Abyssinian Wheatear
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The Abyssinian wheatear (''Oenanthe lugubris''), or Abyssinian black wheatear, is a species of
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
in the family
Muscicapidae The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, Bluethroat (''Luscinia svecica)'' and Norther ...
. It is found from
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
to southern
Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
and northeastern
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...
.


Taxonomy

The Abyssinian wheatear was first formally described in 1837 as ''Saxicola lugubris'' by the German naturalist and
explorer Exploration refers to the historical practice of discovering remote lands. It is studied by geographers and historians. Two major eras of exploration occurred in human history: one of convergence, and one of divergence. The first, covering most ...
Eduard Rüppell Wilhelm Peter Eduard Simon Rüppell (20 November 1794 – 10 December 1884) was a German Natural history, naturalist and List of explorers, explorer. Rüppell is occasionally transliterated to "Rueppell" for the English alphabet, due to german ort ...
with its type locality given as "the rocky valleys of the
Abyssinia The Ethiopian Empire (), also formerly known by the exonym Abyssinia, or just simply known as Ethiopia (; Amharic and Tigrinya: ኢትዮጵያ , , Oromo: Itoophiyaa, Somali: Itoobiya, Afar: ''Itiyoophiyaa''), was an empire that historica ...
n province of Simien, and on the volcanic hills around
Gondar Gondar, also spelled Gonder (Amharic: ጎንደር, ''Gonder'' or ''Gondär''; formerly , ''Gʷandar'' or ''Gʷender''), is a city and woreda in Ethiopia. Located in the North Gondar Zone of the Amhara Region, Gondar is north of Lake Tana on t ...
" (''Vorkommen in der felsigen Thälern der Abyssinischen provinz Simen, und auf der vulkanischen Hügeln um Gondar''). It is regarded, by some authorities, as being the same species as the
mourning wheatear The mourning wheatear (''Oenanthe lugens'') is a bird, one of 14 species of wheatear found in northern Africa and the Middle East. It is a small passerine in a group formerly classed as members of the thrush family Turdidae, but now more generall ...
(''Oenanthe lugens'') of Northern Africa and the Middle East but the I.O.C. regard it as a valid species. The Abyssinian wheatear is regarded as being closely related to the
Arabian wheatear The Arabian wheatear (''Oenanthe lugentoides'') is a species of bird in the family Muscicapidae. It is found in Oman, Saudi Arabia and Yemen Yemen (; ar, ٱلْيَمَن, al-Yaman), officially the Republic of Yemen,, ) is a country in We ...
(''Oenanthe lugentoides''), which has also been regarded as being the same species as the mourning wheatear. These species may form a
species complex In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
.


Subspecies

The I.O.C. recognises the following subspecies: Schalow's wheatear (''O. l. schalowi'') from the highlands of southern Kenya and northeastern Tanzania is regarded as a valid species by some authorities but the I.O.C. has pended its endorsement of this position while it waits for the results of genetic studies to be published. ''O. l. vauriei'' is found in northeastern Somalia.


Description

The Abyssinian wheatear is a dark wheatear which shows
sexual dichromatism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
, with the males being darker than the females. The males have black upperparts, throat and breast and a grey cap with darker streaks. The colour of the belly varies, both within individuals and geographic populations, from white to black. The females are mainly dark brown with heavy streaking on their paler breast and belly. Both males and females have buff under tail coverts, and the tail has a black inverted T pattern on a buff background, best seen in flight. The length of the nominate subspecies is .


Subspecific variation

The three subspecies of Abyssinian wheatear vary as follows: * ''O. l. lugubris'': The males have a grey or greyish brown crown with darker streaks, a black face and upper parts with buff rump and upper tail coverts change in colour from buff to white at the base and the upper sides of the tail. The extent of white on the underparts varies, with most birds being black except for the white under tail coverts, while others may have black restricted to the throat and breast. The females have dark brown upper parts apart from the white, buff, or light orange rump and upper tail coverts, with the underparts being greyish buff with darker streaks on the throats and breasts, which frequently extend onto the whitish flanks. * ''O. l. schalowi'': This subspecies is a little larger than the nominate subspecies and has browner upper parts with a buff orange lower belly and upper tail coverts. * ''O.l. vauriei'': In this subspecies, the males have a pale greyish white to dull sooty buff crown streaked with brown, and a blackish forehead. The crown colour can extend as a diffuse patch onto the upper mantle, while the remainder of the upper parts of the rump is black with pale fringes on the fresh feathers. The rump and upper tail coverts are white to orange buff, a little lighter in colour than those on Schalow's wheatear. The lower head and upper breast are black, with the lower breast and flanks being greyish white with a white belly and buff under tail coverts. The females of this subspecies are paler than female Schalow's wheatear.


Distribution and habitat

The Abyssinian wheatear is found in northeastern Africa, from Eritrea and northwestern Somalia south to northeastern Tanzania. The subspecies ''schalowi'' occurs in southern Kenya and northern Tanzania, while ''vauriei'' in eastern Somalia, and they both have disjunct ranges from the nominate subspecies of Eritrea, Ethiopia, and northwestern Somalia. This bird occurs in rocky areas where there are some trees and bushes.


Behaviour

Abyssinian wheatears are normally solitary outside their breeding season, and in the breeding season, they are typically encountered in pairs. They are generally unafraid of humans and perch in plain sight on rocks, boulders, and shrubs. They may be inactive and shelter during midday. Their diet is mainly invertebrates, caught by dropping on them from a perch or by foraging on the ground. They will also eat fruit, such as berries.


Breeding

The Abyssinian wheatear breeds from the early spring to late summer (March to August) in Ethiopia and Eritrea, in April and May in Somalia, from April to July in the Kenyan
Great Rift Valley The Great Rift Valley is a series of contiguous geographic trenches, approximately in total length, that runs from Lebanon in Asia to Mozambique in Southeast Africa. While the name continues in some usages, it is rarely used in geology as it i ...
, and in March in northern Tanzania. The males display by adopting an upright stance with their heads raised before hopping around the female with droopy, quivering wings, interrupted by quick chases of the female in flight. The female makes the nest in the form of a loose cup with a flat top of the grass and other plant material, lined with softer material such as moss or mammal hair. There may be a small platform of stones either at the entrance to the nest, under it, or around it. The nest is typically situated in a crevice or hole with a maximum depth of . This may be among rocks, on cliffs, in stone walls, on the banks of a river, or side of a wadi. They are double-brooded, and each clutch is 5 to 6 eggs. The juveniles from the first brood have been recorded bringing food to the nestlings of the 2nd brood in Schalow's wheatear.


Migration

Abyssinian wheatears are largely resident, with some post-breeding dispersal. The subspecies ''vauriei'' disappears from its eastern Somali breeding areas from June to January and may migrate to lower altitudes when not breeding.


References

{{Taxonbar, from=Q11858961 Abyssian wheatear Birds of East Africa Abyssian wheatear