conifer
Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
ous tree in the pine family
Pinaceae
The Pinaceae, or pine family, are conifer trees or shrubs, including many of the well-known conifers of commercial importance such as Cedrus, cedars, firs, Tsuga, hemlocks, larches, pines and spruces. The family is included in the order Pinales, ...
. This tree is native to the mountains of western North America, including the
Cascade Range
The Cascade Range or Cascades is a major mountain range of western North America, extending from southern British Columbia through Washington and Oregon to Northern California. It includes both non-volcanic mountains, such as the North Cascades, ...
and southern
Rocky Mountains
The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico in ...
Arizona
Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
,
New Mexico
)
, population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano)
, seat = Santa Fe
, LargestCity = Albuquerque
, LargestMetro = Tiguex
, OfficialLang = None
, Languages = English, Spanish ( New Mexican), Navajo, Ker ...
, and
Northern Mexico
Northern Mexico ( es, el Norte de México ), commonly referred as , is an informal term for the northern cultural and geographical area in Mexico. Depending on the source, it contains some or all of the states of Baja California, Baja California ...
. It naturally occurs at elevations between .
It is popular as an ornamental landscaping tree and as a
Christmas tree
A Christmas tree is a decorated tree, usually an evergreen conifer, such as a spruce, pine or fir, or an artificial tree of similar appearance, associated with the celebration of Christmas. The custom was further developed in early modern ...
.
Description
This large evergreen conifer grows best in the central
Sierra Nevada
The Sierra Nevada () is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primarily ...
of California, where the record specimen was recorded as tall and measured in
diameter at breast height
Diameter at breast height, or DBH, is a standard method of expressing the diameter of the trunk or bole of a standing tree. DBH is one of the most common dendrometric measurements.
Tree trunks are measured at the height of an adult's breast, ...
(dbh) in
Yosemite National Park
Yosemite National Park ( ) is an American national park in California, surrounded on the southeast by Sierra National Forest and on the northwest by Stanislaus National Forest. The park is managed by the National Park Service and covers an ar ...
.American Forestry Association. 1978. National register of big trees. American Forests 84(4):19-47 The typical size of white fir ranges from tall and up to dbh. The largest specimens are found in the central Sierra Nevada, where the largest diameter recorded was found in
Sierra National Forest
Sierra National Forest is a U.S. national forest located on the western slope of central Sierra Nevada in Central California and bounded on the northwest by Yosemite National Park and the south by Kings Canyon National Park. The forest is kno ...
at (1972); the west slope of the Sierra Nevada is also home to the tallest specimen on record, in height. ''Abies concolor'' subsp. ''concolor'' (
Rocky mountain
The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico in ...
white fir) rarely exceeds tall or dbh. Large (but not huge) trees in good soil range from tall and from dbh in California and southwestern Oregon and to tall and dbh in
Arizona
Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
and New Mexico. The dead tree tops sometimes fork into new growth.
The gray bark is usually at least thick, and brown-hued inside. The
leaves
A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are needle-like, flattened, long and wide by thick, green to glaucous blue-green above, and with two glaucous blue-white bands of stomatal bloom below, and slightly notched to bluntly pointed at the tip. The leaf arrangement is spiral on the shoot, but with each leaf variably twisted at the base so they all lie in either two more-or-less flat ranks on either side of the shoot, or upswept across the top of the shoot but not below the shoot.
The
cones
A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, circular) to a point called the apex or vertex.
A cone is formed by a set of line segments, half-lines, or lines conn ...
are long and broad, green or purple ripening pale brown, with about 100–150 scales; the scale bracts are short, and hidden in the closed cone. The winged
seed
A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s are released when the cones disintegrate at maturity about 6 months after pollination.
White fir can live over 300 years.
Subspecies
As treated here, there are two
subspecies
In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
; these are also variously treated at either the lower rank of
variety
Variety may refer to:
Arts and entertainment Entertainment formats
* Variety (radio)
* Variety show, in theater and television
Films
* ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont
* ''Variety'' (1935 film), ...
by some authors, or as distinct species by others:
Botanical collection
White fir was first collected by
Augustus Fendler
Augustus Fendler (January 10, 1813 – November 27, 1883), alternatively written as August Fendler, was a Prussian-born American natural history collector.
