The pied monarch (''Arses kaupi'') is a species of

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...

in the monarch-flycatcher family,

Monarchidae The monarchs (family Monarchidae) comprise a family of over 100 passerine birds which includes shrikebills, paradise flycatchers, and magpie-larks. Monarchids are small insectivorous songbirds with long tails. They inhabit forest or woodland a ...

. It is

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

to coastal

Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...

in Australia.

Taxonomy and systematics

The pied monarch was described by

John Gould John Gould (; 14 September 1804 – 3 February 1881) was an English ornithologist. He published a number of monographs on birds, illustrated by plates produced by his wife, Elizabeth Gould, and several other artists, including Edward Lear, ...

in 1851, who deliberated on placing it in a genus by itself on account of its feet and eye ring. The nest and eggs were undescribed until collected by Robert Hislop on 3 December 1894 near

Bloomfield River The Bloomfield River is a river located in the Wet Tropics of Far North Queensland, Australia, noted for its Bloomfield River cod fish species, found only in the river. Course and features The river rises in the Great Dividing Range below Zi ...

. The pied monarch is closely related to and forms a superspecies with the two other species of

monarch flycatcher The monarchs (family Monarchidae) comprise a family of over 100 passerine birds which includes shrikebills, paradise flycatchers, and magpie-larks. Monarchids are small insectivorous songbirds with long tails. They inhabit forest or woodland a ...

in the genus ''

Arses Arses ( peo, *R̥šā; grc-gre, Ἀρσής), also known by his regnal name Artaxerxes IV (; peo, 𐎠𐎼𐎫𐎧𐏁𐏂𐎠 ; grc-gre, Ἀρταξέρξης), was the twelfth Achaemenid King of Kings from 338 to 336 BC. Arses ascended ...

''. Two subspecies are recognised, however they two intergrade where their ranges meet at Mossman, and they could be treated as a monotypic species. The monarch flycatchers are classified either as a subfamily Monarchinae, together with the

fantail Fantails are small insectivorous songbirds of the genus ''Rhipidura'' in the family Rhipiduridae, native to Australasia, Southeast Asia and the Indian subcontinent. Most of the species are about long, specialist aerial feeders, and named as "f ...

s as part of the drongo family Dicruridae, or as a family Monarchidae in its own right. Molecular research in the late 1980s and early 1990s revealed the monarchs belong to a large group of mainly Australasian birds known as the

Corvida The "Corvida" were one of two "parvorders" contained within the suborder Passeri, as proposed in the Sibley-Ahlquist taxonomy, the other being Passerida. Standard taxonomic practice would place them at the rank of infraorder. More recent resea ...

parvorder Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and ...

comprising many tropical and Australian passerines. More recently, the grouping has been refined somewhat as the monarchs have been classified in a 'Core corvine' group with the crows and ravens, shrikes, birds of paradise, fantails, drongos and mudnest builders. Alternate names include the Australian pied flycatcher, Australian pied monarch, banded monarch, pied monarch-flycatcher, black-breasted flycatcher, Kaup's flycatcher and pied flycatcher.

Subspecies

Two

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

are recognized: * ''A. k. terraereginae'' - Campbell, AJ, 1895: Found on eastern

Cape York Peninsula Cape York Peninsula is a large peninsula located in Far North Queensland, Australia. It is the largest unspoiled wilderness in northern Australia.Mittermeier, R.E. et al. (2002). Wilderness: Earth’s last wild places. Mexico City: Agrupación ...

(north-eastern Australia) * ''A. k. kaupi'' -

Gould Gould may refer to: People * Gould (name), a surname Places United States * Gould, Arkansas, a city * Gould, Colorado, an unincorporated community * Gould, Ohio, an unincorporated community * Gould, Oklahoma, a town * Gould, West Virginia, a ...

, 1851: Found on south-eastern Cape York Peninsula (north-eastern Australia)

Description

The pied monarch is in length and weighs around . The

plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...

sexually dimorphic Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...

