Titanosauria
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Titanosauria
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as ''Patagotitan''—estimated at long with a weight of —and the comparably-sized ''Argentinosaurus'' and ''Puertasaurus'' from the same region. The group's name alludes to the mythological Titans of ancient Greek mythology, via the type genus (now considered a ''nomen dubium)'' ''Titanosaurus''. Together with the brachiosaurids and relatives, titanosaurs make up the larger sauropod clade Titanosauriformes. Titanosaurs have long been a poorly-known group, and the relationships between titanosaur species are still not well-understood. Description Titanosauria have the largest ran ...
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Dreadnoughtus
''Dreadnoughtus'' is a genus of titanosaurian sauropod dinosaur containing a single species, ''Dreadnoughtus schrani''. ''D. schrani'' is known from two partial skeletons discovered in Upper Cretaceous (Campanian to Maastrichtian; approximately 76–70 Ma) rocks of the Cerro Fortaleza Formation in Santa Cruz Province, Argentina. It is one of the largest terrestrial vertebrates known, with the immature type specimen reaching in total body length and possessing the greatest mass of any land animal that can be calculated with reasonable certainty. ''D. schrani'' is known from more complete skeletons than any other gigantic titanosaurian. Drexel University paleontologist Kenneth Lacovara, who discovered the species, chose the name ''Dreadnoughtus'', which means “fears nothing", stating “I think it’s time the herbivores get their due for being the toughest creatures in an environment." Discovery and study American palaeontologist Kenneth Lacovara discovered the remains in th ...
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Patagotitan
''Patagotitan'' is a genus of titanosaurian sauropod dinosaur from the Cerro Barcino Formation in Chubut Province, Patagonia, Argentina. The genus contains a single species known from multiple individuals: ''Patagotitan mayorum'', first announced in 2014 and then validly named in 2017 by José Carballido, Diego Pol, and colleagues. Contemporary studies estimated the length of the type specimen, a young adult, at with a weight estimated at , later revised to . Discovery Remains of ''Patagotitan mayorum'', a part of a lower thighbone, were initially discovered in 2008 by a farm laborer, Aurelio Hernández, in the desert near La Flecha, Argentina, about west of Trelew. Excavation was done by palaeontologists from the Museum of Paleontology Egidio Feruglio. The lead scientists on the excavation were Jose Luis Carballido and Diego Pol, with partial funding from The Jurassic Foundation. Between January 2013 and February 2015, seven paleontological field expeditions were carri ...
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Hamititan
''Hamititan'' (meaning "Hami giant") is a genus of titanosaur sauropod dinosaur from the Shengjinkou Formation of Xinjiang, China. It contains one species, the type species, ''Hamititan xinjiangensis''. Discovery and naming In 2006, a Konservat-Lagerstätte was reported from the Shengjinkou Formation in the Hami region of Xinjiang, China. This consisted of lake sediments allowing for exceptional preservation of fossils. The same year, Qiu Zhanxiang and Wang Banyue started official excavations. Among the excavated fossils were seven caudal vertebrae with three chevrons preserved. These were established as the holotype (HM V22) of the new sauropod taxon, ''Hamititan''. Four sacral elements, specimen IVPP V27875, were not referred. A theropod tooth was found near the sixth caudal vertebra of the holotype. It is likely that this theropod preyed on the holotype shortly after it died. The generic name refers to the city of Hami, where the holotype was found, while the specific nam ...
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Austroposeidon
''Austroposeidon'' is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Presidente Prudente Formation of Brazil. It contains one species, ''Austroposeidon magnificus'' (meaning "Magnificent Southern Poseidon"). Discovery and naming ''Austroposeidon'' is known from a single specimen, MCT 1628-R, which consists of portions of the cervical (neck), dorsal (back), and sacral (hip) vertebrae (including a cervical rib and one complete dorsal vertebra). The specimen was discovered in the Campanian-Maastrichtian Presidente Prudente Formation of the Bauru Group by palaeontologist Llewellyn Ivor Price in 1953, but the remains were not described until 2016. The animal was likely preserved by a crevasse splay on a floodplain, judging by the fine sandstone that the specimen was found in. Unfortunately, the site where the specimen was recovered has now been lost to urban development. The genus name combines ''austro'' ("southern", as in South America, from Latin ''Auster'', ...
