Arbutus Pungens
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''Arbutus'' is a genus of 12 accepted speciesAct. Bot. Mex no.99 Pátzcuaro abr. 2012.
''Arbutus bicolor''
/ref> of flowering plants in the family Ericaceae, native to warm temperate regions of the Mediterranean, western Europe, the
Canary Islands The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Morocc ...
and North America. The name ''Arbutus'' was taken from Latin, where it referred to '' Arbutus unedo''.


Description

''Arbutus'' are small trees or shrubs with red flaking bark and edible red berries.Mabberley, D. J. 1997. ''The plant book: A portable dictionary of the vascular plants''. Cambridge University Press, Cambridge. Fruit development is delayed for about five months after pollination, so that flowers appear while the previous year's fruit are ripening. Peak flowering for the genus is in April with peak fruiting in October.


History

The smooth wood of the tree is mentioned by Theophrastus in his ''Enquiry into Plants'' (''Historia Plantarum'') as formerly being used to make weaving spindles. An article on Arbutus tree cultivation in al-Andalus is brought down in Ibn al-'Awwam's 12th-century agricultural work, ''Book on Agriculture''.


Common names

Members of the genus are called ''madrones'' or ''madronas'' in the United States, from the Spanish name ''madroño'' ( strawberry tree). On the south coast of British Columbia, Canada, where the species is common, ''arbutus'' is commonly used or, rarely and locally, "tick tree". All refer to the same species, ''
Arbutus menziesii ''Arbutus menziesii'' or Pacific madrone (commonly madrone or madrona in the United States and arbutus in Canada), is a species of broadleaf evergreen tree in the family Ericaceae, native to the western coastal areas of North America, from Bri ...
'', native to the Pacific Northwest and Northern and Central California regions. It is Canada's only native broadleaved evergreen tree. Some species in the genera '' Epigaea'', '' Arctostaphylos'' and '' Gaultheria'' were formerly classified in ''Arbutus''. As a result of its past classification, '' Epigaea repens'' (mayflower) has an alternative common name of "trailing arbutus".


Systematics

A study published in 2001 which analyzed ribosomal DNA from ''Arbutus'' and related genera suggests that ''Arbutus'' is
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
and the
Mediterranean Basin In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and w ...
species of ''Arbutus'' are more closely related to '' Arctostaphylos'', '' Arctous'', '' Comarostaphylis'', '' Ornithostaphylos'' and '' Xylococcus'' than to the western North American species of ''Arbutus'', and that the split between the two groups of species occurred at the
Paleogene The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...
/
Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. ...
boundary. The 12 species are as follows:


Afro-Eurasia

* '' Arbutus andrachne'' L. – Greek strawberry tree (Southeastern Europe and southwestern Asia) * ''
Arbutus canariensis ''Arbutus canariensis'', known in Spanish as ''madroño canario'', is a species of shrub or tree in the Ericaceae, heath family. It is Endemism, endemic to the Canary Islands of Spain, specifically Tenerife, La Gomera, Gran Canaria, El Hierro, an ...
'' Duhamel – Canary madrone (
Canary Islands The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Morocc ...
) * '' Arbutus pavarii'' Pampan. ( Libya) * '' Arbutus unedo'' L. – strawberry tree (
Mediterranean Basin In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and w ...
, western France, and western Ireland)


