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''Coffea arabica'' (), also known as the Arabic coffee, is a species of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
in the coffee and madder family
Rubiaceae The Rubiaceae are a family of flowering plants, commonly known as the coffee, madder, or bedstraw family. It consists of terrestrial trees, shrubs, lianas, or herbs that are recognizable by simple, opposite leaves with interpetiolar stipules ...
. It is believed to be the first species of coffee to have been cultivated and is currently the dominant cultivar, representing about 60% of global production.
Coffee Coffee is a drink prepared from roasted coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulant, stimulating effect on humans, primarily due to its caffeine content. It is the most popular hot drink in the world. S ...
produced from the less acidic, more bitter, and more highly caffeinated robusta bean ('' C. canephora'') makes up most of the remaining coffee production. Arabica coffee originates from and was first cultivated in
Yemen Yemen (; ar, ٱلْيَمَن, al-Yaman), officially the Republic of Yemen,, ) is a country in Western Asia. It is situated on the southern end of the Arabian Peninsula, and borders Saudi Arabia to the Saudi Arabia–Yemen border, north and ...
, and documented by the 12th century. ''Coffea arabica'' is called () in
Arabic Arabic (, ' ; , ' or ) is a Semitic languages, Semitic language spoken primarily across the Arab world.Semitic languages: an international handbook / edited by Stefan Weninger; in collaboration with Geoffrey Khan, Michael P. Streck, Janet C ...
, borrowed from the
Amharic Amharic ( or ; (Amharic: ), ', ) is an Ethiopian Semitic language, which is a subgrouping within the Semitic branch of the Afroasiatic languages. It is spoken as a first language by the Amharas, and also serves as a lingua franca for all oth ...
"Buna".


Taxonomy

''Coffea arabica'' was first described scientifically by
Antoine de Jussieu Antoine de Jussieu (6 July 168622 April 1758) was a French naturalist, botanist, and physician. Jussieu was born in Lyon, the son of Christophe de Jussieu (or Dejussieu), an apothecary of some repute, who published a ''Nouveau traité de la ...
, who named it ''Jasminum arabicum'' after studying a specimen from the Botanic Gardens of Amsterdam.
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
placed it in its own genus ''
Coffea ''Coffea'' is a genus of flowering plants in the Family (biology), family Rubiaceae. ''Coffea'' species are shrubs or small trees native to tropical and southern Africa and tropical Asia. The seeds of some species, called coffee beans, are use ...
'' in 1737. ''Coffea arabica'' is the only
polyploid Polyploidy is a condition in which the cells of an organism have more than one pair of ( homologous) chromosomes. Most species whose cells have nuclei ( eukaryotes) are diploid, meaning they have two sets of chromosomes, where each set contain ...
species of the genus ''
Coffea ''Coffea'' is a genus of flowering plants in the Family (biology), family Rubiaceae. ''Coffea'' species are shrubs or small trees native to tropical and southern Africa and tropical Asia. The seeds of some species, called coffee beans, are use ...
,'' as it carries 4 copies of the 11 chromosomes (44 total) instead of the 2 copies of diploid species. Specifically, ''Coffea arabica'' is itself the result of a hybridization between the diploids ''
Coffea canephora ''Coffea canephora'' ( syn. ''Coffea robusta'', commonly known as ''robusta coffee'') is a species of coffee that has its origins in central and western sub-Saharan Africa. It is a species of flowering plant in the family Rubiaceae. Though wid ...
'' and ''Coffea eugenioides,'' thus making it an
allotetraploid Polyploidy is a condition in which the cells of an organism have more than one pair of ( homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two sets of chromosomes, where each set contains ...
, with ''two'' copies of two different genomes. This hybridization event at the origin of ''Coffea arabica'' is estimated between 1.08 million and 543,000 years ago and is linked to changing environmental conditions in East Africa.


