Araripedactylus
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''Araripedactylus'' was a large
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
pterodactyloid Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional ...
pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...
, known from a single wing bone found in the
Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
Santana Formation The Santana Group is a geologic group, formerly included as the middle part of the Araripe Group, in the Araripe Basin of northeastern Brazil. The group comprises the Crato, Ipubi and Romualdo Formations and is dated to the Aptian to Albian stag ...
of
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
. The genus was in 1977 named by
Peter Wellnhofer Peter Wellnhofer (born Munich, 1936) is a German paleontologist at the Bayerische Staatssammlung fur Paläontologie in Munich. He is best known for his work on the various fossil specimens of ''Archaeopteryx'' or "Urvogel", the first known bird. W ...
who, unaware that the name ''
Araripesaurus ''Araripesaurus'' is a genus a pterosaur belonging to the suborder Pterodactyloidea, it was discovered in the Romualdo Formation of the Santana Group in northeastern Brazil, which dates back to the Aptian and Albian of the Early Cretaceous. The ...
'' had been published by Price in 1971, presumed he was naming the first pterosaur genus from Brazil. The genus name refers to the
Araripe Plateau The Chapada do Araripe, also known as the Serra do Araripe, is a ''chapada'' (plateau) in northeastern Brazil. The chapada forms the boundary of Ceará and Pernambuco states, and forms the watershed between the Jaguaribe River of Ceará, which flow ...
and combines this with ''daktylos'', Greek for "finger", a common element in pterosaur names since ''
Pterodactylus ''Pterodactylus'' (from Greek () meaning 'winged finger') is an extinct genus of pterosaurs. It is thought to contain only a single species, ''Pterodactylus antiquus'', which was the first pterosaur to be named and identified as a flying rept ...
''. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is ''Araripedactylus dehmi''; the specific name honours the German paleontologist Richard Dehm, a professor at the
Munich Munich ( ; german: München ; bar, Minga ) is the capital and most populous city of the States of Germany, German state of Bavaria. With a population of 1,558,395 inhabitants as of 31 July 2020, it is the List of cities in Germany by popu ...
institute that acquired the only known specimen in 1975. The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
, BSP 1975 I 166, consists of a single
phalanx The phalanx ( grc, φάλαγξ; plural phalanxes or phalanges, , ) was a rectangular mass military formation, usually composed entirely of heavy infantry armed with spears, pikes, sarissas, or similar pole weapons. The term is particularly use ...
, the first of the right wing finger, embedded in an elongated chalk nodule. When the nodule was split to reveal the fossil, its distal end was damaged. The exemplar is that of an adult individual. The phalanx has a length of 55 centimetres. Its bone walls are described by Wellnhofer as exceptionally thick for a pterosaur, measuring . Wellnhofer, because of the lack of further information, placed ''Araripedactylus'' in a general Pterodactyloidea.
Alexander Kellner Alexander Wilhelm Armin Kellner (born September 26, 1961) is a Brazilian geologist and paleontologist who is a leading expert in the field of studying pterosaurs. His research has focused mainly on fossil reptiles from the Cretaceous Period, i ...
in 2000 assumed that in view of its provenance the species likely belonged to
Ornithocheiroidea Ornithocheiroidea (or ornithocheiroids) is a group of pterosaurs within the extinct suborder Pterodactyloidea. They were typically large pterosaurs that lived from the Early to Late Cretaceous periods (Valanginian to Maastrichtian stages), with f ...
(''sensu'' Kellner) and concluded that the phalanx was hard to distinguish from that of other large pterosaurs from the formation, such as ''
Anhanguera Anhanguera may refer to: People * Bartolomeu Bueno da Silva (1672–1740), a bandeirante Places in Brazil * Anhanguera, Goiás, a municipality in the state of Goiás * Anhanguera (district of São Paulo), a district in São Paulo * Parque Anhan ...
'' or ''
Tropeognathus ''Tropeognathus'' (meaning "keel jaw") is a genus of large pterosaurs from the late Early Cretaceous of South America. This genus is considered to be a member of the family Anhangueridae, however, several studies have also recovered it within a ...
''. He could not confirm the exceptional thickness of the bone walls, nor any other
autapomorphy In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to t ...
of the genus. The wingspan of ''Araripedactylus'' was estimated by Wellnhofer as at least 4.8 meters (15.75 ft), and in another publication at 5 meters (16.5 ft).


References


Literature

*Wellnhofer, P., 1991, "The Santana Formation Pterosaurs", In: J. G. Maisey (ed.), ''Santana fossils: An Illustrated Atlas'', T.F.H. Publications, Neptune, New Jersey, pp. 351–370


See also

*
Timeline of pterosaur research This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic era. Although pterosaurs w ...
*
List of pterosaurs This list of pterosaurs is a comprehensive listing of all genera that have ever been included in the order Pterosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered inva ...
{{Taxonbar, from=Q386325 Ornithocheiroids Early Cretaceous pterosaurs of South America Cretaceous Brazil Fossils of Brazil Romualdo Formation Fossil taxa described in 1977 Taxa named by Peter Wellnhofer