Anserimimus Planinychus
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''Anserimimus'' ( ; "goose mimic") is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
ornithomimid Ornithomimidae (meaning "bird-mimics") is a family of theropod dinosaurs which bore a superficial resemblance to modern ostriches. Ornithomimids were fast, omnivorous or herbivorous dinosaurs known mainly from the Late Cretaceous Period of Laura ...
theropod
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
, from the Late
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
Period of what is now
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
. It was a lanky, fast-running animal, possibly an
omnivore An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutr ...
. From what
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s are known, it probably closely resembled other ornithomimids, except for its more powerful forelimbs.


Discovery

''Anserimimus'' was found in the Mongolian
aimag An aimag (, ; xal, әәмг, ), originally a Mongolian word meaning 'tribe', is an administrative subdivision in Mongolia, Russia, and in the Inner Mongolia region of China. Mongolia In Mongolia, an aimag is the first-level administrative ...
, or province, of
Bayankhongor Bayankhongor ( mn, Баянхонгор, lit=''righ soft'', or ''rich sweetheart'') is the capital of the Bayankhongor Province (aimag) in Mongolia. The administration of the Bayankhongor Sum (district) is also located in the same place. The city ...
during a joint
Soviet The Soviet Union,. officially the Union of Soviet Socialist Republics. (USSR),. was a List of former transcontinental countries#Since 1700, transcontinental country that spanned much of Eurasia from 1922 to 1991. A flagship communist state, ...
-Mongolian expedition to the
Gobi Desert The Gobi Desert (Chinese: 戈壁 (沙漠), Mongolian: Говь (ᠭᠣᠪᠢ)) () is a large desert or brushland region in East Asia, and is the sixth largest desert in the world. Geography The Gobi measures from southwest to northeast an ...
, in the late 1970s. Mongolian
paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
Rinchen Barsbold named ''Anserimimus'' in 1988, combining the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''anser'' meaning 'goose' with the Greek ''mimos'' meaning 'mimic'. ''Anser'' is the generic name of several species of geese. Although ''Anserimimus'' does not specifically resemble a goose, ornithomimosaurs have traditionally been named after different types of birds, such as '' Struthiomimus'' ('ostrich mimic'), ''
Gallimimus ''Gallimimus'' ( ) is a genus of theropod dinosaur that lived in what is now Mongolia during the Late Cretaceous period, about seventy million years ago (mya). Several fossils in various stages of growth were discovered by Polish-Mongolian exped ...
'' ('rooster mimic'), and ''
Pelecanimimus ''Pelecanimimus'' (meaning "pelican mimic") is an extinct genus of basal ("primitive") ornithomimosaurian dinosaur from the Early Cretaceous of Spain. It is notable for possessing more teeth than any other member of the Ornithomimosauria (or an ...
'' ('pelican mimic'). The one named species of ''Anserimimus'' is called ''Anserimimus planinychus''. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
is derived from the Latin ''planus'' meaning 'flat', and the Ancient Greek ὄνυξ, ''onyx'', meaning 'claw', in reference to the peculiar flattened claws which characterize the genus. There is only a single specimen of ''Anserimimus'', its
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
IGM 100/300. It consists of a rather complete and articulated skeleton lacking the skull and lower jaws. Limited information has been published on the anatomy of ''Anserimimus'', as Barsbold did not describe most of these bones, instead focusing on only those with features that set ''Anserimimus'' apart from other ornithomimids. In an unpublished thesis Robert Bronowicz in 2005 gave a detailed description of the species, also referring additional material to it, among which a second partial skeleton, specimen ZPAL MgD-I/65. However, in 2010 Bronowicz concluded this other find, though most closely related to ''Anserimimus planinychus'', probably represented a separate
taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
.


Description

''Anserimimus'' was a medium-sized ornithomimosaurian. Gregory S. Paul in 2010 estimated its length at three metres, its weight at fifty kilogrammes. The thighbone is 435 millimetres long. There are several key differences between it and related species, though most of these are shared with ZPAL MgD-I/65. The claws on the hand are long, low at the back and rather straight, only slightly curved, with the lower surface nearly flat. The forelimb is long and also built more powerfully than that of other ornithomimids, with large crests on the scapulocoracoid of the shoulder and
humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...
(upper arm bone), which provided attachment points for large arm muscles like the
biceps The biceps or biceps brachii ( la, musculus biceps brachii, "two-headed muscle of the arm") is a large muscle that lies on the front of the upper arm between the shoulder and the elbow. Both heads of the muscle arise on the scapula and join ...
. The
metacarpus In human anatomy, the metacarpal bones or metacarpus form the intermediate part of the skeletal hand located between the phalanges of the fingers and the carpal bones of the wrist, which forms the connection to the forearm. The metacarpal bones ...
of the hand is fused, adding to the strength. The foot is strongly arctometatarsalian, with the third metatarsal excluded from the front surface of the metatarsus over 40% of its upper length.