Description

Wild plants grow between tall, and have an open branching system; the
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are opposite, simple elliptic-ovate to oblong, long and broad, glossy dark green. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s are white, 10–15 mm in diameter, and grow in axillary clusters. The seeds are contained in a
drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...
(commonly called a "cherry") 10–15 mm in diameter, maturing bright red to purple and typically containing two
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s, often called
coffee Coffee is a drink prepared from roasted coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulant, stimulating effect on humans, primarily due to its caffeine content. It is the most popular hot drink in the world. S ...
beans.


Distribution and habitat

Endemic to the southwestern
highlands Highland is a broad term for areas of higher elevation, such as a mountain range or mountainous plateau. Highland, Highlands, or The Highlands, may also refer to: Places Albania * Dukagjin Highlands Armenia * Armenian Highlands Australia *Sou ...
of
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
, ''Coffea arabica'' is now rare in Ethiopia, while many populations appear to be of mixed native and planted trees. ''Coffea arabica'' is today grown in dozens of countries between the
Tropic of Capricorn The Tropic of Capricorn (or the Southern Tropic) is the circle of latitude that contains the subsolar point at the December (or southern) solstice. It is thus the southernmost latitude where the Sun can be seen directly overhead. It also reac ...
and the
Tropic of Cancer The Tropic of Cancer, which is also referred to as the Northern Tropic, is the most northerly circle of latitude on Earth at which the Sun can be directly overhead. This occurs on the June solstice, when the Northern Hemisphere is tilted toward ...
. It is commonly used as an understorey shrub. It has also been recovered from the
Boma Plateau The Boma Plateau is a region in the east of South Sudan, located in the Jonglei and Eastern Equatoria provinces. It is inhabited by the Anuak, Murle and Toposa peoples. It contains important wetlands for birdlife in the region. Wildlife is threat ...
in
South Sudan South Sudan (; din, Paguot Thudän), officially the Republic of South Sudan ( din, Paankɔc Cuëny Thudän), is a landlocked country in East Africa. It is bordered by Ethiopia, Sudan, Central African Republic, Democratic Republic of the C ...
. ''Coffea arabica'' is also found on Mount
Marsabit Marsabit is a town in the northern Marsabit County in Kenya. It is situated in the former Eastern Province and is almost surrounded by the Marsabit National Park. The town is located east of the centre of the East African Rift at an elevation o ...
in northern
Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
, but it is unclear whether this is a truly native or naturalised occurrence; recent studies support it being naturalised. The species is widely naturalised in areas outside its native land, in many parts of
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
,
Latin America Latin America or * french: Amérique Latine, link=no * ht, Amerik Latin, link=no * pt, América Latina, link=no, name=a, sometimes referred to as LatAm is a large cultural region in the Americas where Romance languages — languages derived f ...
,
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
, and assorted islands in the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
and in the
Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
. The coffee tree was first brought to
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
in 1813, and it began to be extensively grown by about 1850. It was formerly more widely grown than at present, especially in Kona, and it persists after cultivation in many areas. In some valleys, it is a highly invasive weed. In the Udawattakele and Gannoruwa Forest Reserves near Kandy, Sri Lanka, coffee shrubs are also a problematic invasive species. Coffee has been produced in Queensland and New South Wales of Australia, starting in the 1980s and 90s. The Wet Tropics Management Authority has classified ''Coffea arabica'' as an environmental weed for southeast Queensland due to its invasiveness in non-agricultural areas.


