''Anserimimus'' ( ; "goose mimic") is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
ornithomimid
Ornithomimidae (meaning " bird-mimics") is a family of theropod dinosaurs which bore a superficial resemblance to modern ostriches. Ornithomimids were fast, omnivorous or herbivorous dinosaurs known mainly from the Late Cretaceous Period of ...
theropod
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally ...
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
, from the Late
Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
Period of what is now
Mongolia
Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 millio ...
. It was a lanky,
fast-running animal, possibly an
omnivore
An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nut ...
. From what
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s are known, it probably closely resembled other ornithomimids, except for its more powerful forelimbs.
Discovery
''Anserimimus'' was found in the Mongolian
aimag, or province, of
Bayankhongor during a joint
Soviet
The Soviet Union,. officially the Union of Soviet Socialist Republics. (USSR),. was a transcontinental country that spanned much of Eurasia from 1922 to 1991. A flagship communist state, it was nominally a federal union of fifteen national ...
-Mongolian expedition to the
Gobi Desert
The Gobi Desert ( Chinese: 戈壁 (沙漠), Mongolian: Говь (ᠭᠣᠪᠢ)) () is a large desert or brushland region in East Asia, and is the sixth largest desert in the world.
Geography
The Gobi measures from southwest to northeast ...
, in the late 1970s.
Mongolian
paleontologist
Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of foss ...
Rinchen Barsbold named ''Anserimimus'' in 1988, combining the
Latin
Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...
''anser'' meaning 'goose' with the
Greek
Greek may refer to:
Greece
Anything of, from, or related to Greece, a country in Southern Europe:
*Greeks, an ethnic group.
*Greek language, a branch of the Indo-European language family.
**Proto-Greek language, the assumed last common ancestor ...
''mimos'' meaning 'mimic'. ''Anser'' is the generic name of several species of
geese
A goose ( : geese) is a bird of any of several waterfowl species in the family Anatidae. This group comprises the genera ''Anser'' (the grey geese and white geese) and '' Branta'' (the black geese). Some other birds, mostly related to the she ...
. Although ''Anserimimus'' does not specifically resemble a goose, ornithomimosaurs have traditionally been named after different types of birds, such as ''
Struthiomimus'' ('ostrich mimic'), ''
Gallimimus'' ('rooster mimic'), and ''
Pelecanimimus
''Pelecanimimus'' (meaning "pelican mimic") is an extinct genus of basal ("primitive") ornithomimosaurian dinosaur from the Early Cretaceous of Spain. It is notable for possessing more teeth than any other member of the Ornithomimosauria (or an ...
'' ('pelican mimic'). The one named species of ''Anserimimus'' is called ''Anserimimus planinychus''. The
specific name Specific name may refer to:
* in Database management systems, a system-assigned name that is unique within a particular database
In taxonomy, either of these two meanings, each with its own set of rules:
* Specific name (botany), the two-part (bino ...
is derived from the Latin ''planus'' meaning 'flat', and the Ancient Greek ὄνυξ, ''onyx'', meaning 'claw', in reference to the peculiar flattened claws which characterize the genus.
There is only a single specimen of ''Anserimimus'', its
holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
IGM 100/300. It consists of a rather complete and articulated skeleton lacking the skull and lower jaws. Limited information has been published on the anatomy of ''Anserimimus'', as Barsbold did not describe most of these bones, instead focusing on only those with features that set ''Anserimimus'' apart from other ornithomimids. In an unpublished thesis
Robert Bronowicz
The name Robert is an ancient Germanic given name, from Proto-Germanic "fame" and "bright" (''Hrōþiberhtaz''). Compare Old Dutch ''Robrecht'' and Old High German ''Hrodebert'' (a compound of '' Hruod'' ( non, Hróðr) "fame, glory, honou ...
in 2005 gave a detailed description of the species, also referring additional material to it, among which a second partial skeleton, specimen ZPAL MgD-I/65. However, in 2010 Bronowicz concluded this other find, though most closely related to ''Anserimimus planinychus'', probably represented a separate
taxon
In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...
.
Description
''Anserimimus'' was a medium-sized ornithomimosaurian.
Gregory S. Paul in 2010 estimated its length at three metres, its weight at fifty kilogrammes. The thighbone is 435 millimetres long.
There are several key differences between it and related species, though most of these are shared with ZPAL MgD-I/65. The claws on the hand are long, low at the back and rather straight, only slightly curved, with the lower surface nearly flat. The forelimb is long and also built more powerfully than that of other ornithomimids, with large crests on the
scapulocoracoid of the shoulder and
humerus
The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...
(upper arm bone), which provided attachment points for large arm muscles like the
biceps
The biceps or biceps brachii ( la, musculus biceps brachii, "two-headed muscle of the arm") is a large muscle that lies on the front of the upper arm between the shoulder and the elbow. Both heads of the muscle arise on the scapula and join ...
