plants
Plants are predominantly Photosynthesis, photosynthetic eukaryotes of the Kingdom (biology), kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all curr ...
that grow in an
alpine climate
Alpine climate is the typical weather (climate) for elevations above the tree line, where trees fail to grow due to cold. This climate is also referred to as a mountain climate or highland climate.
Definition
There are multiple definitions of ...
, which occurs at high
elevation
The elevation of a geographic location is its height above or below a fixed reference point, most commonly a reference geoid, a mathematical model of the Earth's sea level as an equipotential gravitational surface (see Geodetic datum § Vert ...
and above the
tree line
The tree line is the edge of the habitat at which trees are capable of growing. It is found at high elevations and high latitudes. Beyond the tree line, trees cannot tolerate the environmental conditions (usually cold temperatures, extreme snowp ...
. There are many different plant species and
taxa
In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
that grow as a
plant community
A plant community is a collection or association of plant species within a designated geographical unit, which forms a relatively uniform patch, distinguishable from neighboring patches of different vegetation types. The components of each plant co ...
in these
alpine tundra
Alpine tundra is a type of natural region or biome that does not contain trees because it is at high elevation, with an associated alpine climate, harsh climate. As the latitude of a location approaches the poles, the threshold elevation for alp ...
. These include
perennial grass
A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also widel ...
es,
sedges
The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, the largest being the "true sedges" genus ''Carex'' wit ...
,
forb
A forb or phorb is an herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in biology and in vegetation ecology, especially in relation to grasslands and understory. Typically these are dicots without woo ...
s,
cushion plant
A cushion plant is a compact, low-growing, mat-forming plant that is found in alpine, subalpine, arctic, or subarctic environments around the world. The term "cushion" is usually applied to woody plants that grow as spreading mats, are limited in ...
s,
moss
Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hor ...
es, and
lichen
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.. Alpine plants are adapted to the harsh conditions of the alpine environment, which include low temperatures, dryness, ultraviolet radiation, wind, drought, poor nutritional soil, and a short growing season.
Some alpine plants serve as
medicinal plants
Medicinal plants, also called medicinal herbs, have been discovered and used in traditional medicine practices since prehistoric times. Plants synthesize hundreds of chemical compounds for various functions, including defense and protection ag ...
.
Ecology
Alpine plants occur in a
tundra
In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless moun ...
: a type of
natural region
A natural region (landscape unit) is a basic geographic unit. Usually, it is a region which is distinguished by its common natural features of geography, geology, and climate.
From the ecology, ecological point of view, the naturally occurring fl ...
or biome that does not contain trees. Alpine tundra occurs in mountains worldwide. It transitions to subalpine forests below the tree line; stunted forests occurring at the forest-tundra
ecotone
An ecotone is a transition area between two biological communities, where two communities meet and integrate. It may be narrow or wide, and it may be local (the zone between a field and forest) or regional (the transition between forest and gras ...
are known as ''
Krummholz
''Krummholz'' (german: krumm, "crooked, bent, twisted" and ''Holz'', "wood") — also called ''knieholz'' ("knee timber") — is a type of stunted, deformed vegetation encountered in the subarctic and subalpine
Montane ecosystems are fou ...
''. With increasing elevation, it ends at the
snow line
The climatic snow line is the boundary between a snow-covered and snow-free surface. The actual snow line may adjust seasonally, and be either significantly higher in elevation, or lower. The permanent snow line is the level above which snow wil ...
where snow and ice persist through summer, also known as the Nival Zone.
Alpine plants are not limited to higher
elevation
The elevation of a geographic location is its height above or below a fixed reference point, most commonly a reference geoid, a mathematical model of the Earth's sea level as an equipotential gravitational surface (see Geodetic datum § Vert ...
s. However, high-elevation areas have different ecology than those growing at higher latitudes. One of the biggest distinctions is that the lower bound of a tropical alpine area is difficult to define due to a mixture of human disturbances, dry climates, and a naturally lacking tree line. The other major difference between tropical and arctic-alpine ecology is the temperature differences. The tropics have a summer/winter cycle every day, whereas the higher latitudes stay cold both day and night.