History

The first written record of coffee made from
roasted Roasting is a cooking method that uses dry heat where hot air covers the food, cooking it evenly on all sides with temperatures of at least from an open flame, oven, or other heat source. Roasting can enhance the flavor through caramelization ...
coffee beans (botanical seeds) comes from
Arab The Arabs (singular: Arab; singular ar, عَرَبِيٌّ, DIN 31635: , , plural ar, عَرَب, DIN 31635: , Arabic pronunciation: ), also known as the Arab people, are an ethnic group mainly inhabiting the Arab world in Western Asia, ...
scholars, who wrote that it was useful in prolonging their working hours. The Arab innovation in Yemen of making a brew from roasted beans, spread first among the
Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Mediter ...
ians and
Turks Turk or Turks may refer to: Communities and ethnic groups * Turkic peoples, a collection of ethnic groups who speak Turkic languages * Turkish people, or the Turks, a Turkic ethnic group and nation * Turkish citizen, a citizen of the Republic o ...
, and later on found its way around the world. Other scholars believe that the coffee plant was introduced from Yemen, based on a Yemeni tradition that slips of both coffee and '' qat'' were planted at 'Udein' ('the two twigs') in Yemen in pre-Islamic times.
Arabica coffee production in Indonesia Indonesia was the fourth-largest producer of coffee in the world in 2014.http://www.ico.org/prices/po-production.pdf Coffee cultivation in Indonesia began in the late 1600s and early 1700s, in the early Dutch colonial period, and has played an ...
began in 1699 through the spread of Yemen's trade.
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
n coffees, such as
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
n and Java, are known for heavy body and low acidity. This makes them ideal for blending with the higher acidity coffees from
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
and
East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...
.


Cultivation and use

''Coffea arabica'' accounts for 60% of the world's coffee production. ''C. arabica'' takes approximately seven years to mature fully, and it does best with of rain, evenly distributed throughout the year. It is usually cultivated at an altitude between , but there are plantations that grow it as low as sea level and as high as . The plant can tolerate low temperatures, but not frost, and it does best with an average temperature between . Commercial
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...
s mostly only grow to about 5 m, and are frequently trimmed as low as 2 m to facilitate harvesting. Unlike ''
Coffea canephora ''Coffea canephora'' ( syn. ''Coffea robusta'', commonly known as ''robusta coffee'') is a species of coffee that has its origins in central and western sub-Saharan Africa. It is a species of flowering plant in the family Rubiaceae. Though wid ...
,'' ''C. arabica'' prefers to be grown in light shade. Two to four years after planting, ''C. arabica'' produces small, white, highly fragrant flowers. The sweet fragrance resembles the sweet smell of
jasmine Jasmine ( taxonomic name: ''Jasminum''; , ) is a genus of shrubs and vines in the olive family (Oleaceae). It contains around 200 species native to tropical and warm temperate regions of Eurasia, Africa, and Oceania. Jasmines are widely cultiva ...
flowers. Flowers opening on sunny days result in the greatest number of berries. This can be problematic and deleterious, however, as coffee plants tend to produce too many berries; this can lead to an inferior harvest and even damage yield in the following years, as the plant will favor the ripening of berries to the detriment of its own health. On well-kept plantations, overflowering is prevented by pruning the tree. The flowers only last a few days, leaving behind only the thick, dark-green leaves. The berries then begin to appear. These are as dark green as the foliage until they begin to ripen, at first to yellow and then light red and finally darkening to a glossy, deep red. At this point, they are called "