Americas

* '' Arbutus arizonica'' (A.Gray) Sarg. – Arizona madrone ( New Mexico, Arizona and western Mexico south to
Jalisco Jalisco (, , ; Nahuatl: Xalixco), officially the Free and Sovereign State of Jalisco ( es, Estado Libre y Soberano de Jalisco ; Nahuatl: Tlahtohcayotl Xalixco), is one of the 31 states which, along with Mexico City, comprise the 32 Federal En ...
) * '' Arbutus bicolor'' S. González, M. González et P. D. Sørensen ( Mexico) * ''
Arbutus madrensis ''Arbutus madrensis'' is a Mexican species of trees in the Ericaceae, heath family. It is found in the Sierra Madre Occidental of Durango, Jalisco, Nayarit, and Sinaloa in western Mexico.Sørensen, P. D. 1995. ''Arbutus'' Linnaeus. Flora Neotropi ...
'' M. González – western Mexico * ''
Arbutus menziesii ''Arbutus menziesii'' or Pacific madrone (commonly madrone or madrona in the United States and arbutus in Canada), is a species of broadleaf evergreen tree in the family Ericaceae, native to the western coastal areas of North America, from Bri ...
'' Pursh – Pacific madrone (West coast of
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
from southern British Columbia to central (less frequently southern) California, on the west slopes of the
Sierra Nevada The Sierra Nevada () is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primarily ...
and
Pacific Coast Range The Pacific Coast Ranges (officially gazetted as the Pacific Mountain System in the United States) are the series of mountain ranges that stretch along the West Coast of North America from Alaska south to Northern and Central Mexico. Although the ...
mountains) * ''
Arbutus mollis ''Arbutus mollis'' is a species of plant in the heath family. It is found in Mexico.Acta Botanica Mexicana 101: 49-81 (2012).''Arbutus mollis''/ref> References mollis Mollis is a former municipality in the canton of Glarus in Switzerla ...
'' Kunth ( Mexico) * ''
Arbutus occidentalis ''Arbutus occidentalis'' or the Mexican Madrone is a small tree species in the heath family, that is endemic to Mexico. It is only known from a few areas in Western Mexico where it grows on rocky slopes. It produces red edible berries that are v ...
'' McVaugh & Rosatti - western Mexico * ''
Arbutus tessellata ''Arbutus tessellata'' is a Mexican species of shrubs in the heath family. It is widespread across much of northern and central Mexico from Chihuahua to Jalisco, Tlaxcala, and Veracruz Veracruz (), formally Veracruz de Ignacio de la Llave ...
'' (Mexico) * '' Arbutus xalapensis'' Kunth (syn. ''A. texana'', ''A. glandulosa'', ''A. peninsularis'') – Texas madrone ( Texas, New Mexico and northeastern Mexico)


Hybrids

* ''Arbutus'' × ''andrachnoides'' Link (''A. andrachne × A. unedo''): this hybrid has gained the Royal Horticultural Society's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
. * ''Arbutus'' × ''androsterilis'' (''A. canariensis'' × ''A. unedo'') in Canary Islands * ''Arbutus'' × ''thuretiana'' Demoly (''A. andrachne'' × ''A. canariensis'') * ''Arbutus'' × ''reyorum'' (''A. andrachne'' × ''A. canariensis'') × ''A. unedo''


Formerly placed here

*''
Arctostaphylos tomentosa ''Arctostaphylos tomentosa'' is a species of manzanita known by the common name woollyleaf manzanita or woolley manzanita. This shrub is endemic to California. It is a resident of chaparral canyons, foothills, and lower-elevation mountains. One ...
'' (Pursh) Lindl. (as ''A. tomentosa'' Pursh) *'' Arctostaphylos uva-ursi'' (L.) Spreng. (as ''A. uva-ursi'' L.) *''
Comarostaphylis discolor ''Comarostaphylis'' is a genus of shrubs in the heath family native to the Americas from California in the United States to Panama. These are hairy, glandular shrubs to small trees with shreddy bark, often quite similar to their close relatives, ...
'' (Hook.) Diggs (as ''A. discolor'' Hook.) *''
Gaultheria phillyreifolia ''Gaultheria'' is a genus of about 135 species of shrubs in the family Ericaceae. The name commemorates Jean François Gaultier of Quebec, an honour bestowed by the Scandinavian Pehr Kalm in 1748 and taken up by Carl Linnaeus in his '. These pl ...
'' (Pers.) Sleumer (as ''A. phillyreifolia'' Pers.)


Natural history

''Arbutus'' species are used as food plants by some
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
species including emperor moth, '' Pavonia pavonia'' and the
madrone butterfly ''Eucheira socialis'', commonly known as the Madrone butterfly is a lepidopteran that belongs to the family Pieridae. It was first described by Westwood in 1834. Locally known as ''Mariposa del madroño'' or ''tzauhquiocuilin'', it is endemic to ...
. The distribution of the latter species is in fact heavily affected by the distribution of the madrone. For Athenaios, it is the tree which Asclepiades of Myrlea talks about (Deiphnosophists, II.35)