In the northern latitudes, the main factor to overcome is the cold. Intense
frost action
Frost weathering is a collective term for several mechanical weathering processes induced by stresses created by the freezing of water into ice. The term serves as an umbrella term for a variety of processes such as frost shattering, frost wedg ...
processes have a profound effect on what little soil there is and the vegetation of arctic-alpine regions. Tropical alpine regions are subject to these conditions as well, but they seldom happen. Because northern alpine areas cover a massive area it can be difficult to generalize the characteristics that define the ecology. One factor in alpine ecology is wind in an area. Wind
pruning
Pruning is a horticultural, arboricultural, and silvicultural practice involving the selective removal of certain parts of a plant, such as branches, buds, or roots.
The practice entails the ''targeted'' removal of diseased, damaged, dead, ...
is a common sight within northern alpine regions. Along with wind pruning, wind erosion of vegetation mats is a common sight throughout
Alaska
Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
.
Growth
Long-lived perennial herbs are the most common type of plant in alpine environments, with most having a large, well-developed
root
In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the sur ...
and/or
rhizome
In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
system. These underground systems store
carbohydrate
In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may or ma ...
s through the winter which are then used in the spring for new shoot development. Some species of saxifrages have small root systems, but are
evergreen
In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
. The leaves of these plants store energy in the form of carbohydrates and
lipid
Lipids are a broad group of naturally-occurring molecules which includes fats, waxes, sterols, fat-soluble vitamins (such as vitamins A, D, E and K), monoglycerides, diglycerides, phospholipids, and others. The functions of lipids include ...
photoperiod
Photoperiodism is the physiological reaction of organisms to the length of night or a dark period. It occurs in plants and animals. Plant photoperiodism can also be defined as the developmental responses of plants to the relative lengths of light a ...
.
Seedling
A seedling is a young sporophyte developing out of a plant embryo from a seed. Seedling development starts with germination of the seed. A typical young seedling consists of three main parts: the radicle (embryonic root), the hypocotyl (embry ...
establishment is very slow and occurs less often than
vegetative reproduction
Vegetative reproduction (also known as vegetative propagation, vegetative multiplication or cloning) is any form of asexual reproduction occurring in plants in which a new plant grows from a fragment or cutting of the parent plant or spec ...
. In the first year of growth of perennial alpine plants, most of the photosynthate is used in establishing a stable root system which is used to help prevent desiccation and for carbohydrate storage over winter. In this year, the plant may produce a few true leaves, but usually, only the
cotyledon
A cotyledon (; ; ; , gen. (), ) is a significant part of the embryo within the seed of a plant, and is defined as "the embryonic leaf in seed-bearing plants, one or more of which are the first to appear from a germinating seed." The numb ...
s are produced. It usually takes a few years for plants to become well established.
Adaptations
Alpine plants can exist at very high elevations, from , depending on location. For example, there is a
moss
Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hor ...
that grows at on
Mount Everest
Mount Everest (; Tibetan: ''Chomolungma'' ; ) is Earth's highest mountain above sea level, located in the Mahalangur Himal sub-range of the Himalayas. The China–Nepal border runs across its summit point. Its elevation (snow heig ...
. ''
Arenaria bryophylla
''Eremogone bryophylla'' is a flowering plant in the sandwort ('' Eremogone'') genus. It occurs in the Himalaya of China, Tibet, Nepal, and India
India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the s ...
'' is the highest flowering plant in the world, occurring as high as .
To survive, alpine plants are adapted to the conditions at high elevations, including cold, dryness, high levels of
ultraviolet radiation
Ultraviolet (UV) is a form of electromagnetic radiation with wavelength from 10 nm (with a corresponding frequency around 30 PHz) to 400 nm (750 THz), shorter than that of visible light, but longer than X-rays. UV radiation i ...
, and difficulty of reproduction. These conditions are linked to topographical slope, ultimately affecting plant diversity and distribution. This is due to steeper slopes causing faster soil erosion which in turn impedes plant growth, seed distribution, and seed settlement. Furthermore, the slope of the topography directly affects many other abiotic factors including temperature, solar radiation, moisture content, and nutritional content in the soil.