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Jiangshanosaurus
''Jiangshanosaurus'' is a genus of herbivorous titanosauriform sauropod dinosaur that lived in China approximately 92-88 million years ago, during the Turonian- Coniacian stage of the Late Cretaceous. Discovery and naming In 1977 and 1978 a sauropod skeleton was excavated by paleontologists Wei Feng, Wu Weitang and Kang Ximin in the Jinhua Formation of Lixian Village, Jiangshan county, in the eastern Chinese province of Zhejiang. The type and only named species, ''Jiangshanosaurus lixianensis'', was formally described by Tang Feng, Kang, Jin Xingsheng, Wei and Wu in 2001. The holotype, ZNM M1322, of ''J.lixianensis'' includes elements of the left shoulder, five back vertebrae, three tail vertebrae, the pubic bones, the ischia, and a left femur.Feng Tang, Xi-Min Kang, Xing-Sheng Jin, Feng Wei, Wei-Tang Wu (2001"A New Sauropod Dinosaur of Cretaceous From Jiangshan, Zhejiang Province"in: ''Vertebrata PalAsiatica.'' Bd. 39, Nr. 4, pp. 272–281. The genus name refers to Jiangsh ...
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Dongyangosaurus
''Dongyangosaurus'' is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous. The only species is ''Dongyangosaurus sinensis'', from which only a single fragmentary skeleton is known, coming from the Zhejiang province of eastern China. It was described and named by Lü Junchang and colleagues. Like other sauropods, ''Dongyangosaurus'' would have been a large quadrupedal herbivore. Description The only skeleton (holotype DYM 04888) is stored in the Dongyang Museum (Dongyang, Zhejiang). It consists of ten dorsal vertebrae, the sacrum, two caudal vertebrae as well as the complete pelvis. The skeleton was found articulated. ''Dongyangosaurus'' was a midsized sauropod, measuring approximately 50 ft (15 m) in length and 15 ft (5 m) in height. The dorsal vertebrae were characterized by eye shaped pleurocoels and low bifurcated neural spines. The sacrum consisted of six fused sacral vertebrae, a feature unique to somphospondylans. The caudal vertebrae were amphic ...
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Daxiatitan
''Daxiatitan'' (; meaning "Daxia giant" after a tributary of the Yellow River) is a genus of sauropod dinosaur known from the Lower Cretaceous of Gansu, China. Its type and only species is ''Daxiatitan binglingi'' (). It is known from a single partial skeleton consisting of most of the neck and back vertebrae, two tail vertebrae, a shoulder blade, and a thigh bone. At the time of its discovery in 2008, ''Daxiatitan'' was regarded as potentially the largest known dinosaur from China. Taxonomy ''Daxiatitan'' and its type and only species ''Daxiatitan binglingi'' were named by You Hailu, Li Daqing, Zhou Lingqi, and Ji Qiang in 2008. The holotype of ''D. binglingi'', GSLTZP03-001, was collected from the Hekou Group, in Gansu Province, and consists of ten cervical, ten dorsal, and two caudal vertebrae, cervical and dorsal ribs, a haemal arch, a scapulocoracoid, and a femur. The genus name refers to the Daxia River, a tributary of the Yellow River that runs through the area where th ...
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Choconsaurus
''Choconsaurus'' is an extinct genus of herbivorous sauropod dinosaur belonging to the group Titanosauriformes, which lived in the area of present-day Argentina at the end of the Cretaceous. Discovery and naming The holotype (specimen MMCh-PV 44/10) was found in Villa El Chocón in Neuquén province, Argentina by Viviana Moro before 1996. Subsequent field campaigns carried out by the Ernesto Bachmann Paleontological Museum and the Museum of Geology and Paleontology of the National University of Comahue between 1996 and 2002 discovered more remains pertaining to ''Choconsaurus'', with several of the specimens described by Calvo (1999) under "Titanosauridae indet.".Calvo, J.O. (1999). Dinosaurs and other vertebrates of the Lake Ezequiel Ramos Mexía Area, Neuquén, Patagonia, Argentina. ''National Museum Monographs'' 15: 13–45. In 2017, the species type ''Choconsaurus baileywillisi'' was named and described by Edith Simón, Leonardo Salgado and Jorge Orlando Calvo. The genu ...
