Anasazisaurus Horneri
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''Anasazisaurus'' ( ; "Anasazi lizard") is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
saurolophine Saurolophinae is a subfamily (biology), subfamily of hadrosaurid dinosaurs. It has since the mid-20th century generally been called the Hadrosaurinae, a group of largely non-crested hadrosaurs related to the crested sub-family Lambeosaurinae. How ...
hadrosaurid Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which inclu ...
("duckbill")
ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (), that started out as small, bipedal running grazers and grew in size and numbers until they became one of the most successful groups of herbivores in the Cretaceous world ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
that lived about 74 million years ago, in the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
Period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...
. It was found in the Farmington Member of the
Kirtland Formation The Kirtland Formation (originally the Kirtland Shale) is a sedimentary geological formation. Description The Kirtland Formation is the product of alluvial muds and overbank sand deposits from the many channels draining the coastal plain that ...
, in the
San Juan San Juan, Spanish for Saint John, may refer to: Places Argentina * San Juan Province, Argentina * San Juan, Argentina, the capital of that province * San Juan, Salta, a village in Iruya, Salta Province * San Juan (Buenos Aires Underground), ...
Basin of
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
. Only a partial skull has been found to date. It was first described as a specimen of ''Kritosaurus'' by Jack Horner, and has been intertwined with ''
Kritosaurus ''Kritosaurus'' is an incompletely known genus of hadrosaurid (duck-billed) dinosaur. It lived about 74.5-66 million years ago, in the Late Cretaceous of North America. The name means "separated lizard" (referring to the arrangement of the cheek ...
'' since its description. It is known for its short nasal crest, which stuck out above and between its eyes for a short distance.


History

Adrian Hunt Adrian is a form of the Latin given name Adrianus or Hadrianus. Its ultimate origin is most likely via the former river Adria from the Venetic and Illyrian word ''adur'', meaning "sea" or "water". The Adria was until the 8th century BC the main ...
and
Spencer G. Lucas Spencer George Lucas is an American paleontologist and stratigrapher, and curator of paleontology at the New Mexico Museum of Natural History and Science. His main areas of study are late Paleozoic, Mesozoic and early Cenozoic vertebrate fossils ...
,
American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, pe ...
paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
s, named this dinosaur in 1993. Its name is derived from the
Anasazi The Ancestral Puebloans, also known as the Anasazi, were an ancient Native American culture that spanned the present-day Four Corners region of the United States, comprising southeastern Utah, northeastern Arizona, northwestern New Mexico, a ...
, an outdated term for the Ancestral Pueblo Native American people, and the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
word ''sauros'' ("lizard"). The Ancestral Puebloans were famous for their
cliff-dwelling In archeology, archaeology, cliff dwellings are dwellings formed by using niches or caves in high cliffs, and sometimes with excavation or additions in the way of masonry. Two special types of cliff dwelling are distinguished by archaeologist ...
s, such as those in
Chaco Canyon Chaco Culture National Historical Park is a United States National Historical Park in the American Southwest hosting a concentration of pueblos. The park is located in northwestern New Mexico, between Albuquerque and Farmington, in a remote ca ...
, near the location of
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
''Anasazisaurus'' remains. The term "Anasazi" itself is actually a
Navajo language Navajo or Navaho (; Navajo: or ) is a Southern Athabaskan language of the Na-Dené family, through which it is related to languages spoken across the western areas of North America. Navajo is spoken primarily in the Southwestern United States ...
word, ''anaasází'' ("enemy ancestors"). There is one known
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
(''A. horneri''), which is named in honor of Jack Horner, an influential paleontologist who first described the skull in 1992. The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
skull (and only known specimen) was collected in the late 1970s by a
Brigham Young University Brigham Young University (BYU, sometimes referred to colloquially as The Y) is a private research university in Provo, Utah. It was founded in 1875 by religious leader Brigham Young and is sponsored by the Church of Jesus Christ of Latter-day ...
field party working in San Juan County, and is housed at BYU as BYU 12950. Horner originally assigned the skull to ''
Kritosaurus navajovius ''Kritosaurus'' is an incompletely known genus of hadrosaurid (duck-billed) dinosaur. It lived about 74.5-66 million years ago, in the Late Cretaceous of North America. The name means "separated lizard" (referring to the arrangement of the cheek ...
''. Later, Hunt and Lucas claimed that the ''Kritosaurus'' holotype was undiagnostic and judged the genus to be a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a s ...
''. They decided to give BYU 12950 the new name ''Anasazisaurus horneri''. Some later authors, including those of the most recent review, followed this decision. However, others disagreed, including Thomas Williamson, who made the most detailed published case. In a paper published in 2014, Alberto Prieto-Marquez agreed that ''Anasazisaurus horneri'' is similar to ''Kritosaurus navajovius'', but found it distinct enough to be considered a valid species of ''Kritosaurus'', as ''K. horneri''. While this conclusion has been accepted in later papers, one study finds it to be a member of the
Saurolophini Saurolophini is a tribe of saurolophine hadrosaurids native to the Americas and Asia. It includes ''Saurolophus'' (from Canada and Mongolia), ''Augustynolophus'' (from the United States), and ''Prosaurolophus'' (from Alberta, Canada, and Montana, ...
, although it did not include ''Kritosaurus navajovius''.