Surviving low temperature extremes
Most alpine plants are faced with low-temperature extremes at some point in their lives. There are several ways that a plant can survive these extremes. Plants can avoid exposure to low temperature by using different forms of seasonal
phenology
Phenology is the study of periodic events in biological life cycles and how these are influenced by seasonality, seasonal and interannual variations in climate, as well as environmental factor, habitat factors (such as elevation).
Examples includ ...
,
morphology
Morphology, from the Greek and meaning "study of shape", may refer to:
Disciplines
* Morphology (archaeology), study of the shapes or forms of artifacts
* Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
, or by variable growth form preference. They can also avoid the freezing of their exposed tissues by increasing the number of solutes in their tissues, known as
freezing-point depression
Freezing-point depression is a drop in the minimum temperature at which a substance freezes, caused when a smaller amount of another, non- volatile substance is added. Examples include adding salt into water (used in ice cream makers and for ...
. Another, somewhat similar, method plants may use to avoid freezing is
supercooling
Supercooling, also known as undercooling, is the process of lowering the temperature of a liquid or a gas below its melting point without it becoming a solid. It achieves this in the absence of a seed crystal or nucleus around which a crystal ...
, which prevents
ice
Ice is water frozen into a solid state, typically forming at or below temperatures of 0 degrees Celsius or Depending on the presence of impurities such as particles of soil or bubbles of air, it can appear transparent or a more or less opaq ...
crystallization within plant tissues. These methods are only sufficient when the temperature is only moderately cold. In the alpine zone, temperatures are often low enough that these methods are not sufficient.. When plants need a more permanent solution, they can develop freeze tolerance. Plants can also
dehydrate
In physiology, dehydration is a lack of total body water, with an accompanying disruption of metabolic processes. It occurs when free water loss exceeds free water intake, usually due to exercise, disease, or high environmental temperature. Mil ...
their cells by moving
water
Water (chemical formula ) is an inorganic, transparent, tasteless, odorless, and nearly colorless chemical substance, which is the main constituent of Earth's hydrosphere and the fluids of all known living organisms (in which it acts as a ...
cell
Cell most often refers to:
* Cell (biology), the functional basic unit of life
Cell may also refer to:
Locations
* Monastic cell, a small room, hut, or cave in which a religious recluse lives, alternatively the small precursor of a monastery ...
where
ice crystals
Ice crystals are solid ice exhibiting atomic ordering on various length scales and include hexagonal columns, hexagonal plates, dendritic crystals, and diamond dust.
Formation
The hugely symmetric shapes are due to depositional growth, na ...
will not cause damage. When all of these strategies fail to prevent
frost
Frost is a thin layer of ice on a solid surface, which forms from water vapor in an above-freezing atmosphere coming in contact with a solid surface whose temperature is below freezing, and resulting in a phase change from water vapor (a gas) ...
damage, alpine plants often have the capacity to repair or replace the
organs
In biology, an organ is a collection of tissues joined in a structural unit to serve a common function. In the hierarchy of life, an organ lies between tissue and an organ system. Tissues are formed from same type cells to act together in a f ...
damaged. As it is often difficult to prevent damage, many alpine plants depend on the replacement of their organs. They help make this possible by placing their
meristems
The meristem is a type of tissue found in plants. It consists of undifferentiated cells (meristematic cells) capable of cell division. Cells in the meristem can develop into all the other tissues and organs that occur in plants. These cells conti ...
below ground, where temperatures are generally warmer.
Photosynthesis and respiration rates
Photosynthesis
Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored i ...
and
respiration
Respiration may refer to:
Biology
* Cellular respiration, the process in which nutrients are converted into useful energy in a cell
** Anaerobic respiration, cellular respiration without oxygen
** Maintenance respiration, the amount of cellul ...
rates are not uniform throughout the growing season. At the start of the growing season, new shoots have low net photosynthesis rates and high respiration rates due to rapid growth of new shoots. As the temperature rises in a plants
microclimate
A microclimate (or micro-climate) is a local set of atmospheric conditions that differ from those in the surrounding areas, often with a slight difference but sometimes with a substantial one. The term may refer to areas as small as a few squ ...