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Campylodoniscus
''Campylodoniscus'' is a genus of titanosaur sauropod dinosaur from the Late Cretaceous Period of what is now Argentina. The type species was first named and described by Friedrich von Huene in 1929 as ''Campylodon ameghinoi'', the genus name meaning 'bent tooth', from Greek καμπυλος, 'bent' or 'curved' (as of a bow) and ὀδών meaning 'tooth'. The specific name honours Florentino Ameghino. In 1961 H. Haubold and O. Kuhn noted that the name was pre-occupied by a fish and renamed the genus into ''Campylodoniscus'', the diminutive.Haubold, H. & Kuhn, O., 1961, ''Lebensbilder und Evolution fossiler Saurier, Amphibien und Reptilien'', Wittenberg : Ziemsen The fossil remains of ''Campylodoniscus'' were found in the Bajo Barreal Formation and consist of a single jaw bone, the maxilla, holding seven teeth. ''Campylodoniscus'' dates to the Cenomanian (95 Ma). It is sometimes estimated as being around twenty meters in length. ''Campylodoniscus'' is probably a member of ...
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Brasilotitan
''Brasilotitan'' (meaning "Brazil giant") is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous (early Maastrichtian) Adamantina Formation of Brazil. The type species is ''Brasilotitan nemophagus''. Material and morphology The holotype was collected near Presidente Prudente city, São Paulo state. It consists of a dentary, cervical and sacral vertebrae, one ungual, and remains of the pelvic region. The mandible has an 'L' shaped morphology, with the symphyseal region of the dentary slightly twisted medially, a feature never recorded before in any titanosaur. Phylogeny Although the phylogenetic position of ''Brasilotitan'' is difficult to establish, the new species is neither basal nor a derived member of Titanosauria. Based on lower jaw morphology, it appears to be closely related to ''Antarctosaurus ''Antarctosaurus'' (; meaning "southern lizard") is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. T ...
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Bonatitan
''Bonatitan'' is a genus of titanosaurian dinosaur from the Late Cretaceous Allen Formation of Argentina. It was named in 2004. Description The type species is ''Bonatitan reigi'', first described by Martinelli and Forasiepi in 2004. The specific epithet honours Osvaldo Reig. The holotype, MACN-PV RN 821, originally included a braincase and caudal vertebrae as well as limb elements. However, Salgado et al. (2014) emended the holotype to include the braincase only, and treated other elements catalogued under MACN-PV RN 821 as belonging to separate individual based on size and relative proportions.Salgado L., Gallina P.A. and Paulina Carabajal A. 2014. "Redescription of ''Bonatitan reigi'' (Sauropoda: Titanosauria), from the Campanian–Maastrichtian of the Río Negro Province (Argentina)". Historical Biology: An International Journal of Paleobiology 27(5): 525-548 The genus and species names honor the famous Argentine paleontologists José Fernando Bonaparte and Osvaldo Reig O ...
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Kaijutitan
''Kaijutitan'' (meaning "Kaiju titan" after the type of Japanese movie monsters) is a genus of basal titanosaur dinosaur from the Sierra Barrosa Formation from Neuquén Province in Argentina. The type and only species is ''Kaijutitan maui''. Discovery and naming ''Kaijutitan'' was discovered by a team of researchers from the Museo Municipal Argentino Urquiza and the Museo Provincial de Ciencias Naturales “Prof. Dr. Juan Olsacher" from a layer from the Sierra Barrosa Formation, in Cañadón Mistringa, about 9 km southwest of the city of Rincón de los Sauces, in Neuquen, Argentina. Description ''Kaijutitan'' is known from the holotype MAU-Pv-CM-522, stored at the Museo Municipal Argentino Urquiza, Paleontología de Vertebrados, Cañadón Mistringa, which is a partial skeleton preserving elements from several parts of the skeleton. It can be distinguished from other titanosaurians through the possession of unique traits: the width between the basal tuberosities is almost fo ...
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