Description

The anatomy of ''Anasazisaurus'' is poorly known. The skull is somewhat poorly preserved, lacks the lower jaw, beak, and quadrate, and was only recently fully prepared. It has a sort of tab or flange of bone, from the nasals, that rises between and above the eyes and folds back under itself. This unique crest allows it to be distinguished from similar hadrosaurs, like ''
Gryposaurus ''Gryposaurus'' (meaning "hooked-nosed (Ancient Greek, Greek ''grypos'') lizard"; sometimes incorrectly translated as "griffin (Latin ''gryphus'') lizard") was a genus of hadrosaur, duckbilled dinosaur that lived about 80 to 75 million years ...
''. The top of the crest is roughened, and the maximum preserved length of the skull is ~90 centimeters (~35 in). According to
Gregory S. Paul Gregory Scott Paul (born December 24, 1954) is an American freelance researcher, author and illustrator who works in paleontology, and more recently has examined sociology and theology. He is best known for his work and research on theropod dino ...
, it was about 7.5 meters long and weigh up to 2500 kg.Paul, G.S., 2010, ''The Princeton Field Guide to Dinosaurs'', Princeton University Press p. 306


Classification

''Anasazisaurus'' is a saurolophine hadrosaurid, meaning it lacked a hollow crest. The most recent review recognized it as distinct, but did not try to place it beyond
Hadrosaurinae Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which includ ...
.