, the net photosynthesis rates will increase as long as ample water is available and will peak during flowering. Alpine plants are able to start photosynthesizing and reach maximum photosynthesis rates at lower temperatures compared to plants adapted to lower elevations and warmer climates. This is due to the combined effects of
genotype
The genotype of an organism is its complete set of genetic material. Genotype can also be used to refer to the alleles or variants an individual carries in a particular gene or genetic location. The number of alleles an individual can have in a ...
and environmental factors.
Avoidance of desiccation
In alpine areas, water availability is often variable.
Bryophyte
The Bryophyta s.l. are a proposed taxonomic division containing three groups of non-vascular land plants (embryophytes): the liverworts, hornworts and mosses. Bryophyta s.s. consists of the mosses only. They are characteristically limited in ...
s and
lichen
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.desiccation
Desiccation () is the state of extreme dryness, or the process of extreme drying. A desiccant is a hygroscopic (attracts and holds water) substance that induces or sustains such a state in its local vicinity in a moderately sealed container.
...
tolerance, which contributes to their abundance throughout all alpine areas habitats. Among
higher plants
Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They ...
, tissue desiccation is rare at high elevation. If it does occur, it normally happens to plants growing on exposed sites, where
wind
Wind is the natural movement of air or other gases relative to a planet's surface. Winds occur on a range of scales, from thunderstorm flows lasting tens of minutes, to local breezes generated by heating of land surfaces and lasting a few hou ...
stress is increased. Alpine plants avoid water loss by deep
root
In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the sur ...
ing and increased
stoma
In botany, a stoma (from Greek ''στόμα'', "mouth", plural "stomata"), also called a stomate (plural "stomates"), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is bor ...
tal control. Plants at low elevation normally reach a maximum stomatal opening in the morning while alpine plants reach maximum opening mid-day when the temperature is greatest. Alpine
succulent plants
In botany, succulent plants, also known as succulents, are plants with parts that are thickened, fleshy, and engorged, usually to retain water in arid climates or soil conditions. The word ''succulent'' comes from the Latin word ''sucus'', meani ...
often utilize
CAM photosynthesis
Crassulacean acid metabolism, also known as CAM photosynthesis, is a carbon fixation pathway that evolved in some plants as an adaptation to arid conditions that allows a plant to photosynthesize during the day, but only exchange gases at night. ...
to avoid water loss.
Avoidance of ultraviolet radiation
Because
ultraviolet
Ultraviolet (UV) is a form of electromagnetic radiation with wavelength from 10 nanometer, nm (with a corresponding frequency around 30 Hertz, PHz) to 400 nm (750 Hertz, THz), shorter than that of visible light, but longer than ...
radiation tends to increase with elevation, it is often assumed to be a stress factor among alpine plants. In the past, there have been many attempts to research how ultraviolet radiation may influence alpine plant forms. However, it is uncertain if the growth and development of plants are affected by ultraviolet radiation. It is also not clear if the radiation is responsible for promoting genetic differentiation, leading to stunted growth forms.
Reproduction
Alpine plants use both
sexual reproduction
Sexual reproduction is a type of reproduction that involves a complex life cycle in which a gamete ( haploid reproductive cells, such as a sperm or egg cell) with a single set of chromosomes combines with another gamete to produce a zygote tha ...
and
asexual reproduction
Asexual reproduction is a type of reproduction that does not involve the fusion of gametes or change in the number of chromosomes. The offspring that arise by asexual reproduction from either unicellular or multicellular organisms inherit the fu ...
. Sexual reproduction has limits in high alpine areas, especially in areas with a short
growing season
A season is a division of the year marked by changes in weather, ecology, and the amount of daylight. The growing season is that portion of the year in which local conditions (i.e. rainfall, temperature, daylight) permit normal plant growth. Whil ...
in alpine zones at high latitudes. In tropical alpine zones with a year-round growing season, such as the northern
Andes
The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
, plants can flower year-round. Regardless of when alpine plants flower, pollinators are often scarce. The activity of pollinators decreases with increasing elevation.. The most common
pollinators
A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains.