Fendler gained his first taste of exploration as a physician's assistant. He partook in a ...
New Mexico
)
, population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano)
, seat = Santa Fe
, LargestCity = Albuquerque
, LargestMetro = Tiguex
, OfficialLang = None
, Languages = English, Spanish ( New Mexican), Navajo, Ker ...
in 1846–1847. Fender's patron George Engelmann, a St. Louis area physician and botanist, then described the plant. This tree was first collected in California by
William Lobb
William is a male given name of Germanic origin.Hanks, Hardcastle and Hodges, ''Oxford Dictionary of First Names'', Oxford University Press, 2nd edition, , p. 276. It became very popular in the English language after the Norman conquest of Engl ...
on his expedition to California of 1849–1853, after it was overlooked by David Douglas in his 1825–27 expedition to the Pacific coast region.
The specific epithet ''concolor'' means "all one color".
Distribution
This tree is native to the mountains of western North America from the southern Cascade Range in
Oregon
Oregon () is a U.S. state, state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington (state), Washington, while the Snake River delineates much of it ...
, south throughout
California
California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
and into the
Sierra de San Pedro Mártir
Sierra de San Pedro Mártir ( Kiliwa: ʔxaal haq, en, mountains of Saint Peter the Martyr) is a mountain range located within southern Ensenada Municipality and southern Baja California state, of northwestern Mexico.
It is a major mountain ran ...
in northern
Baja California
Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
Rocky Mountains
The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico in ...
in
Utah
Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...
and
Colorado
Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...
Arizona
Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
, New Mexico, and
Northern Mexico
Northern Mexico ( es, el Norte de México ), commonly referred as , is an informal term for the northern cultural and geographical area in Mexico. Depending on the source, it contains some or all of the states of Baja California, Baja California ...
. It naturally occurs between above sea level.
Ecology
White fir is very closely related to
grand fir
''Abies grandis'' (grand fir, giant fir, lowland white fir, great silver fir, western white fir, Vancouver fir, or Oregon fir) is a fir native to the Pacific Northwest and Northern California of North America, occurring at altitudes of sea leve ...
(''Abies grandis''), with subspecies ''lowiana'' being particularly similar to the interior variety of grand fir, ''A. grandis'' var. ''idahoensis'', intergrading with it where they meet in the Cascades of Central Oregon. White fir occupies a broader range, including dryer and colder sites, being more drought tolerant and having thicker, fire-resistant bark. In Mexico, it is replaced by further close relatives, Durango fir (''A. durangensis'') and
Mexican fir
Mexican may refer to:
Mexico and its culture
*Being related to, from, or connected to the country of Mexico, in North America
** People
*** Mexicans, inhabitants of the country Mexico and their descendants
*** Mexica, ancient indigenous peopl ...
(''A. mexicana'').
Like grand fir, white fir is more
shade tolerant
In ecology, shade tolerance is a plant's ability to tolerate low light levels. The term is also used in horticulture and landscaping, although in this context its use is sometimes imprecise, especially in labeling of plants for sale in commercial ...
than
Douglas fir
The Douglas fir (''Pseudotsuga menziesii'') is an evergreen conifer species in the pine family, Pinaceae. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Oregon pine, and Columbian pine. There are three va ...
, but less so than
western hemlock
''Tsuga heterophylla'', the western hemlock or western hemlock-spruce, is a species of hemlock native to the west coast of North America, with its northwestern limit on the Kenai Peninsula, Alaska, and its southeastern limit in northern Sonoma ...
and
western redcedar
''Thuja plicata'' is an evergreen coniferous tree in the cypress family Cupressaceae, native to western North America. Its common name is western redcedar (western red cedar in the UK), and it is also called Pacific redcedar, giant arborvitae, w ...
. White fir is a
climax species
Climax species, also called late seral, late-successional, K-selected or equilibrium species, are plant species that can germinate and grow with limited resources; e.g., they need heat exposure or low water availability. They are the species wi ...
, which means the forest has reached complex maturity in forest succession in western coniferous forests of the U.S. White fir and
yellow pine
In ecology and forestry, yellow pine refers to a number of conifer species that tend to grow in similar plant communities and yield similar strong wood. In the Western United States, yellow pine refers to Jeffrey pine or ponderosa pine. In the ...
