Anostoma Depressum
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''Anostoma depressum'' is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of air-breathing
land snail A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial gastropod mollusks that have shells (those without shells are known as ...
, a terrestrial
pulmonate Pulmonata or pulmonates, is an informal group (previously an order, and before that a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group includ ...
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is esti ...
in the family
Odontostomidae Odontostomidae is a taxonomic family of medium-sized to large, air-breathing, tropical and sub-tropical land snails, terrestrial pulmonate gastropod molluscs in the superfamily Orthalicoidea.Breure A. S. H. & Romero P. (2012). "Support and su ...
.


Distribution

This species occurs in
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
.


Original description

Lamarck originally described ''Anostoma depressum'' under name ''Anostoma depressa'' as follows: (French description add here) Translated from French language: "Shell suborbicular, convex on both sides, a little depressed, obtusely carinated, imperforate, glabrous; whitish with a circular reddish line above; aperture five toothed; lip strongly reflexed."


Shell description

The shell of this species is biconvex, the altitude about half the diameter, angular at the periphery, whitish, more or less brown-tinted, the base dappled with oblong spots or streaks arranged concentrically, having a dark band along the basal suture, the upper surface sparsely spotted near the periphery, and having a brown band revolving below the suture, usually with a fainter one above it. Surface is finely striate above and below, fresh specimens showing dense fine spiral lines, especially on the last whorl. The shell is composed of 4
whorls A whorl ( or ) is an individual circle, oval, volution or equivalent in a whorled pattern, which consists of a spiral or multiple concentric objects (including circles, ovals and arcs). Whorls in nature File:Photograph and axial plane floral d ...
, that are nearly flat. The last whorl is angular or carinated at the periphery, its last half is straightened and turned upward, showing two or three grooves or pits behind the lip. The shell
aperture In optics, an aperture is a hole or an opening through which light travels. More specifically, the aperture and focal length of an optical system determine the cone angle of a bundle of rays that come to a focus in the image plane. An opt ...
is subhorizontal, semicircular, obstructed by 5 or 6 teeth. Peristome is expanded and reflexed, pinkish-brown or flesh tinted. Parietal margin is bearing two or three teeth; a minute tubercular angle-lamella is often obsolete, an erect, compressed parietal lamella, and a small infraparietal lamella at about the middle of the parietal margin. Outer margin with three (or four) folds; columellar fold usually wanting; upper and lower palatal folds well developed, the former a little larger and more oblique; suprapalatal fold tubercular, small or minute. The width of adult shells is 31–37 mm, the height is 15-17.5 mm. Pilsbry H. A. 1901. in Tryon G. W. & Pilsbry H. A. 1901-1902
''Oriental Bulimoid Helicidae. Odontostominae. Cerionidae'' Volume 14
Manual of Conchology. Second series: Pulmonata. page 112-114.
The immature shell measures 21 by 18 mm in width, the height is 10.5 mm. The axis is hollow and the mouth possesses no lamellae. Distinguished from ''
Anostoma octodentatum ''Anostoma octodentatum'', common name Brazilian up-mouth snail, is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Odontostomidae. ''Anostoma octodentatum'' is the type species of the genus '' Anos ...
'' chiefly by the narrower and tinted lip, and fewer teeth. It is usually smaller, more angular at the periphery, and less densely mottled beneath.


Anatomy

Anatomy of ''Anostoma depressum'' was examined by Harold Heath in 1914. Heath H. 1914. ''The anatomy of two Brazilian land-shells, Anostoma depressum and
Tomigerus clausus ''Tomigerus clausus'' is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Odontostomidae. Distribution This species occurs in Brazil. Anatomy The anatomy of ''Tomigerus clausus'' was examined by ...
''
Proceedings of the Academy of Natural Sciences of Philadelphia, 65688
692
plate 31
913 __NOTOC__ Year 913 ( CMXIII) was a common year starting on Friday (link will display the full calendar) of the Julian calendar. Events By place Byzantine Empire * June 6 – Emperor Alexander III dies of exhaustion while playing ...
The
kidney The kidneys are two reddish-brown bean-shaped organs found in vertebrates. They are located on the left and right in the retroperitoneal space, and in adult humans are about in length. They receive blood from the paired renal arteries; blood ...
, placed between the pericardial and body walls on the anterior face of the last fold, presents the usual sac-like appearance. Evidently the reno-pericardial opening is minute. The ureter, likewise, is difficult to trace throughout a portion of its course. Its external opening is immediately behind that of the
alimentary canal The gastrointestinal tract (GI tract, digestive tract, alimentary canal) is the tract or passageway of the digestive system that leads from the mouth to the anus. The GI tract contains all the major organs of the digestive system, in humans ...
, and from this point is readily followed to the region of the
pericardium The pericardium, also called pericardial sac, is a double-walled sac containing the heart and the roots of the great vessels. It has two layers, an outer layer made of strong connective tissue (fibrous pericardium), and an inner layer made of ...
, where it decreases in caliber and passes into a network of
blood vessel The blood vessels are the components of the circulatory system that transport blood throughout the human body. These vessels transport blood cells, nutrients, and oxygen to the tissues of the body. They also take waste and carbon dioxide away ...
s. It appears that the kidney is in contact, along its inner face, with a slender sack or tube with which the ureter communicates. There are no special features of the digestive system of ''Anostoma depressum''. The description of the reproductive system of '' Ringicella ringens'' as described by Fischer (1869) applies in all essential details to ''Anostoma depressum'' with the exception of a penis retractor muscle attached to the vas deferens. In ''Anostoma depressum'' the external reproductive pore, located on the right side of the head, leads into a small atrium with which the penis, hermaphroditic duct, and seminal receptacle attach. The first named is a heavy, muscular organ, accompanied by the usual slender vas deferens, their union being effected at some distance from the distal end of the penis, which thus forms a flagellum. The penis and flagellum are both sharply defined and appear to be without muscular attachments, though bound to the body wall by delicate connective tissue strands. In the neighborhood of the outer opening, however, the penis sheath affords attachment for three or four slender muscle bands that Heath inclined to regard as penis retractors. The hermaphroditic duct, a highly glandular duct of comparatively even diameter, traverses the greater portion of the first coil of the body to enter the accessory glands. All of these structures are in a quiescent condition, and in size and configuration probably fall far short of their fully developed state in the sexually mature condition. The same is likewise true of the gonad. While the duct leading to it from the accessory glands is clearly apparent throughout the first part of its course, it gradually approaches the vanishing point, and cannot with certainty be traced to a gonad, which is accordingly drawn in its hypothetical position.


Feeding habits

''Anostoma depressum'' is phytophagous feeding on vegetables and moss.


References

This article incorporates public domain text from references


Further reading

* Fischer. 1869. Jour. de Conchyl., page 261.


External links


Photo of the shell of ''Anostoma depressum''

Four fine views of one shell of ''Anostoma depressum''

Photo of a shell of ''Anostoma depressum''
{{Taxonbar, from=Q4770185 Odontostomidae Gastropods described in 1822