Insects are the maj ...
in the alpine zone are
bumblebees
A bumblebee (or bumble bee, bumble-bee, or humble-bee) is any of over 250 species in the genus ''Bombus'', part of Apidae, one of the bee families. This genus is the only extant group in the tribe Bombini, though a few extinct related gener ...
and
flies
Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced ...
. Plants utilize different strategies to deal with these limits, including alternate flowering time and clonal propagation.
Early flowering plants
Some plants flower immediately after snow melting or soil thawing. These early flowering plants always form their flowers in the previous season, called preformation. This flower
primordium
A primordium (; plural: primordia; synonym: anlage) in embryology, is an organ or tissue in its earliest recognizable stage of development. Cells of the primordium are called primordial cells. A primordium is the simplest set of cells capable o ...
is produced one to three years before flowering which ensures that flowering is not delayed after snowmelt and that with the right environmental conditions, there will be enough time for seed set. Consequently, they risk frost damage to the preformed
inflorescence
An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
. In order to minimize frost damage, preformed flowers are often surrounded by tightly packed
bracts
In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
that are densely covered in
trichomes
Trichomes (); ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a plant ...
. This helps to keep the interior of a flower
bud
In botany, a bud is an undeveloped or embryonic shoot and normally occurs in the axil of a leaf or at the tip of a stem. Once formed, a bud may remain for some time in a dormant condition, or it may form a shoot immediately. Buds may be spec ...
warm. Because of early-season pollinator limitation, plants that bloom early generally have a low rate of reproductive success. One advantage of flowering early is that seeds that are produced have a greater chance of developing to maturity before the next freeze. They also have a high
outcrossing
Out-crossing or out-breeding is the technique of crossing between different breeds. This is the practice of introducing distantly related genetic material into a breeding line, thereby increasing genetic diversity.
Outcrossing can be a useful ...
rate, which helps to increase
genetic diversity
Genetic diversity is the total number of genetic characteristics in the genetic makeup of a species, it ranges widely from the number of species to differences within species and can be attributed to the span of survival for a species. It is dis ...
. Speed and time of flowering is dependent on the time of snowmelt, temperature, and photoperiod, but usually occurs 10 to 20 days after snowmelt.
Mid-season flowering
Approximately half of all alpine species flower in mid-season. Flowering at the seasonal peak combines some of the advantages and risks of early flowering and late flowering plants. Some mid-season plants pre-form their inflorescences, but not all do.
Late flowering
Late flowering occurs after the main growing season ends. They have a high seed output but their seeds have a reduced rate of maturing because of time constraints. These plants tend towards
self pollination
Self-pollination is a form of pollination in which pollen from the same plant arrives at the Stigma (botany), stigma of a flower (in flowering plants) or at the ovule (in gymnosperms). There are two types of self-pollination: in autogamy, pollen i ...
,
apomixis
In botany, apomixis is asexual reproduction without fertilization. Its etymology is Greek for "away from" + "mixing". This definition notably does not mention meiosis. Thus "normal asexual reproduction" of plants, such as propagation from cuttin ...
, and
vivipary
In plants, vivipary occurs when seeds or embryos begin to develop before they detach from the parent. Plants such as some Iridaceae and Agavoideae grow cormlets in the axils of their inflorescences. These fall and in favourable circumstances the ...
.
Clonal propagation
Because investment in flowers and seed production can be costly for alpine plants, they often use
clonal propagation
Vegetative reproduction (also known as vegetative propagation, vegetative multiplication or cloning) is any form of asexual reproduction occurring in plants in which a new plant grows from a fragment or cutting of the parent plant or spec ...
. This strategy becomes increasingly more frequent as elevation increases, and is most common among
cryptogams
A cryptogam (scientific name Cryptogamae) is a plant (in the wide sense of the word) or a plant-like organism that reproduces by spores, without flowers or seeds. The name ''Cryptogamae'' () means "hidden reproduction", referring to the fact ...
and
grasses
Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns and ...
. Some alpine plants use it as their predominant method of reproduction. In these plants, sexual reproduction is rare and does not contribute significantly to reproductive output. An example of such a plant is ''
Carex
''Carex'' is a vast genus of more than 2,000 species of grass-like plants in the family Cyperaceae, commonly known as sedges (or seg, in older books). Other members of the family Cyperaceae are also called sedges, however those of genus ''Carex'' ...