(
ponderosa pine
''Pinus ponderosa'', commonly known as the ponderosa pine, bull pine, blackjack pine, western yellow-pine, or filipinus pine is a very large pine tree species of variable habitat native to mountainous regions of western North America. It is the ...
/
Jeffrey pine
''Pinus jeffreyi'', also known as Jeffrey pine, Jeffrey's pine, yellow pine and black pine, is a North American pine tree. It is mainly found in California, but also in the westernmost part of Nevada, southwestern Oregon, and northern Baja Califo ...
) have co-existed for millennia in old growth forests throughout their range. In the presence of logging of large diameter trees and exclusion of cleansing wildfires, young trees have become abundant over the past two centuries. White fir had been regarded as a pest in the past by those in the lumber industry, but this opinion has changed. White fir is now one of the most important of all commercial softwoods according to the Western Wood Products Association.
The white fir trait of retaining lower limbs creates an escape route for medium-to-small forest birds (such as
spotted owl
The spotted owl (''Strix occidentalis'') is a species of true owl. It is a resident species of old-growth forests in western North America, where it nests in tree hollows, old bird of prey nests, or rock crevices. Nests can be between high a ...
) from larger flying predators and provides a drip zone around the roots for collecting moisture. The retained limbs can become a
fuel ladder
A fuel ladder or ladder fuel is a firefighting term for live or dead vegetation that allows a fire to climb up from the landscape or forest floor into the tree canopy. Limbing-up white fir, instead of removing medium to large diameter trees, in areas where the public is more likely to start fires can help keep other trees and specifically giant sequoia from experiencing canopy fire. Recent concern for sequoia groves has caused agencies to call for removal of white fir in the Sierra Nevada. While sequoia seedlings and young saplings are highly susceptible to mortality or serious injury by fire; mature sequoias are fire adapted with: fire-resistant bark, elevated canopies, self-pruning lower branches, latent buds, and serotinous cones. The sequoia ecosystem is incomplete without the mixed pine/fir and oak that make up the mid and understory.
Giant Sequoia
''Sequoiadendron giganteum'' (giant sequoia; also known as giant redwood, Sierra redwood, Sierran redwood, California big tree, Wellingtonia or simply big treea nickname also used by John Muir) is the sole living species in the genus ''Sequoiade ...
's cones release seeds when the heat of fire triggers them to open while the thick bark protects the inner
cambium
A cambium (plural cambia or cambiums), in plants, is a tissue layer that provides partially undifferentiated cells for plant growth. It is found in the area between xylem and phloem. A cambium can also be defined as a cellular plant tissue from w ...
from fire damage.
This tree is host to fir mistletoe (''Phoradendron pauciflorum''), a
parasitic plant
A parasitic plant is a plant that derives some or all of its nutritional requirements from another living plant. They make up about 1% of angiosperms and are found in almost every biome. All parasitic plants develop a specialized organ called the ...
. It is attacked by many types of insects, such as the fir engraver (''Scolytus ventralis'').
Dependent species
Mature white fir–yellow pine forests support
old-growth
An old-growth forestalso termed primary forest, virgin forest, late seral forest, primeval forest, or first-growth forestis a forest that has attained great age without significant disturbance, and thereby exhibits unique ecological feature ...
dependent wildlife species such as California spotted owl (''Strix occidentalis occidentalis''), Mount Pinos
sooty grouse
The sooty grouse (''Dendragapus fuliginosus'') is a species of forest-dwelling grouse native to North America's Pacific Coast Ranges.del Hoyo, J., Elliott, A., & Sargatal, J., eds. (1994). ''Handbook of the Birds of the World'' 2: 401-402. Lynx ...