cherries A cherry is the fruit of many plants of the genus ''Prunus'', and is a fleshy drupe (stone fruit). Commercial cherries are obtained from cultivars of several species, such as the sweet ''Prunus avium'' and the sour ''Prunus cerasus''. The nam ...
", which fruit they then resemble, and are ready for picking. The berries are oblong and about 1 cm long. Inferior coffee results from picking them too early or too late, so many are picked by hand to be able to better select them, as they do not all ripen at the same time. They are sometimes shaken off the tree onto mats, which means ripe and unripe berries are collected together. The trees are difficult to cultivate and each tree can produce from of dried beans, depending on the tree's individual character and the climate that season. The most valuable part of this
cash crop A cash crop or profit crop is an Agriculture, agricultural crop which is grown to sell for profit. It is typically purchased by parties separate from a farm. The term is used to differentiate marketed crops from staple crop (or "subsistence crop") ...
is the beans inside. Each berry holds two
locule A locule (plural locules) or loculus (plural loculi) (meaning "little place" in Latin) is a small cavity or compartment within an organ or part of an organism (animal, plant, or fungus). In angiosperms (flowering plants), the term ''locule'' usu ...
s containing the beans. The coffee beans are actually two seeds within the fruit; sometimes, a third seed or one seed, a
peaberry Peaberry, known in Spanish as caracolillo, is a type of coffee bean. Normally the fruit ("cherry") of the coffee plant contains two seeds ("beans") that develop with flattened facing sides, but sometimes only one of the two seeds is fertilized, ...
, grows in the fruit at the tips of the branches. These seeds are covered in two membranes; the outer one is called the "parchment coat" and the inner one is called the "silver skin". On
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's List ...
, trees are planted at all times of the year and are harvested year-round. In parts of
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
, however, the trees have a season and are harvested only in winter. The plants are vulnerable to damage in such poor growing conditions as cold or low pH soil, and they are also more vulnerable to pests than the '' C. robusta'' plant. It is expected that a medium-term depletion of indigenous populations of ''C. arabica'' may occur, due to projected
global warming In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to E ...
, based on
IPCC The Intergovernmental Panel on Climate Change (IPCC) is an intergovernmental body of the United Nations. Its job is to advance scientific knowledge about climate change caused by human activities. The World Meteorological Organization (WMO) a ...
modelling. Climate change—rising temperatures, longer droughts, and excessive rainfall—appears to threaten the sustainability of arabica coffee production, leading to attempts to breed new cultivars for the changing conditions. Gourmet coffees are almost exclusively high-quality mild varieties of arabica coffee, and among the best known arabica coffee beans in the world are those from Jamaican Blue Mountain, Colombian Supremo, Tarrazú,
Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the no ...
,
Guatemala Guatemala ( ; ), officially the Republic of Guatemala ( es, República de Guatemala, links=no), is a country in Central America. It is bordered to the north and west by Mexico; to the northeast by Belize and the Caribbean; to the east by H ...
n
Antigua Antigua ( ), also known as Waladli or Wadadli by the native population, is an island in the Lesser Antilles. It is one of the Leeward Islands in the Caribbean region and the main island of the country of Antigua and Barbuda. Antigua and Bar ...
, and Ethiopian Sidamo.


