowering Plants'' (1968; 2nd edition, 1988) a ...

(1981) and as

Sympetalae Sympetaly (fused petals) is a flower characteristic that historically was used to classify a grouping of plants termed Sympetalae, but this term has been abandoned in newer molecular based classifications, although the grouping has similarity to ...

in earlier systems. The name asterids (not necessarily capitalised) resembles the earlier

botanical name A botanical name is a formal scientific name conforming to the '' International Code of Nomenclature for algae, fungi, and plants'' (ICN) and, if it concerns a plant cultigen, the additional cultivar or Group epithets must conform to the ''Intern ...

but is intended to be the name of a clade rather than a formal ranked name, in the sense of the ''

ICBN The ''International Code of Nomenclature for algae, fungi, and plants'' (ICN) is the set of rules and recommendations dealing with the formal botanical names that are given to plants, fungi and a few other groups of organisms, all those "trad ...

''.

History

Genetic analysis carried out after

APG II The APG II system (Angiosperm Phylogeny Group II system) of plant classification is the second, now obsolete, version of a modern, mostly molecular-based, system of plant taxonomy that was published in April 2003 by the Angiosperm Phylogeny Gr ...

maintains that the sister to all other asterids are the

Cornales The Cornales are an order of flowering plants, early diverging among the asterids, containing about 600 species. Plants within the Cornales usually have four-parted flowers, drupaceous fruits, and inferior to half-inferior gynoecia topped with d ...

. A second order that split from the base of the asterids are the

Ericales The Ericales are a large and diverse order of dicotyledons. Species in this order have considerable commercial importance including for Camellia sinensis, tea, persimmon, blueberry, kiwifruit, Brazil nuts, Argania, argan, and azalea. The order i ...

. The remaining orders cluster into two clades, the lamiids and the campanulids. The structure of both of these clades has changed in

APG III APG is an abbreviation with several different meanings: * Aberdeen Proving Ground, a United States Army installation in Aberdeen, Maryland, also ** Phillips Army Airfield, the airfield of the above, from its IATA airport code * Aboriginal Provisiona ...

. In the APG III system, the following clades were renamed: : euasterids I → lamiids : euasterids II → campanulids

Phylogeny

The phylogenetic tree presented hereinafter has been proposed by the APG IV project.

Subdivision

;lamiids The lamiid subclade consists of about 40,000 species and account for about 15% of angiosperm diversity, characterized in general by superior

ovaries The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. T ...

and corollas with any fusion of the petals () occurring late in the process of development. The major part of lamiid diversity occurs in the group of five orders from Boraginales to Solanales, referred to informally as "core lamiids" (sometimes called Laminae), although Vahliales consists of the single small genus

Vahlia :''The later homonym Vahlia Dahl is now known as Dombeya.'' ''Vahlia'' is a genus of herbs and subshrubs that grow in Africa and the Indian subcontinent. There are at least five species. The genus is placed alone in family ''Vahliaceae''. This ...

. The remainder of the lamiids are referred to as "basal lamiids", in which

Garryales The Garryales are a small order of dicotyledons, including only two families and three genera. Description Garryales are woody plants that are either hairless or have very fine hairs. Members of the family Garryaceae are evergreen, whereas thos ...

is the

sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...

to the core lamiids. It has been suggested that the core lamiids radiated from an ancestral line of tropical trees in which the flowers were inconspicuous and the fruit large,

drupaceous In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or ''pyrena'') of hardened endocarp with a seed (''kernel'') ...

and often single-seeded.

References

Bibliography

External links

*
Asterids
in Stevens, P. F. (2001 onwards)
Angiosperm Phylogeny Website
Version 7, May 2006. {{Taxonbar, from=Q747502 Plant unranked clades