Phylogeny

''Anserimimus'' was by Barsbold assigned to the
Ornithomimidae Ornithomimidae (meaning "bird-mimics") is a family of theropod dinosaurs which bore a superficial resemblance to modern ostriches. Ornithomimids were fast, omnivorous or herbivorous dinosaurs known mainly from the Late Cretaceous Period of Lauras ...
, a group of
derived Derive may refer to: * Derive (computer algebra system), a commercial system made by Texas Instruments * ''Dérive'' (magazine), an Austrian science magazine on urbanism *Dérive, a psychogeographical concept See also * *Derivation (disambiguatio ...
Ornithomimosauria Ornithomimosauria ("bird-mimic lizards") are theropod dinosaurs which bore a superficial resemblance to the modern-day ostrich. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period (geology), Period of Laurasia (now Asia ...
. This has been confirmed by modern cladistic analysis. Its closest relative may be ''Gallimimus'' from the same formation albeit from a different area (Kobayashi & Lu, 2003; Kobayashi & Barsbold, 2005). Other studies have been unable specifically to determine its relationships or those of any other ornithomimids (Ji ''et al.'', 2003; Makovicky ''et al.'', 2004). The following cladogram is based on Xu ''et al.'', 2011:


Paleobiology

The function of the powerful arm, with its straightened claws, remains unknown. It may indicate a different diet of food-gathering strategy than other ornithomimids, although its diet is difficult to determine, since the animal's skull is unknown. Scientists have long
hypothesized A hypothesis (plural hypotheses) is a proposed explanation for a phenomenon. For a hypothesis to be a scientific hypothesis, the scientific method requires that one can test it. Scientists generally base scientific hypotheses on previous obser ...
that ornithomimids, descended from
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
theropod ancestors, were actually omnivores or even
herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
s.


Paleoecology

''Anserimimus'' was recovered from the Nemegt Formation of
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
. The Nemegt is thought to represent
alluvial plain An alluvial plain is a largely flat landform created by the deposition of sediment over a long period of time by one or more rivers coming from highland regions, from which alluvial soil forms. A floodplain is part of the process, being the sma ...
s containing
meander A meander is one of a series of regular sinuous curves in the channel of a river or other watercourse. It is produced as a watercourse erodes the sediments of an outer, concave bank ( cut bank) and deposits sediments on an inner, convex bank ...
ing
river A river is a natural flowing watercourse, usually freshwater, flowing towards an ocean, sea, lake or another river. In some cases, a river flows into the ground and becomes dry at the end of its course without reaching another body of wate ...
s. The layer ''Anserimimus'' was found, is dating from the early
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
stage of the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
, or about 70
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
. Aside from ''Gallimimus'', other theropods found in the Nemegt Formation include the gigantic '' Tarbosaurus'' and ''
Deinocheirus ''Deinocheirus'' ( ) is a genus of large ornithomimosaur that lived during the Late Cretaceous around 70 million years ago. In 1965, a pair of large arms, shoulder girdles, and a few other bones of a new dinosaur were first discovered in the Ne ...
'', as well as smaller
dromaeosaurid Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning ...
s, oviraptorosaurs, troodontids, and
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s.
Herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
s are represented by the
hadrosaurid Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which inclu ...
s ''
Barsboldia ''Barsboldia'' (meaning "of Barsbold", a well-known Mongolian paleontologist) is a genus of large hadrosaurid dinosaur from the early Maastrichtian Nemegt Formation of Ömnogöv', Mongolia. It is known from a partial vertebral column, parti ...
'' and ''
Saurolophus ''Saurolophus'' (; meaning "lizard crest") is a genus of large hadrosaurid dinosaur from the Late Cretaceous period of Asia and North America, that lived in what is now the Horseshoe Canyon and Nemegt formations about 70 million to 68 million ...
'', the ankylosaurid '' Tarchia'' and several titanosaurian
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...
s and pachycephalosaurians. The rock
facies In geology, a facies ( , ; same pronunciation and spelling in the plural) is a body of rock with specified characteristics, which can be any observable attribute of rocks (such as their overall appearance, composition, or condition of formatio ...
of this formation suggest the presence of stream and river channels, mudflats, and shallow lakes. Sediments also indicate that there existed a rich habitat, offering diverse food in abundant amounts that could sustain massive Cretaceous dinosaurs.Novacek, M. (1996). Dinosaurs of the Flaming Cliffs. Bantam Doubleday Dell Publishing Group Inc. New York, New York.


See also

*
Timeline of ornithomimosaur research This timeline of ornithomimosaur research is a chronological listing of events in the history of paleontology focused on the ornithomimosaurs, a group of bird-like theropods popularly known as the ostrich dinosaurs. Although fragmentary, probable, ...


References


External links

*
Anserimimus
' at The Theropod Database {{Taxonbar, from1=Q132481 Late Cretaceous dinosaurs of Asia Fossil taxa described in 1988 Ornithomimids Taxa named by Rinchen Barsbold Maastrichtian life Nemegt fauna