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''Andalucia'' is a genus of
jakobid Jakobids are an order of free-living, heterotrophic, flagellar eukaryotes in the supergroup Excavata. They are small (less than 15 μm), and can be found in aerobic and anaerobic environments. The order Jakobida, believed to be monophyletic, con ...
s.


Classification

The morphology of ''Andalucia'' broadly resembles that of other
jakobid Jakobids are an order of free-living, heterotrophic, flagellar eukaryotes in the supergroup Excavata. They are small (less than 15 μm), and can be found in aerobic and anaerobic environments. The order Jakobida, believed to be monophyletic, con ...
s. Molecular data has not always been conclusive, but recent phylogenomic analyses indicate that ''Andalucia'' is a sister group to the other jakobids, in other words more closely related to them than to the
Heterolobosea The Percolozoa are a group of colourless, non-photosynthetic Excavata, including many that can transform between amoeboid, flagellate, and cyst stages. Characteristics Most Percolozoa are found as bacterivores in soil, fresh water and occasionall ...
or
Euglenozoa Euglenozoa are a large group of flagellate Discoba. They include a variety of common free-living species, as well as a few important parasites, some of which infect humans. Euglenozoa are represented by three major clades, i.e., Kinetoplastea, Di ...
(the other two groups in the
Discoba Excavata is a major supergroup of unicellular organisms belonging to the domain Eukaryota. It was first suggested by Simpson and Patterson in 1999 and introduced by Thomas Cavalier-Smith in 2002 as a formal taxon. It contains a variety of fre ...
). The α-
tubulin Tubulin in molecular biology can refer either to the tubulin protein superfamily of globular proteins, or one of the member proteins of that superfamily. α- and β-tubulins polymerize into microtubules, a major component of the eukaryotic cytoske ...
gene of ''Andalucia'' more closely resembles that of
opisthokont The opisthokonts () are a broad group of eukaryotes, including both the animal and fungus kingdoms. The opisthokonts, previously called the "Fungi/Metazoa group", are generally recognized as a clade. Opisthokonts together with Apusomonadida and ...
s and
diplomonad The diplomonads (Greek for "two units") are a group of flagellates, most of which are parasitic. They include ''Giardia duodenalis'', which causes giardiasis in humans. They are placed among the metamonads, and appear to be particularly close r ...
s than its closer relatives, the apparent result of
horizontal gene transfer Horizontal gene transfer (HGT) or lateral gene transfer (LGT) is the movement of genetic material between Unicellular organism, unicellular and/or multicellular organisms other than by the ("vertical") transmission of DNA from parent to offsprin ...
.


Species

As of 2009, the genus contains two species: * '' A. incarcerata'' was until 2006 known as ''
Jakoba ''Jakoba'' is a genus in the taxon Excavata, and currently has a single described species, ''Jakoba libera'' described by Patterson in 1990, and named in honour of Dutch botanist (Algology, Myology and Lichenology) Jakoba Ruinen. (Previously desc ...
incarcerata''. It lives in
sulfide Sulfide (British English also sulphide) is an inorganic anion of sulfur with the chemical formula S2− or a compound containing one or more S2− ions. Solutions of sulfide salts are corrosive. ''Sulfide'' also refers to chemical compounds lar ...
-rich sediments, especially marine
intertidal The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of Marine habitat, habitats with var ...
ones. Currently it is the type species of the genus '' Stygiella'' Pánek, Táborský & Čepička 2015. * '' A. godoyi'' is a soil
heterotroph A heterotroph (; ) is an organism that cannot produce its own food, instead taking nutrition from other sources of organic carbon, mainly plant or animal matter. In the food chain, heterotrophs are primary, secondary and tertiary consumers, but ...
. Analysis of DNA sequences from the environment suggest at least two additional species which have not been isolated or formally described.


References

Excavata genera Jakobids {{Excavata-stub