. The
metacarpus
In human anatomy, the metacarpal bones or metacarpus form the intermediate part of the skeletal hand located between the phalanges of the fingers and the carpal bones of the wrist, which forms the connection to the forearm. The metacarpal bones ...
of the hand is fused, adding to the strength. The foot is strongly arctometatarsalian, with the third
metatarsal excluded from the front surface of the
metatarsus over 40% of its upper length.
Phylogeny
''Anserimimus'' was by Barsbold assigned to the
Ornithomimidae, a group of
derived Ornithomimosauria. This has been confirmed by modern
cladistic
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived ch ...
analysis. Its closest relative may be ''Gallimimus'' from the same formation albeit from a different area (Kobayashi & Lu, 2003; Kobayashi & Barsbold, 2005). Other studies have been unable specifically to determine its relationships or those of any other ornithomimids (Ji ''et al.'', 2003; Makovicky ''et al.'', 2004).
The following cladogram is based on Xu ''et al.'', 2011:
Paleobiology
The function of the powerful arm, with its straightened claws, remains unknown. It may indicate a different diet of food-gathering strategy than other ornithomimids, although its diet is difficult to determine, since the animal's skull is unknown. Scientists have long
hypothesized
A hypothesis (plural hypotheses) is a proposed explanation for a phenomenon. For a hypothesis to be a scientific hypothesis, the scientific method requires that one can test it. Scientists generally base scientific hypotheses on previous obser ...
that ornithomimids, descended from
carnivorous
A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other ...
theropod ancestors, were actually omnivores or even
herbivore
A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthp ...
s.
Paleoecology
''Anserimimus'' was recovered from the
Nemegt Formation
The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians ...
of
Mongolia
Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 millio ...
. The Nemegt is thought to represent
alluvial plain
An alluvial plain is a largely flat landform created by the deposition of sediment over a long period of time by one or more rivers coming from highland regions, from which alluvial soil forms. A floodplain is part of the process, being the s ...
s containing
meander
A meander is one of a series of regular sinuous curves in the channel of a river or other watercourse. It is produced as a watercourse erodes the sediments of an outer, concave bank ( cut bank) and deposits sediments on an inner, convex ban ...
ing
river
A river is a natural flowing watercourse, usually freshwater
Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the ...
s. The layer ''Anserimimus'' was found, is dating from the early
Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interv ...
stage of the
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
, or about 70
million years ago
The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds.
Usage
Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
.
Aside from ''Gallimimus'', other theropods found in the Nemegt Formation include the gigantic ''
Tarbosaurus'' and ''
Deinocheirus
''Deinocheirus'' ( ) is a genus of large ornithomimosaur that lived during the Late Cretaceous around 70 million years ago. In 1965, a pair of large arms, shoulder girdles, and a few other bones of a new dinosaur were first discovered in the Ne ...
'', as well as smaller
dromaeosaurids,
oviraptorosaurs,
troodontid
Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil disco ...
s, and
bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s.
Herbivore
A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthp ...
s are represented by the
hadrosaurid
Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which incl ...
s ''
Barsboldia'' and ''
Saurolophus
''Saurolophus'' (; meaning "lizard crest") is a genus of large hadrosaurid dinosaur from the Late Cretaceous period of Asia and North America, that lived in what is now the Horseshoe Canyon and Nemegt formations about 70 million to 68 millio ...
'', the
ankylosaurid ''
Tarchia'' and several
titanosaurian
sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their ...
s and
pachycephalosaurians. The rock
facies
In geology, a facies ( , ; same pronunciation and spelling in the plural) is a body of rock with specified characteristics, which can be any observable attribute of rocks (such as their overall appearance, composition, or condition of formatio ...
of this formation suggest the presence of stream and river channels, mudflats, and shallow lakes. Sediments also indicate that there existed a rich habitat, offering diverse food in abundant amounts that could sustain massive Cretaceous dinosaurs.
[Novacek, M. (1996). Dinosaurs of the Flaming Cliffs. Bantam Doubleday Dell Publishing Group Inc. New York, New York. ]
See also
*
Timeline of ornithomimosaur research
This timeline of ornithomimosaur research is a chronological listing of events in the history of paleontology focused on the ornithomimosaurs, a group of bird-like theropods popularly known as the ostrich dinosaurs. Although fragmentary, probable, ...
References
External links
*
Anserimimus' at The Theropod Database
{{Taxonbar, from1=Q132481
Late Cretaceous dinosaurs of Asia
Fossil taxa described in 1988
Ornithomimids
Taxa named by Rinchen Barsbold
Maastrichtian life
Nemegt fauna