Paleoecology and paleobiology

''Anasazisaurus'' was discovered in the Farmington Member of the
Kirtland Formation The Kirtland Formation (originally the Kirtland Shale) is a sedimentary geological formation. Description The Kirtland Formation is the product of alluvial muds and overbank sand deposits from the many channels draining the coastal plain that ...
, which is one of the lower members. This formation dates from the late
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...
stages of the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
Period (73-74 million years ago), and is also the source of several other dinosaurs, like ''
Bistahieversor ''Bistahieversor'' (meaning "Bistahi destroyer"), also known as the "Bisti Beast", is a genus of eutyrannosaurian tyrannosauroid dinosaur; the genus contains only a single known species, ''B. sealeyi'', described in 2010, from the Late Cretaceo ...
'', ''
Kritosaurus ''Kritosaurus'' is an incompletely known genus of hadrosaurid (duck-billed) dinosaur. It lived about 74.5-66 million years ago, in the Late Cretaceous of North America. The name means "separated lizard" (referring to the arrangement of the cheek ...
'', ''
Naashoibitosaurus ''Naashoibitosaurus'' (from Navajo language, Navajo —"creek lizard") is a genus of hadrosaurid dinosaur that lived about 73 million years ago, in the Late Cretaceous, and was found in the Kirtland Formation of the San Juan, New Mexico, San ...
'', ''
Parasaurolophus ''Parasaurolophus'' (; meaning "near crested lizard" in reference to '' Saurolophus)'' is a genus of herbivorous hadrosaurid ornithopod dinosaur that lived in what is now North America and possibly Asia during the Late Cretaceous Period, abou ...
'', ''
Pentaceratops ''Pentaceratops'' ("five-horned face") is a genus of herbivorous ceratopsid dinosaur from the late Cretaceous Period of what is now North America. Fossils of this animal were first discovered in 1921, but the genus was named in 1923 when its typ ...
'', ''
Nodocephalosaurus ''Nodocephalosaurus'' (meaning "knob headed lizard") is a monospecific genus of ankylosaurid dinosaur from New Mexico that lived during the Late Cretaceous (late Campanian to early Maastrichtian stage, 73.49 to 73.04 Ma) in what is now the De-na- ...
'', ''
Saurornitholestes ''Saurornitholestes'' ("lizard-bird thief") is a genus of carnivorous dromaeosaurid theropod dinosaur from the late Cretaceous of Canada (Alberta) and the United States (Montana, New Mexico, Alabama, North Carolina, and South Carolina). Two spec ...
'', and ''
Ziapelta ''Ziapelta'' is an extinct genus of ankylosaurid. Its fossils have been found in the Hunter Wash and De-na-zin members of the Kirtland Formation of Upper Cretaceous (Campanian) New Mexico. It was named in 2014, in a research paper led by ankyl ...
''.Weishampel, David B.; Barrett, Paul M.; Coria, Rodolfo A.; Le Loeuff, Jean; Xu Xing; Zhao Xijin; Sahni, Ashok; Gomani, Elizabeth, M.P.; and Noto, Christopher R. (2004). "Dinosaur Distribution". ''The Dinosauria'' (2nd). 517–606. As a hadrosaurid, ''Anasazisaurus'' would have been a large
bipedal Bipedalism is a form of terrestrial locomotion where an organism moves by means of its two rear limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' 'double' ...
/
quadruped Quadrupedalism is a form of locomotion where four limbs are used to bear weight and move around. An animal or machine that usually maintains a four-legged posture and moves using all four limbs is said to be a quadruped (from Latin ''quattuor' ...
al
herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
, eating plants with a sophisticated skull that permitted a grinding motion analogous to
chewing Chewing or mastication is the process by which food is crushed and ground by teeth. It is the first step of digestion, and it increases the surface area of foods to allow a more efficient break down by enzymes. During the mastication process, th ...
. Its
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
were continually replacing and packed into
dental batteries Dinosaur teeth have been studied since 1822 when Mary Ann Mantell (1795-1869) and her husband Dr Gideon Algernon Mantell (1790-1852) discovered an ''Iguanodon'' tooth in Sussex in England. Unlike mammal teeth, individual dinosaur teeth are gener ...
that contained hundreds of teeth, only a relative handful of which were in use at any time. Plant material would have been cropped by its beak, and held in the jaws by a muscular
cheek The cheeks ( la, buccae) constitute the area of the face below the eyes and between the nose and the left or right ear. "Buccal" means relating to the cheek. In humans, the region is innervated by the buccal nerve. The area between the inside ...
. Feeding would have been from the ground up to ~4 meters (13 ft) above.


See also

*
Timeline of hadrosaur research A timeline is a display of a list of events in chronological order. It is typically a graphic design showing a long bar labelled with dates paralleling it, and usually contemporaneous events. Timelines can use any suitable scale representin ...


References

{{Taxonbar, from=Q1705543 Saurolophines Late Cretaceous dinosaurs of North America Natural history of San Juan County, New Mexico Taxa named by Spencer G. Lucas Fossil taxa described in 1993 Paleontology in New Mexico Campanian genus first appearances Campanian genus extinctions Ornithischian genera