(''Dendragapus fuliginosus howardi''), and Pacific fisher (''Pekania pennanti''). The spotted owl and fisher utilize cavities in
decadent
The word decadence, which at first meant simply "decline" in an abstract sense, is now most often used to refer to a perceived decay in standards, morals, dignity, religious faith, honor, discipline, or skill at governing among the members of ...
large-diameter white fir for nesting and denning. The Mount Pinos sooty grouse requires large diameter trees for thermal cover and its winter diet consists of mostly white fir and yellow pine needles. This subspecies of sooty grouse has been extirpated along with a significant number of large diameter white fir from much of its range. Other subspecies of sooty grouse also utilize Douglas fir, which does not occur in the range of Mount Pinos sooty grouse. Squirrel also frequent the tree's branches.
Deer browse the foliage of this species and porcupines chew the bark. Songbirds, grouse, and various mammals eat the seeds.
Uses
White fir and grand fir were used by Native Americans for medicinal use involving powdered bark or pitch to treat tuberculosis or skin ailments. The
Nlaka'pamux
The Nlaka'pamux or Nlakapamuk ( ; ), also previously known as the ''Thompson'', ''Thompson River Salish'', ''Thompson Salish'', ''Thompson River Indians'' or ''Thompson River people'', and historically as the ''Klackarpun'', ''Haukamaugh'', ''Kni ...
used the bark to cover lodges and make canoes. Branches were also used as bedding.
White fir is a preferred construction species because of its nail-holding ability, lightness in weight, and resistance to split, twist, and pitch. It is straight-grained, non-resinous, fine-textured, stiff, and strong.
It is popular as a
Christmas tree
A Christmas tree is a decorated tree, usually an evergreen conifer, such as a spruce, pine or fir, or an artificial tree of similar appearance, associated with the celebration of Christmas. The custom was further developed in early modern ...
and for
Christmas
Christmas is an annual festival commemorating Nativity of Jesus, the birth of Jesus, Jesus Christ, observed primarily on December 25 as a religious and cultural celebration among billions of people Observance of Christmas by country, around t ...
decoration owing to its soft needles, generally excellent needle retention and abundance. It is often marketed as concolor or white fir.
Cultivation
White fir is widely planted as an
ornamental tree
Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that i ...
in parks and larger
garden
A garden is a planned space, usually outdoors, set aside for the cultivation, display, and enjoyment of plants and other forms of nature. The single feature identifying even the wildest wild garden is ''control''. The garden can incorporate bot ...
s, particularly some
cultivar
A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...
s of subsp. ''concolor'' selected for very bright glaucous blue foliage, such as cv. 'Violacea'. The dwarf cultivar 'Compacta', growing to a maximum height and spread of , has gained the
Royal Horticultural Society
The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity.
The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nort ...
's
Award of Garden Merit
The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions.
History
The Award of Garden Merit ...
.
Gallery
Image:Abies_concolor_8076.jpg, ''A. concolor'' subsp. ''lowiana'' foliage upperside
Image:Abies_concolor_8065.jpg, ''A. concolor'' subsp. ''lowiana'' foliage underside
Image:Baby whitefir.jpg, A young sapling of subsp. ''lowiana'' on
Mount Whitney
Mount Whitney (Paiute: Tumanguya; ''Too-man-i-goo-yah'') is the highest mountain in the contiguous United States and the Sierra Nevada, with an elevation of . It is in East–Central California, on the boundary between California's Inyo and Tu ...
Image:White fir MN 2007.JPG, White fir in garden environment at
Minnesota Landscape Arboretum
The Minnesota Landscape Arboretum is a horticultural garden and arboretum located about west of Chanhassen, Minnesota at 3675 Arboretum Drive, Chaska, Minnesota. It is part of the Department of Horticultural Science in the College of Food, Agri ...
Image:Young Abies concolor.jpg, Small stand of young white firs on
Cuyamaca Peak
Cuyamaca Peak is a mountain peak of the Cuyamaca Mountains range, in San Diego County, Southern California.
Geography
At , its summit is the second highest point in San Diego County.
Cuyamaca Peak is located roughly from the Pacific Ocean, wi ...
California
Image:Toro Peak 537.jpg, White firs at
Toro Peak
Toro Peak, in Southern California, is the highest mountain in the Santa Rosa Mountain Range. It is located south of Palm Springs, west of the Salton Sea, and east of Temecula, in the County of Riverside, California.
Part of the mountain is u ...
, CA.
File:Arceuthobium abietinum 1.jpg, Foliage, with '' Arceuthobium abietinum'' infestation