. The upperparts and head of the male are black, as is the tail, and the wings are brownish-black with white scapulars (which shows as a white crescent across the back when folded). The collar, which is erectable, is white, and it joins through the neck with a white throat. The breast-band is black, and the belly and underside are white. In the female the white on the throat and collar is less distinct and covers less area, and the collar is incomplete around the neck. The eye is black, and is surrounded by a blue coloured eye-ring, which is less distinct in the female. The

bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...

is blue-grey, and the legs are black. Immature birds look like females but with duller plumage, no blue in the eye-rings, and a horn-coloured bill. Like other members of the genus it has long hind-toes and claws, and in general the size and shape of the feet resemble those of the

Australasian treecreeper There are seven species of Australasian treecreeper in the passerine bird family Climacteridae. They are medium-small, mostly brown birds with patterning on their underparts, and all are endemic to Australia-New Guinea. They resemble, but are no ...

s, and the feet are used for feeding in the same manner on the trunks of trees.

Distribution and habitat

The pied monarch is found in tropical forest edge habitats and secondary growth in coastal north eastern

, from

Cooktown Cooktown is a coastal town and locality in the Shire of Cook, Queensland, Australia. Cooktown is at the mouth of the Endeavour River, on Cape York Peninsula in Far North Queensland where James Cook beached his ship, the Endeavour, for repairs ...

to Ingham. It ranges from sea level up to . It also occurs in palm-vine scrub, gallery forest and along rivers. The species is mostly non- migratory, but some birds disperse to the ''

Eucalyptus ''Eucalyptus'' () is a genus of over seven hundred species of flowering trees, shrubs or mallees in the myrtle family, Myrtaceae. Along with several other genera in the tribe Eucalypteae, including '' Corymbia'', they are commonly known as euca ...

'' woodland in the

Atherton Tableland The Atherton Tableland is a fertile plateau which is part of the Great Dividing Range in Queensland, Australia. The principal river flowing across the plateau is the Barron River. It was dammed to form an irrigation reservoir named Lake Tina ...

during the winter. The species has a tiny global range, and is described as uncommon and occurring at low densities throughout its range. However much of its range is protected within

national park A national park is a nature park, natural park in use for conservation (ethic), conservation purposes, created and protected by national governments. Often it is a reserve of natural, semi-natural, or developed land that a sovereign state dec ...

s or

World Heritage Site A World Heritage Site is a landmark or area with legal protection by an international convention administered by the United Nations Educational, Scientific and Cultural Organization (UNESCO). World Heritage Sites are designated by UNESCO for h ...

s, and its habitat is thought to be secure. The species is considered to be safe at the moment, and is listed as

Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...

by the

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

Behaviour

The pied monarch is insectivorous, with

beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...

s (

Coleoptera Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...

) and moths and butterflies (

Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...

) being recorded in its diet. It is usually seen as singles or pairs and small groups (of three to five birds, which may be family groups). They join

mixed-species foraging flock A mixed-species feeding flock, also termed a mixed-species foraging flock, mixed hunting party or informally bird wave, is a flock of usually insectivorous birds of different species that join each other and move together while foraging. These ar ...

s with other

s, whistlers and shrikethrushes. Within the forest they usually feed at the mid level, and rarely close to the ground. One foraging method typical to the genus is to climb up trunks and the larger branches in the manner of a treecreeper (''

Climacteris ''Climacteris'' is a genus of bird in the family Climacteridae. These birds and the other members of the family, genus '' Cormobates'', are similar to Northern Hemisphere creepers, Certhiidae, in climbing helically up tree trunks looking for ins ...

'') and probing the bark and lichens, but they also catch prey from the air.

Breeding

Breeding season is October to January with one brood raised. The nest is a shallow cup made of vines and sticks, woven together with

spider web A spider web, spiderweb, spider's web, or cobweb (from the archaic word '' coppe'', meaning "spider") is a structure created by a spider out of proteinaceous spider silk extruded from its spinnerets, generally meant to catch its prey. Spid ...

s and shredded plant material, and decorated with

lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.John Gould John Gould (; 14 September 1804 – 3 February 1881) was an English ornithologist. He published a number of monographs on birds, illustrated by plates produced by his wife, Elizabeth Gould, and several other artists, including Edward Lear, ...

References

{{Taxonbar, from=Q777968

pied monarch The pied monarch (''Arses kaupi'') is a species of bird in the monarch-flycatcher family, Monarchidae. It is endemic to coastal Queensland in Australia. Taxonomy and systematics The pied monarch was described by John Gould in 1851, who delibe ...

Birds of Queensland Endemic birds of Australia

Taxonomy articles created by Polbot