Strains

One
strain Strain may refer to: Science and technology * Strain (biology), variants of plants, viruses or bacteria; or an inbred animal used for experimental purposes * Strain (chemistry), a chemical stress of a molecule * Strain (injury), an injury to a mu ...
of ''Coffea arabica'' naturally contains very little caffeine. While beans of normal ''C. arabica'' plants contain 12 mg of caffeine per gram of dry mass, these mutants contain only 0.76 mg of caffeine per gram, but with a taste similar to normal coffee.


Threats

Although it presently has a huge wild population of 13.5 to 19.5 billion individuals throughout its native range, ''C. arabica'' is still considered
Endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inva ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
due to numerous threats it faces. Due to being an
understory In forestry and ecology, understory (American English), or understorey (Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the forest canopy without penetrating it to any great extent, but abov ...
plant, it requires standing forest, making it highly susceptible to the historically significant
deforestation Deforestation or forest clearance is the removal of a forest or stand of trees from land that is then converted to non-forest use. Deforestation can involve conversion of forest land to farms, ranches, or urban use. The most concentrated d ...
levels in Ethiopia; prior to major deforestation, forest cover was thought to number between 25 - 31% of Ethiopia's total land surface, but now numbers just 4%, and deforestation still continues. In addition,
climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to E ...
may have a major effect on growing areas for wild ''C. arabica'' in Ethiopia due to its high-temperature sensitivity, and estimates indicate that population numbers could reduce by 50 - 80% with a 40 - 50% reduction in
area of occupancy Area is the quantity that expresses the extent of a region on the plane or on a curved surface. The area of a plane region or ''plane area'' refers to the area of a shape or planar lamina, while '' surface area'' refers to the area of an open s ...
by 2088; climate change can also impact reproductive success. In addition, the main pest of coffee, the coffee berry borer (''Hypothenemus hampei'') may stand to benefit from climate change and colonize higher altitudes that were formerly too cold for it, which can also impact coffee populations. The conservation of the genetic variation of ''C. arabica'' relies on conserving healthy populations of wild coffee in the
Afromontane The Afromontane regions are subregions of the Afrotropical realm, one of the Earth's eight biogeographic realms, covering the plant and animal species found in the mountains of Africa and the southern Arabian Peninsula. The Afromontane regions ...
rainforests Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainforest ...
of Yemen. Genetic research has shown coffee cultivation is threatening the genetic integrity of wild
coffee Coffee is a drink prepared from roasted coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulant, stimulating effect on humans, primarily due to its caffeine content. It is the most popular hot drink in the world. S ...
because it exposes wild
genotypes The genotype of an organism is its complete set of genetic material. Genotype can also be used to refer to the alleles or variants an individual carries in a particular gene or genetic location. The number of alleles an individual can have in a ...
to
cultivars A cultivar is a type of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and when Plant propagation, propagated retain those traits. Methods used to propagate cultivars include: division, root and st ...
. Nearly all of the coffee that has been cultivated over the past few centuries originated with just a handful of wild plants from Yemen, and today the coffee growing on plantations around the world contains less than 1% of the diversity contained in the wild in Yemen alone. Climate change also serves as a threat to cultivated ''C. arabica'' due to their temperature sensitivity, and some studies estimate that by 2050, over half of the land used for cultivating coffee could be unproductive. The more heat-tolerant ''
Coffea stenophylla ''Coffea stenophylla'', also known as highland coffee or Sierra Leone coffee, is a species of ''Coffea'' originating from West Africa. It is currently not commercially cultivated due to its low yield and small berries, which makes it inferior to ...
'' may replace ''C. arabica'' as the dominant coffee species in cultivation in order to guard against this.


Gallery

File:Coffee germination.jpg, Coffee germinating. File:Starr 070617-7322 Coffea arabica.jpg, ''Coffea arabica'' growing at
Olinda Olinda () is a historic city in Pernambuco, Brazil, in the Northeast Region, Brazil, Northeast Region. It is located on the country's northeastern Atlantic Ocean coast, in the Recife metropolitan area, Metropolitan Region of Recife, the state capi ...
,
Maui The island of Maui (; Hawaiian: ) is the second-largest of the islands of the state of Hawaii at 727.2 square miles (1,883 km2) and is the 17th largest island in the United States. Maui is the largest of Maui County's four islands, which ...
. File:Coffee arabica flower.jpg, Coffee flowers. File:Coffee arabica cherry.jpg, Fresh coffee fruits. File:Open coffee cherry.jpg, Fresh coffee seeds ("beans"). File:Fermented coffee beans.jpg, Fermented coffee seeds. File:Green arabica coffee beans.jpg, Fermented coffee (green) seeds without hulls. File:Roasted arabica coffee beans Minca.jpg, Fermented and roasted coffee seeds. File:Unroasted coffee.jpg, Unroasted ("green") coffee (''Coffea arabica'') seeds from
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
.


See also

* Maraba coffee


References


Further reading

* *


External links


World Checklist of Rubiaceae

Understanding the difference between Arabica and Robusta


{{Authority control arabica Flora of East Tropical Africa Flora of Northeast Tropical Africa Endemic flora of Ethiopia Flora of Kenya Flora of Sudan Crops originating from Africa Crops originating from Ethiopia Agriculture in Brazil Agriculture in the Caribbean Agriculture in China Agriculture in Central America Agriculture in Mexico Agriculture in South America Bale Mountains Ethiopian Highlands Invasive plant species in Sri Lanka Plants described in 1753 Taxa named by Carl Linnaeus Coffee