'' ''curvula'', which is estimated to have a clonal age of approximately 2000 years.
After establishment, each year new shoot growth occurs rapidly from the perennating bud which is usually located close to the soil surface. This growth occurs after snowmelt when the soil temperature is above 0 °C. Some species, like '' Erythronium grandiflorum'', can begin new shoot growth before snowmelt as they have their perennating buds located in
bulb
In botany, a bulb is structurally a short stem with fleshy leaves or leaf basesBell, A.D. 1997. ''Plant form: an illustrated guide to flowering plant morphology''. Oxford University Press, Oxford, U.K. that function as food storage organs duri ...
s buried deep in the soil. As new leaves protrude from the snow, the new shoots give off heat from thermal reradiation and/or respiratory heat which melts the surrounding snow. This exposes more soil to
solar radiation
Solar irradiance is the power per unit area (surface power density) received from the Sun in the form of electromagnetic radiation in the wavelength range of the measuring instrument.
Solar irradiance is measured in watts per square metre (W/m ...
, heating it up and allowing new growth to accelerate.
Medicinal alpine plants
There are a number of alpine plants that are used
economically
Economics () is the social science that studies the production, distribution, and consumption of goods and services.
Economics focuses on the behaviour and interactions of economic agents and how economies work. Microeconomics analyzes ...
. In the
Himalayas
The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...
, hundreds of species are traded for medicinal and aromatic uses. It is estimated that the annual trade of these plants amounts to millions of US dollars. Many households in rural
Nepal
Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne,
सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is mai ...
and
India
India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
rely on
medicinal
Medicine is the science and practice of caring for a patient, managing the diagnosis, prognosis, prevention, treatment, palliation of their injury or disease, and promoting their health. Medicine encompasses a variety of health care practic ...
alpine plant trade as a source of income. This creates an increased need to focus on plant
conservation
Conservation is the preservation or efficient use of resources, or the conservation of various quantities under physical laws.
Conservation may also refer to:
Environment and natural resources
* Nature conservation, the protection and managem ...
in these areas, ensuring
sustainable harvest
The sustainable yield of natural capital is the ecological yield that can be extracted without reducing the base of capital itself, i.e. the surplus required to maintain ecosystem services at the same or increasing level over time. The term only ...
as well as
ecosystem
An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syste ...
Rheum australe
''Rheum australe'', synonym ''Rheum emodi'', is a flowering plant in the family Polygonaceae. It is commonly known as Himalayan rhubarb, Indian rhubarb and Red-veined pie plant. It is a medicinal herb used in the Indian Unani system of medici ...
'', and ''
Bergenia
''Bergenia'' (elephant-eared saxifrage, elephant's ears) is a genus of ten species of flowering plants in the family Saxifragaceae, native to central Asia, from Afghanistan to China and the Himalayan region.
Description
They are clump-forming, ...
''. In the Indian Himalayas, the alpine medicinal plants such as ''
Dactylorhiza hatagirea
''Dactylorhiza hatagirea'' is a species of orchid generally found growing in the Himalayas, from Pakistan to SE Tibet, at altitudes of . It is locally called 'salam panja' or 'hatta haddi'. It is called 'panchaule' (पाँचऔंले) in ...
Alpinum (garden)
An alpine garden (or alpinarium, alpinum) is a domestic or botanical garden, or more often a part of a larger garden, specializing in the collection and cultivation of alpine plants growing naturally at high altitudes around the world, such as in ...
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Alpine Garden Society
The Alpine Garden Society headquarters are at Pershore, Worcestershire. It is an "International Society for the cultivation, conservation and exploration of alpine and rock garden plants, small hardy herbaceous plants, hardy and half-hardy bulb ...
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Flora of the Alps
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The Alps are one of the great mountain range systems of Europe stretching approximately 1,200 kilometres (750 mi) across eight Alpine countries from Austria and Slovenia in the east, Switzerland, Liechtenstein, Germany, France to th ...