Uses and symbolism

Several species are widely cultivated as ornamental plants outside of their natural ranges, though cultivation is often difficult due to their intolerance of root disturbance. The hybrid ''Arbutus'' 'Marina' is much more adaptable and thrives under garden conditions. The ''Arbutus unedo'' tree makes up part of
the coat of arms ''The'' () is a grammatical Article (grammar), article in English language, English, denoting persons or things that are already or about to be mentioned, under discussion, implied or otherwise presumed familiar to listeners, readers, or speak ...
(''El oso y el madroño'', The Bear and the Strawberry Tree) of the city of Madrid, Spain. A statue of a bear eating the fruit of the madroño tree stands in the center of the city (
Puerta del Sol The Puerta del Sol (English: "Gate of the Sun") is a public square in Madrid, one of the best known and busiest places in the city. This is the centre ('' Km 0'') of the radial network of Spanish roads. The square also contains the famous clo ...
). The image appears on city crests, taxi cabs, man-hole covers, and other city infrastructure. The ''Arbutus'' was important to the Straits
Salish Salish () may refer to: * Salish peoples, a group of First Nations/Native Americans ** Coast Salish peoples, several First Nations/Native American groups in the coastal regions of the Pacific Northwest ** Interior Salish peoples, several First Nat ...
people of Vancouver Island, who used arbutus bark and leaves to create medicines for colds, stomach problems, and tuberculosis, and as the basis for contraceptives. The tree also figured in myths of the Straits Salish. The fruit is edible but has minimal flavour and is not widely eaten. In Portugal, the fruit is sometimes
distilled Distillation, or classical distillation, is the process of separating the components or substances from a liquid mixture by using selective boiling and condensation, usually inside an apparatus known as a still. Dry distillation is the heating ...
(legally or not) into a potent brandy known as ''
medronho The Aguardente de Medronhos is a strong spirit, a traditional fruit brandy from Portugal, obtained from arbutus berries - the fruit of the Medronho tree, ''Arbutus unedo'' - also called strawberry tree. The ''Arbutus unedo'' grow wild on the poor ...
''. In Madrid, the fruit is distilled into madroño, a sweet, fruity liqueur. ''Arbutus'' is a good fuelwood tree since it burns hot and long. Many Pacific Northwest states in the United States use the wood of ''A. menziesii'' primarily as a heat source, as the wood holds no value in the production of homes since it does not grow in straight timbers. "My love's an arbutus" is the title of a poem by the Irish writer Alfred Perceval Graves (1846–1931), set to music by his compatriot Charles Villiers Stanford (1852–1924) The Canadian songwriter, singer and painter
Joni Mitchell Roberta Joan "Joni" Mitchell ( Anderson; born November 7, 1943) is a Canadian-American musician, producer, and painter. Among the most influential singer-songwriters to emerge from the 1960s folk music circuit, Mitchell became known for her sta ...
(born 1943), includes a reference to the "arbutus rustling" in her song, "For The Roses". It sounded like applause. She calls the arbutus tree her "favorite all-time tree". She had one outside her door in a house she built.


Cultural significance

According to the Straits
Salish Salish () may refer to: * Salish peoples, a group of First Nations/Native Americans ** Coast Salish peoples, several First Nations/Native American groups in the coastal regions of the Pacific Northwest ** Interior Salish peoples, several First Nat ...
, an anthropomorphic form of pitch would go fishing, but return to shore before it got too hot. One day he was too late getting back to shore and melted from the heat and several anthropomorphic trees rushed to get him – the first was
Douglas fir The Douglas fir (''Pseudotsuga menziesii'') is an evergreen conifer species in the pine family, Pinaceae. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Oregon pine, and Columbian pine. There are three va ...
, who took most of the pitch, the grand fir received a small portion, and the madrone received none – which is why they say it still has no pitch. Also, according to the Great Flood legends of several bands in the northwest, the madrone helped people survive by providing an anchor on top of a mountain. Because of this the Saanich people do not burn madrone out of thanks for saving them.''Plants of the Pacific Northwest Coast: Washington, Oregon, British Columbia & Alaska'', Paul Alaback,


Gallery

File:Ab plant 1359.jpg, '' Arbutus andrachne'' File:Arbutus canariensis kz7.JPG, ''
Arbutus canariensis ''Arbutus canariensis'', known in Spanish as ''madroño canario'', is a species of shrub or tree in the Ericaceae, heath family. It is Endemism, endemic to the Canary Islands of Spain, specifically Tenerife, La Gomera, Gran Canaria, El Hierro, an ...
'' File:Arbutus menziesii 3208s.JPG, ''
Arbutus menziesii ''Arbutus menziesii'' or Pacific madrone (commonly madrone or madrona in the United States and arbutus in Canada), is a species of broadleaf evergreen tree in the family Ericaceae, native to the western coastal areas of North America, from Bri ...
'' File:Cireres de pastor Cirerer d'arboç Arbutus unedo.jpg, '' Arbutus unedo'' File:Arbutus xalapensis Guadalupe Mountains.jpg, '' Arbutus xalapensis'' File:Arbutus × andrachnoides in Hackfalls Arboretum (2).jpg, The hybrid ''Arbutus'' × ''andrachnoides'' File:Arbutus x thuretiana2-Clapiers-7636~2020 03 23.jpg, The hybrid ''Arbutus'' × ''thuretiana''


See also

*'' Myrica rubra'', a different plant bearing a similar fruit, whose name is sometimes inaccurately translated from Chinese as ''Arbutus''


References

*


External links

{{Taxonbar, from=Q430057 Ericaceae genera